Conicera, Meigen, Meigen, 1830

Liu, Guang-Chun, 2023, Revision of the genus Conicera Meigen (Diptera, Phoridae) from China, Zootaxa 5380 (5), pp. 401-445 : 440-443

Key to species (♂)

1. Vein Rs with basisetula ( Fig. 30D )....................................................................... 17

Vein Rs without basisetula.............................................................................. 2

2. Posterior face of mid femur with sense-organ ( Figs. 24D , 27D )................................................. 3

Posterior face of mid femur without sense-organ............................................................. 7

3. Sense-organ of mid femur including a rounded pit (or groove) and a apical process................................. 4

Sense-organ of mid femur only with a groove, but without a apical process........................................ 6

4. Sense-organ composed of a long groove and a long apical process ( Fig. 24D )...................... C. formosensis Brues

Sense-organ with a rounded pit and a short apical process..................................................... 5

5. Sensory pit large, its diameter more than 1/2 width of mid femur ( Fig. 26D ); the aspect ratio of first flagellomere is 3:1.......................................................................................... C. tibialis Schmitz

Sensory pit small, its diameter less than 1/2 width of mid femur ( Fig. 25D ); the aspect ratio of first flagellomere equaled to or less than 2:1.......................................................................... C. simili s (Haliday)

6. The naked area at the base of posterior face of mid femur extends to half length of mid femur; the sensory groove is short and broad ( Fig. 27D ); basidorsal process of right surstylus long and finger-like ( Fig. 27C ).................. C. kempi Brunetti

The naked area at the base of posterior face of mid femur extends to one third of mid femur; the sensory groove is long and narrow ( Fig. 28B ), basidorsal process of right surstylus short, rectangular ( Figs. 28D, F ).............. C. rectangularis Liu

7. The base part of left surstylus is equaled to or broader than the apical part........................................ 8

The base part of left surstylus is significantly contracted and narrower than the apical part........................... 11

8. Left surstylus with a long, band-shaped basiprocess; posterior edge of left and right surstyli with 7–8 teeth respectively.... 9

Left surstylus without a long band-shaped basiprocess; posterior edge of left and right Surstyli only with1–4 teeth respectively......................................................................................... 10

9. Both anterior and posterior edges of surstylus are similar length, so the surstylus looks square ( Figs. 4B–E ).... C. ulrichi Liu

Anterior edge of surstylus longer than posterior edge, so the surstylus looks rhombic ( Figs.3B–E ) C. gracilis Michailovskaya

10. Apex of surstyli pointed ( Figs. 1D–G ); fine veins whitish gray( Fig. 29A )........................... C. dauci (Meigen)

Apex of surstyli blunt ( Figs. 2B–E ); wing with light-brown fine veins ( Fig. 29C ).................. C. obtusifinis sp. nov.

11. Both surstyli with concave, petal-like apical edge ( Figs. 5B–E ).................................. C. petalina sp. nov.

Surstyli without this features........................................................................... 12

12. Right surstylus long, slender, base part broader than the apex; left surstylus short.................................. 13

Right surstylus various shapes, base part narrower than the apex............................................... 15

13. Right surstylus parallel-sided, and the apex beak-shaped, with 3 long, thick, curved inner teeth ( Figs. 9A–C, E ).................................................................................................. C. spinifer a Liu

Right surstylus thin and lanceolate, and the apex not beak-shaped.............................................. 14

14. The apex of right surstylus suddenly becomes thinner, without inner teeth ( Fig. 10D ).................. C. longilobusa Liu

The apex of right surstylus gradually tapered, with 3 inner teeth ( Fig. 10A ).................. C. pacifica Michailovskaya

15. Fore femur with a brownish spot ( Fig. 6A ); left surstylus short, extending downwards, pointed apically, with a basidorsal and a adjacent small processes ( Figs. 6D, F )............................................... C. maculifemoris sp. nov.

Fore femur without a brownish spot...................................................................... 16

16. Right surstylus is split into a rectangular upper branch and a crescent lower branch ( Figs. 7C, E ); left surstylus large and oval ( Figs. 7B, D )........................................................................... C. forcipis sp. nov.

