Monacha (Metatheba) perfrequens ( Hesse, 1914 )

Hausdorf, Bernhard, 2000, genus Monacha in the Western Caucasus Gastropoda: Hygromiidae), Journal of Natural History 34, pp. 1575-1594 : 1589-1590

Monacha (Metatheba) perfrequens ( Hesse, 1914)

(®gures 6, 15, 18)

Helix frequens, Mousson 1863: 318 [partim, non Mousson, 1859].

Helix frequens var. obscura Mousson, 1863: 318 [non O. F. MuÈller, 1774]. Locus typicus:`ReÂduktaleh’, Georgia.

Helix (Carthusiana) frequens, O. Boettger 1881: 206 [non Mousson, 1859].

Helix (Carthusiana) frequens f. minor O. Boettger, 1881: 206 [non C. B. Adams, 1851]. Locus typicus:`Gordi’, Georgia.

Helix (Carthusiana) frequens, O. Boettger 1883: 158 [non Mousson, 1859].

Helix (Carthusiana) frequens f. minor, O. Boettger 1884: 149.

Helix perfrequens O. Boettger, 1891: 83 [nomen nudum]. Locus typicus:`Kutais’, Georgia.

Helix frequens, Klika 1893: 5 [non Mousson, 1859].

Theba samsunensis, Hesse 1914: 257 , ®g. 7±9 [partim, non L. PfeiOEer, 1868].

Theba samsunensis perfrequens Hesse, 1914: 259 [partim]. Locus typicus (restrictus, Hausdorf, in press):`Kutais’, Georgia.

Theba samsunensis obscura, Hesse 1914: 259 .

Helix (Theba) samsunensis var. perfrequens, Retowski 1914: 285 .

Metatheba samsunensis, Hesse 1931: 43 , pl. 6, ®g. 55 [non L. PfeiOEer, 1868].

Theba (Metatheba) samsunensis, Likharev and Rammelmeier 1952: 433 , ®g. 358 [partim, non L. PfeiOEer, 1868].

Theba (Metatheba) samsunensis var. perfrequens, Likharev and Rammelmeier 1952: 434 .

Metatheba (Metatheba) samsunensis, Hudec and Lezhawa 1969: 125 , text-®g. 22, pl. 10, ®g. 25 [non L. PfeiOEer, 1868].

Monacha (Metatheba) samsunensis, Schileyko 1978: 288 , text-®g. 377±378, pl. 16, ®g. 159 [partim, non L. PfeiOEer, 1868].

Monacha samsunensis, Clauss 1990: 64 [non L. PfeiOEer, 1868].

Shell (®gure 6). Depressed conical±globular; with 4×±5× convex whorls; teleoconch with wrinkled growth-ridges; upper whorls often, sometimes also lower whorls, with hair-scars (or very short hairs); especially lower whorls with Ô distinct incised spiral striae; whitish or light corneous, then a whitish peripheral band might be discerned; body whorl rounded or with a slight blunt edge at the beginning, which disappears towards the aperture; aperture elliptical; upper insertion of the peristome slightly descending; peristome sharp, hardly expanded, with a whitish internal rib; umbilicus very narrow, partly or almost completely obscured by the reēcted columellar edge.

Measurements. Golovinka (n 5 20): D: 8.5±11.6 mm, x Å 5 9.6 mm; H: 5.8 ± 7.6 mm, x Å 5 6.4 mm; D/H: 1.42±1.55, x Å 5 1.49. Cap Pitsunda (n 5 30): D: 9.8±15.0 mm, x Å 5 12.8 mm; H: 6.6±10.1 mm, x Å 5 8.8 mm; D/H: 1.34±1.57, x Å 5 1.46.

Genitalia (®gure 15). The penial papilla is cylindrical or club-shaped. The two glandulae mucosae are little rami®ed. The bursa of the bursa copulatrix is oval or Ô rounded triangular. The right ommatophoral retractor passes to the left of the genitalia. The measurements of some sections of the genitalia are compiled in table 3 and the variation of the proportions of some sections of the genitalia is compiled in table 4.

