Cyrtodactylus intermedius Smith

Murdoch, Matthew L., Grismer, L. Lee, Wood Jr, Perry L., Neang, Thy, Poyarkov, Nikolay A., Tri, Ngo Van, Nazarov, Roman A., Aowphol, Anchalee, Pauwels, Olivier S. G., Nguyen, Hung Ngoc & Grismer, Jesse L., 2019, Six new species of the Cyrtodactylus intermedius complex (Squamata: Gekkonidae) from the Cardamom Mountains and associated highlands of Southeast Asia, Zootaxa 4554 (1), pp. 1-62 : 20-21

Cyrtodactylus intermedius Smith

Intermediate Bow-fingered Gecko

Fig. 9 , Table 8

Gymnodactylus intermedius Smith 1917: 221

Cyrtodactylus intermedius Taylor 1963: 734

Gymnodactylus (Cyrtodactylus) intermedius Wermuth 1965: 54

Cyrtodactylus intermedius Cox et al. 1998: 86

Cyrtodactylus (Cyrtodactylus) intermedius Rösler 2000: 66

Cyrtodactylus intermedius Nguyen et al. 2009

Syntypes. MCZ R-39040 and UMMZ 78687 ; collected by M.A. Smith, March 1916 from “Khao Sebab, SE Siam ” Additional specimen . FMNH 215981 ; collected by D.L. Damman, September of 1979 from Khao Soi Daow Wildlife Sanctuary

Diagnosis. Adult males reaching 78.7 mm SVL; eight supralabials, 9 or 10 infralabials; 31–33 paravertebral tubercles; 19 or 20 longitudinal rows of dorsal tubercles; 42–44 rows of ventral scales; seven or eight expanded subdigital lamellae proximal to the digital inflection, 12–14 unmodified, distal, subdigital lamellae; 1 9–21 total subdigital lamellae on fourth toe; enlarged femoral and precloacal scales continuous; 23 or 24 enlarged femoral scales; proximal femoral scales one-half the size of distal femorals; 8–10 enlarged precloacal scales with pores on each in males; three rows of enlarged post-precloacal scales; two or three postcloacal tubercles; no pocketing between digits of hind or forefeet; dark pigmented blotches absent on top of head; posterior border of nuchal loop rounded; and four dark body bands ( Table 8). These characters are scored across all species of the Cyrtodactylus intermedius complex in Table 7.

locality of Khao Sebab. Abbreviations are listed in the Materials and Methods. R = right, L = left, / = data unobtainable,

r = regenerated, b = broken.

Remarks. No tissues were available from the type locality of Khao Sebab, Chanthaburi, Thailand however a specimen, FMNH 215981, and genetic material, LSUHC 9513, from Khao Soi Dao Wildlife Sanctuary in the adjacent mountain 30km north of the type locality were available. Upon morphological comparison of the two syntypes, UMMZ 78687 and MCZ R-39040, with FMNH 215981 from Khao Soi Dao the two populations show no morphological differences and are thus recognized here as conspecific. Thus, it is highly probable that if genetic analyses of the Khao Sebab population were conducted it would be closely related to or conspecific with the Khao Soi Dao population which falls within the Thai clade. The Thai clade is a member of the western group and the sister clade to a lineage containing [ C. thylacodactylus sp. nov. [ C. intermedius complex incertae sedis 1 and C. cardamomensis sp. nov.]] from which it is separated by 5.4%–6.9% sequence divergence ( Table 4).

Figures

FIGURE 9. Cyrtodactylus intermedius from Khao Kitchakoot, Chanthaburi, Thailand. (photo by P. L. Wood Jr.)