Right surstylus trapezoid ( Figs. 8C, E ); left surstylus enlarged apically, anchor-shaped ( Figs. 8B, D )...................................................................................................... C. ancoraria sp. nov.

17. Left and right surstyli similar in shape, triangular or long banded.............................................. 18

Left and right surstyli very different...................................................................... 22

18. Both surstyli are long banded........................................................................... 19

Both left and right surstyli are nearly triangular............................................................ 20

19. Right surstylus with basidorsal process ( Figs. 11C, E ); left surstylus straight ( Figs. 11B, D )............ C. angusta sp. nov.

Right surstylus lacking basidorsal process ( Figs. 12C, E ); left surstylus S-form curved ( Figs. 12B, D )... C. tortuosa sp. nov.

20. Dorsal edge of left surstylus shorter than ventral edge; left and right surstyli symmetrical inversely ( Figs. 14B–E ).............................................................................................. C. convallis sp. nov.

Dorsal edge of left surstylus equaled to ventral edge......................................................... 21

21. Left surstylus with a short dorso-apical process, on which there are about 10 inner teeth ( Figs. 15B, G ); the dorso-apical corner of right surstylus pointed, with about 8 inner teeth ( Figs. 15C, H ).............................. C. chayuensis sp. nov.

Left surstylus without dorso-apical process and with 4–6 inner teeth ( Figs. 15E, I ); right surstylus with 6–8 inner teeth ( Figs. 15F, J )............................................................................ C. triangularis sp. nov.

22. Right surstylus with a basidorsal process ( Figs. 23F, H )...................................................... 28

Right surstylus without a basidorsal process............................................................... 23

23. Right surstylus oval or near square; inner teeth spread on the inner surface....................................... 24

Right surstylus trapezoidal or palm; inner teeth are mostly distributed on the edge................................. 25

24. Left surstylus oval, without a basidorsal process; with 16 inner teeth ( Figs. 13B, E )............... C. dentisparsa sp. nov.

Left surstylus kidney-shaped, with a basidorsal process and a depression at the Apex; scattered with 1–5 inner teeth ( Fig. 19B )..................................................................... C. quadrata Mostovski et Disney

25. Left surstylus axe-shaped ( Figs. 17A, D )................................................. C. securiclata sp. nov.

Left surstylus without this feature....................................................................... 26

26. Right surstylus large, palmar-shaped, and lacking inner teeth ( Figs. 18B, E )........................ C. palmata sp. nov.

Right surstylus small, with teeth........................................................................ 27

27. The apical edge of right surstylus wavy, with short setulae on surface ( Figs. 16A, C, E )............ C. margiflucta sp. nov.

The apical edge of right surstylus slightly concaved; with long setulae on surface ( Figs. 19F, H )......... C. longicilia Liu

28. Basidorsal process of right surstylus membranous.......................................................... 29

Basidorsal process of right surstylus sclerotized............................................................ 30

29. Left surstylus with a broad edge on basidorsal part ( Figs. 20G, I ); right surstylus with 2 small blunt basiprocess ( Figs. 20H, J ).................................................................................. C. marginata sp. nov.

Left surstylus only with a small basiprocess ( Figs. 20B, D ); right surstylus with 2 pointed basiprocess ( Figs. 20C, E )............................................................................................. C. reniformis Liu

30. Frons without lower fronto-orbital setae; basidorsal process of left surstylus with inner teeth ( Figs. 21B, D )................................................................................................ C. denticulata sp. nov.

Frons with lower fronto-orbital setae; basidorsal process of left surstylus without inner teeth......................... 31

31. The base of left surstylus not contracted ( Figs. 21F, I, K ); the inner teeth of the left and right surstyli thin and not sclerotized (same color with other setulae) ( Fig. 21H )............................................. C. microspinulosa sp. nov.