Remarks. Monacha perfrequens diOEers from all other Caucasian Monacha species in the very narrow, partly or almost completely obscured umbilicus.

M. perfrequens diOEers from M. samsunensis in the lower epiphallus: penial papilla ratio and the on average longer vagina. It cannot be reliably distinguished from M. samsunensis by shell characters, however the umbilicus of M. samsunensis is usually wider.

The proportions of the genitalia of M. perfrequens are very similar to those of a new Central Anatolian Metatheba species. It diOEers from that species in the course of the right ommatophoral retractor which crosses with the penial retractor in the Central Anatolian species and in the narrower umbilicus.

Distribution and material (®gure 18). Monacha perfrequens is widespread in the Western Caucasus. It is known from Golovinka in the north to Batumi and Kutaisi in the south and east.

Russia. Golovinka, EJ 34 (det. anat.; CLA); Sochi, EJ 52 (det. anat.; SMF 278694; ZIRAN).

Georgia. Zhove Kvara valley near Gagra, EH99 (det. anat.; IZPAN); rocks at the shore W of Gagra, EH99 ( CLA); bank of the Malaya Khosta, EJ 72 ( ZMUM 8421, 19104); Kheivani (5 Yermolovka), EJ 80 ( ZMUM 8451); Mys Pitsunda, FH 07 (det. anat.; CLA); Bzyb’ valley, Bzyb’, FH 08 ( CLA); Gagra, FH 09 (det. anat.; CLA; IZPAN; SMF 208905; ZIRAN); Gagryps valley near Gagra, FH 09 (det. anat.; CLA); Pitsunda, FH 18 (det. anat.; CLA; IZPAN; ZMH 2772); Khip’st’a (5 Belaya) valley W of Gudauta, FH 27 ( CLA); Akhali Af ’oni (5 Novyy Afon), FH 47 (paralectotype s of perfrequens: ZIRAN ; ZMH 2773); Akhali Af ’ oni, Monastir (paralectotypes of perfrequens: SMF 69914 (also specimens to O. Boettger 1883); SMF 100051); Akhali Af ’oni, Iberian mountain ( ZIRAN); Gumista valley near Sukhumi, FH 66 ( CLA); Sukhumi, FH 66 ( SMF 69906, 69923, specimens to O. Boettger 1884); Kelasuri valley NE of Sukhumi, FH 76 ( CLA); headwaters of the Macara NE of Sukhumi, FH 76 ( CLA); Bzyb’ valley, Goluboye Ozero, FJ10 ( CLA); middle Bzyb’ valley (above branch towards Gega valley), FJ20 ( CLA); Qulevi (5 Redut-Kale), GG18 (lectotype of obscura (design. nov.): ZMZ 545882a, measurements: D5 11.8 mm, H5 8.9 mm; paralectotypes: ZMZ 505716, 545882b); Batumi, mountain with vantage point, GG21 (det. anat.; CLA, specimens to Clauss 1990); Ch’ak’vi, tea-plantation, GG22 (det. anat.; CLA, specimens to Clauss 1990); Ts’ikhisdziri, ruins, GG22 ( CLA, specimens to Clauss 1990); P’ot’i (Kulumbar), GG27 ( ZMUM 8449); Kobuleti, GG33 ( Hudec and Lezhawa 1969); Och’amch’ ire, GH03 ( Hudec and Lezhawa 1969); Senaki (5 Tskhakaya), KM 58 ( ZMUM 8418); Nakalakevi, KM 69 ( ZIRAN); Gordi, KN90 (lectotype of minor (design. nov.): SMF 69922, measurements: D 5 7.8 mm, H5 5.4 mm); national park Sat’ap’liis Nakrdzali, LM08 ( ZMUM 8427); Kutaisi, LM18 (lectotype of perfrequens (design. Hausdorf in press): SMF 6572 (also specimen to O. Boettger 1881); paralectotypes of perfrequens: SMF 6573, 6574 and 100015 (also specimens to O. Boettger 1881); SMF 69903, 287038; ZIRAN; ZMH 2774; ZMZ 505713, 545884 specimens to Mousson 1863); Gelat’i near Kutaisi, LM18 (det. anat.; CLA).