Osmylidia donnae, Makarkin & Archibald & Mathewes, 2021

Makarkin, Vladimir N., Archibald, S. Bruce & Mathewes, Rolf W., 2021, New Protosmylinae (Neuroptera: Osmylidae) from the early Eocene of western North America, with taxonomic remarks, Zootaxa 4980 (1), pp. 142-156 : 146

Osmylidia donnae sp. nov.

Fig. 2

Type material. Holotype Q-0317, collected by RWM, October 2001; deposited in SFU. A rather well-preserved almost complete hind wing.

Type locality and horizon. Quilchena , British Columbia, Canada; Eocene (mid-Ypresian) .

Etymology. From the first name of Donna Mathewes, for her support of fieldwork at Quilchena.

Diagnosis. May be distinguished from hind wings of other species of the genus by long distal crossvein between posterior branch of MA, MP [other species: this crossvein short or lacking as these veins fused for short distance], and membrane slightly infuscate [other species: hyaline].

Description. Hind wing 13.5 mm as preserved (estimated complete length about 14 mm), 4.3 mm wide. Costal space relatively narrow, with simple, relatively closely spaced subcostal veinlets. Sc fused with RA far from wing apex. Sc+RA veinlets simple, closely spaced. Subcostal space narrow; crossveins not detected. RA space relatively narrow, with four widely spaced crossveins. RP with 10 branches, RP8 deeply forked, other branches shallowly forked. Basal crossvein 1r-m preserved fragmentarily, near RP. M forked near base wing, slightly proximad origin of RP. MA deeply forked, proximad level of proximal crossvein between RP1, RP2. All crossveins in radial, radiomedial spaces arranged in three gradate series: proximal series with two crossveins; intermediate series with five crossveins; outer series with eight preserved crossveins (region partly missing or damaged). Anterior branch of MA pectinately branched with four branches; posterior branch of MA with terminal fork, one incompletely preserved branch. MP pectinately branched with two branches. Two crossveins (one fragmentarily preserved) between MA, MP proximad forking of MA; three crossveins between anterior, posterior branches of MA, between posterior branch of MA, MP (including long distal crossvein, i.e., posterior branch of MA, MP not fused). CuA strongly pectinately branched, with seven preserved branches (probably eight in life). CuP relatively short, forked once distally. Two crossveins between CuA, CuP. 1A with two pectinate branches. Membrane mostly slightly infuscate (as preserved; perhaps uniformly infuscate in life?).

Figures

FIGURE 2. Osmylidia donnae sp. nov., holotype hind wing Q-0317. A, photograph; B, drawing of venation (colouration omitted, refer to photograph). Posterior branch of MA in red. Scale bar = 5 mm.