The base of left surstylus contracted; the inner teeth of the left and right surstyli thick and sclerotized (darker than other setulae)............................................................................................ 32

32. Terminalia Large ( Fig. 22A ), length 0.40–0.44 mm; right surstylus with 14–18 inner teeth arranged in two rows ( Figs. 22C, H )............................................................................. C. grandicaudata sp. nov. Terminalia small, length less than 0.35 mm; right surstylus with less than 10 inner teeth............................ 33

33. Posterior edge of right surstylus with 7 inner teeth, the undermost one is pointed ( Figs. 22D, F, J ); terminalia 0.31–0.33 mm ................................................................................. C. schnittmanni Schmitz

Posterior edge of right surstylus with 8–9 inner teeth, which all of them similar................................... 34

34. Basidorsal process of right surstylus short, inner teeth robust and blunt ( Figs. 23A, C ); terminalia 0.31–0.33 mm ........................................................................................... C. obtusidentis sp. nov.

Basidorsal process of right surstylus long, inner teeth slender and pointed ( Figs. 23D, F, H ); terminalia 0.31–0.33 mm ...................................................................................... C. digitalis Liu et Zhang

Figures

FIGURES 1A–G. C. dauci (Meigen), male. A. body, left view; B. head, anterior view; C. terminalia, dorsal view; D, F. left surstylus; E, G. right surstylus. Scale bars: A = 0.5 mm; B–G= 0.1mm.

FIGURES 2A–E. C. obtusifinis sp. nov., male. A. terminalia, dorsal view; B, D. left surstylus; C, E. right surstylus. Scale bars= 0.1mm.

FIGURES 3A–E. C. gracilis Michailovskaya, male.A. terminalia, dorsal view; B, D. left surstylus; C, E. right surstylus. Scale bars= 0.1mm.

FIGURES 4A–E. C. ulrichi Liu, male. A. terminalia, dorsal view; B, D. left surstylus; C, E. right surstylus. Scale bars= 0.1mm.

FIGURE 5A–E. C. petalina sp. nov., male. A. terminalia, dorsal view; B, D. left surstylus; C, E. right surstylus. Scale bars= 0.1mm.

FIGURES 6A–G. C. maculifemoris sp. nov., male. A. body, left view; B. antenna, left view; C–E. terminalia; C. dorsal view; D. left view; E. right view; F. left surstylus; G. right surstylus. Scale bars: A = 0.5mm; B–G = 0.1 mm.

FIGURES 7A–E. C. forcipis sp. nov., male. A. body, left view; B–C, terminalia; B. left view; C. right view; D. left surstylus; E. right surstylus. Scale bars: A = 0.5 mm; B–E = 0.1 mm.

FIGURES 8A–E. C. ancoraria sp. nov., male.A. body, left view; B–C. terminalia; B. left view; C. right view; D. left surstylus; E. right surstylus. Scale bars: A = 0.5 mm; B–E = 0.1mm.

FIGURES 9A–E. C. spinifera Liu., male.A–C. terminalia; A. posterior view; B. right view; C. ventral view; D. left surstylus; E. right surstylus. Scale bars= 0.1 mm.

FIGURES 10A–F. Male terminalia.A–C. C. pacifica Michailovskaya; D–F. C. longilobusa Liu; A, D. posterior view; B, E. left view; C, F. right view. Scale bars = 0.1 mm.

FIGURES 11A–E. C. angusta sp. nov., male. A–C. terminalia; A. dorsal view; B. left view; C. right view; D. left surstylus; E. right surstylus. Scale bars = 0.1mm.

FIGURES 12A–E. C. tortuosa sp. nov., male. A–C. terminalia; A. dorsal view; B. left view; C. right view; D. left surstylus; E. right surstylus. Scale bars = 0.1mm.

FIGURES 13A–F. C. dentisparsa sp. nov., male. A. body, left view; B. left surstylus; C. right surstylus; D–F. terminalia; D. dorsal view; E. left view; F. right view. Scale bars: A= 0.5 mm; B–F = 0.1 mm.

FIGURES 14A–E. C. convallis sp. nov., male. A. body, left view; B. left surstylus; C. right surstylus; D. terminalia, left view; E. terminalia, right view. Scale bars: A= 0.5 mm; B–F = 0.1 mm.

