Eurynassa Thomson, 1864

Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong & Ślipiński, Adam, 2020, A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae), Annales Zoologici 70 (1), pp. 33-96 : 56-57

publication ID	https://doi.org/ 10.3161/00034541ANZ2020.70.1.003
DOI	https://doi.org/10.5281/zenodo.3795846
persistent identifier	https://treatment.plazi.org/id/03ED87B7-CB63-1B36-F96B-F97BFAA9FDDA
treatment provided by	Felipe
scientific name	Eurynassa Thomson, 1864
status

Treatment

Eurynassa Thomson, 1864 View in CoL

( Figs. 8 View Figure 8 , 14H, 15K View Figure 15 , 17 View Figure 17 C–D)

Eurynassa Thomson, 1864: 303 View in CoL . Type species Eurynassa servillei Thomson, 1864 View in CoL , by original designation.

Eurynassa: Lacordaire, 1868: 111 View in CoL ; Lameere, 1903a: 11.

Diagnostic combination (Male). Large beetles, 30–60 mm long. Dorsum usually with dark brown to black head and prothorax, and yellowish- or reddishbrown elytra. Sexually dimorphic dense punctures present on most of the pronotum, scutellum, all thoracic sterna and at least part of abdominal ventrites. Head narrower than prothorax; frontoclypeal suture arcuate to broadly angulate; median groove complete. Antennal tubercles prominent, rounded at apex. Eyes moderately large, weakly emarginate near antennal insertion, relatively distant dorsally. Mandibles about half of head capsule length, not sexually dimorphic, broad at base, not bent towards ventral side, and unidentate apically. Antenna extend to the middle of elytra. Scape about 1/3 of head capsule length, posteriorly usually extending to posterior margin of eye, gradually expanded; antennomere 3 shorter than scape, as long as or longer than antennomere 4. Mentum not fused to sub-mentum. Terminal palpomere of maxillary and labial palps elongate oval or weakly expanded, apex rounded. Prothorax transverse with lateral carina distinct, regularly dentate; anterior and posterior margins with complete beads, posterior angles relatively blunt. Pronotal disc uneven with weakly elevated smooth and coarsely punctate callosities on densely punctate, relatively dull background. Prosternal process subparallel, extending beyond procoxae, rounded apically. Elytral surfaces coriaceous or finely punctate; inner apical angle with sharp spine. Legs relatively long and strong; rows of or at least few spines present on femora and tibiae. Protibia surface rough and densely setose apically, with pair of subequal spurs; tarsomere 3 moderately lobed, tarsomere 5 shorter than tarsomeres 1 and 2 combined.

Description. Male. Length 30–60 mm. Head and prothorax usually dark brown to black, elytra often reddish or yellowish brown. Head approximately as long as wide, narrower than pronotum. Mandibles about half of head capsule length, weakly curved inwards, without sexual dimorphism; each mandible wedge-shaped, very thick and strong on lateral side and near the base, unidentate at apex, with two or three teeth on incisor edge; dorsal surface with distinct carinae in E. servillei and E. tuberculicollis . Labrum weakly concave near apex, anterior margin weakly pointed with very dense setae; labrum separated from clypeus by a less sclerotised deep groove; dorsal surface of clypeus almost glabrous. Frontoclypeal suture arcuate to broadly angulate; median groove distinct and complete. Antennal tubercles relatively prominent, apex rounded. Antenna 11-segmented, filiform, extending to the middle of elytra; scape moderately long and gradually expanded, reaching to or slightly beyond posterior margin of eye; antennomere 3 shorter than scape, as long as or slightly longer than antennomere 4; antennomeres 5–11 with rather blunt posterior angles. Eyes moderately large, transverse, coarsely facetted, slightly emarginate near antennal foramen; relatively distant dorsally. Submentum curved at apex, mentum well exposed. Terminal palpomere of maxillary and labial palps elongate oval to weakly expanded, apex rounded. Pronotum transverse, usually sub rectangular, weakly trapezoid in E. stigmosa , with posterior margin slightly broader than anterior margin; complete bead present on both anteri- or and posterior margins; lateral margin distinct with regular sharp teeth, with weak teeth in E. stigmosa . Disc surface very densely punctate and matt except for shiny, relatively smooth islands consisting of two triangular admedian areas, two smaller lateral areas and transverse area in front of the posterior margin. Prosternum covered with fine and deep punctures; hypomeron well-defined, slightly broader than prosternal process. Prosternal process parallel sided, extending beyond procoxae towards mesoventrite, rounded apically. Mesoventral process sub-parallel and emarginate apically. Ventral side of thorax and coxae with dense golden hairs. Metaventrite with shiny arrowshaped area medially, finely and densely setose elsewhere. Metanepisternum weakly constricted at base. Scutellum rounded apically; surface with very fine, dense punctures. Elytral surfaces relatively shiny, covered with irregular coriaceous sculpture or very fine punctures; without weak traces of venation; elytral apices rounded with sharp sutural spine; epipleuron complete, narrowing apically. Legs very strong; femur sub-parallel or weakly constricted near both ends, surface rough, rows of spines present on ventral side; tibia expanded towards apex, with spines along outer margin, bearing sensory setae on ventral side apically; each tibia with distinct apical projection and a pair of subequal spurs; tarsi strong; tarsomere 3 moderately lobed, tarsomere 5 shorter than tarsomeres 1 and 2 combined. Abdomen with ventrites 1–5 densely punctate basally with smooth and appearing less sclerotised apical part; ventral surface almost glabrous, only bearing long setae along the edge, and very densely setose on the apex of ventrite 5. Male genitalia ( Fig. 14H View Figure 14 ). Tegmen longer than sternite VII; parameres relatively short, about 0.13 times length of entire tegmen, round- ed apically with triangular projections at the base; penis as long as tegmen, dorsal apex truncate while ventral apex narrowly rounded.

Female. Body usually more elongate than male, antennae usually extending to elytra humeral angle or just reach the middle of elytra. Pronotal disc entirely shiny, surfaces irregular with smooth central area and coarsely punctate lateral area; fine and dense punctures on thorax and abdomen absent. Legs much slenderer than in male. Ovipositor ( Fig. 15K View Figure 15 ) long, apical sclerosed part about 2/3 length of baculus, distal gonocoxites usually slender, relatively short and stout in E. stigmosa , apex rounded in most species, but weakly truncate in E. stigmosa ; stylus small, inserted laterally and moderately distant to gonocoxite apex, weakly expanded apically.

Remarks. Eurynassa can be distinguished from the remaining genera of the Australian Macrotomini by having a short antennal scape, usually posteriorly extending to the middle of eye, antennomere 3 shorter than the scape; lateral margin of the pronotum moderately serrate; femur and tibia at least with some spines along external margins. Males always with distinguishable fine sexual punctures on ventral surface.