FIGURES 15A–J. Male terminalia. A–C, G–H. C. chayuensis sp. nov.; D–F, I–J. C. triangularis sp. nov.; A, D. dorsal view; B, E, G, I. left surstylus; C, F, H, J. right view. Scale bars = 0.1 mm.

FIGURES 16A–E. C. margiflucta sp. nov., male. A–C. terminalia; A. posterior view; B. left view; C. right view; D. left surstylus; E. right surstylus; F. antenna. Scale bars = 0.1 mm.

FIGURES 17A–E. C. securiclata sp. nov., male.A–C. terminalia; A. left view; B. right view; C. dorsal view; D. left surstylus; E. right surstylus. Scale bars = 0.1 mm.

FIGURES 18A–E. C. palmata sp. nov., male. A–C. terminalia; A. left view; B. right view; C. dorsal view; D. left surstylus; E. right surstylus. Scale bars = 0.1 mm.

FIGURES 19A–H. Male terminalia. A–C. C. qudrata Mostovski et Disney; D–H. C. longicilia Liu; A, D. dorsal view; B, E. G. left view; C, F, H. right view. Scale bars = 0.1 mm.

FIGURES 20A–J. Male terminalia. A–E. C. reniformis Liu; F–J. C. marginata sp. nov.; A, F. dorsal view; B, D, G, I. left surstylus; C, E, H, J. right surstylus. Scale bars = 0.1 mm.

FIGURES 21A–L. Male terminalia.A–E. C. denticulata sp. nov.; F–L. C. microspinulosa sp. nov. A. Dorsal view; H. posterior view; F. left view; G. right view; B, D, K, I. left surstylus; C, E, J, L. right surstylus. Scale bars = 0.1 mm.

FIGURES 22A–J. Male terminalia. A–C, G–H. C. grandicaudata sp. nov.; D–F, I–J. C. schnittmanni Schmitz; A, D. dorsal view; B, E, G, I. left surstylus; C, F, H, J. right surstylus. Scale bars = 0.1 mm.

FIGURES 23A–H. Male terminalia. A–C. C. obtusidentis sp. nov.; D–H. C. digitalis Liu et Zhang; A, D. dorsal view; E. left view; F. right view; B, G. left surstylus; C, H. right surstylus. Scale bars = 0.1 mm.

FIGURES 24A–D. C. formosensis Brues, male. A. terminalia, dorsal view; B. left surstylus; C. right surstylus; D. mid femur, posterior view. Scale bars = 0.1 mm.

FIGURES 25A–D. C. similis (Haliday), male. A. terminalia, dorsal view; B. left surstylus; C. right surstylus; D. mid femur, posterior view. Scale bars = 0.1 mm.

FIGURES 26A–D. C. tibialis Schmitz, male. A–C. terminalia; A. dorsal view; B. left view; C. right view; D. mid femur, posterior view. Scale bars = 0.1 mm.

FIGURES 27A–D. C. kempi Brunetti, male. A. terminalia, dorsal view; B. left surstylus; C. right surstylus; D. mid femur, posterior view. Scale bars = 0.1 mm.

FIGURES 28A–F. C. rectangularis Liu, male. A. antenna; B. mid femur, posterior view; C, E. left surstylus; D, F. right surstylus. Scale bars = 0.1 mm.

FIGURES 29A–J. Male wings.A. C. dauci (Meigen); B. C. gracilis Michailovskaya; C. C. obtusifinis sp. nov.; D. C. petalina sp. nov.; E. C. spinifera Liu; F. C. angusta sp. nov.; G. C. tortuosa sp. nov.; H. C. triangularis sp. nov.; I. C. palmata sp. nov.; J. C. quadrata Mostovski et Disney. Scale bars = 0.2 mm.

FIGURES 30A–H. Male wings. A. C. marginata sp. nov.; B. C. grandicaudata sp. nov.; C. C. schnittmanni Schmitz; D. C. obtusidentis sp. nov.; E. C. formosensis Brues; F. C. similis (Haliday); G. C. tibialis Schmitz; H. C. kempi Brunetti. Scale bars = 0.2 mm.