Veronica teucrium Linnaeus (1762: 16)

Rojas-Andrés, Blanca M. & Martínez-Ortega, Montserrat, 2016, Taxonomic revision of Veronica subsection Pentasepalae (Veronica, Plantaginaceae sensu APG III), Phytotaxa 285 (1), pp. 448-450 : 448-450

16. Veronica teucrium Linnaeus (1762: 16) . ( Fig. 25 )

Stems (15) 30–70 (120) cm long, decumbent to erect, covered by long and short eglandular hairs (0.2) 0.5–0.7 (1.3) mm long, the shortest ones incurvate, appressed and antrorse, generally concentrated in 2 opposite lines along the stem; apical shoot bearing 1–7 pairs of leaves. Leaves (3) 5–55 (70) × (2) 7–25 (40) mm, the basal and medium ones similar, narrowly elliptic or lanceolate to widely ovate or even cordate-deltate, subcordate, truncate or rounded at the base, occasionally subamplexicaul, crenate-serrate to incised-dentate (rarely subentire), subglabrous or pilose, sparsely covered by hairs (up to 0.15–0.9 mm long) on the adaxial face (hairs at least on the primary vein and along the margin) and more densely pilose on the abaxial face, sessile or rarely subsessile, usually non-revolute; apical shoot leaves opposite, ± widely ovate, ovate or elliptic, crenate to serrate, rarely subentire but non-revolute. Racemes axillary, opposite, bearing 15–60 (110) flowers, loosely to densely arranged; peduncles 1.7–11 cm long, usually glabrous on the lower part and covered by the same type of indument as that of the stem on the upper part; bracts (2) 4–8 (14) mm long, linear-lanceolate or narrowly elliptic, entire, usually subglabrous or with crooked hairs on the primary vein and along the margin; pedicels (1) 3–7 (14) mm long. Calyx (1.5) 4–5 (6.5) mm long; (4) 5 sepals, linear-lanceolate or narrowly elliptic, almost always shorter than the capsule, glabrous or with ± crooked eglandular hairs. Corolla (9) 12–17 (18) mm in diameter, light or intense blue to blue-violet. Capsule (3) 4–5.5 (6) × (2.5) 3.5–5 (5.5) mm, usually longer than wide, exceptionally wider than long, widely obovoid or obcordate, ± rounded at the base, usually emarginate, sinus up to 0.7 (0.9) mm depth, glabrous or pilose, sparsely covered by eglandular hairs. Style 5–6 mm long. Seeds 1.2–1.8 × 1.0– 1.4 mm, ca. 2–18 per capsule ( Kulpa 1968).

Chromosome Number: —2 n = 64.

Habitat: —Meadows, edge forests, shrublands, pastures, preferentially calcicolous; 0–1,900 (3,400) m.

Distribution: —C and E Europe and Asia (Siberia). Austria, Belarus, Belgium, NW Bulgaria, Czech Republic, N and E France, Germany, Hungary, N Italy, Lithuania, Poland, Romania, Russia, Serbia, Slovakia, N Slovenia, Switzerland and Ukraine. According to Borissova (1955) extending also to W Siberia ( Fig. 26 ).

Phenology: —Flowering from (April) May to July (August).

Notes: —Several infraspecific taxa have been described and subordinated under V. teucrium throughout history ( Rojas-Andrés et al. 2016). Some of them, such as V. crinita , have recently been shown to be monophyletic independent taxonomic entities ( Rojas-Andrés et al. 2015). After a detailed study of the morphological variation of V. teucrium (herbarium specimens and living populations), we consider that only two infraspecific entities could be recognized at the most at the varietal level: V. teucrium var. teucrium and V. teucrium var. angustifolia . They differ mainly in characters that according to our field observations might be related to sun exposure and soil humidity. Veronica teucrium var. teucrium has generally erect and robust stems (> 35 cm), leaves (20) 30–55 (70) × (3) 15– 25 (40) mm, widely ovate or very widely ovate to cordate-deltate. Veronica teucrium var. angustifolia has decumbent to ascending and ± flexuous stems (<30–35 cm), leaves (3) 5–25 (30) × (2) 7–20 (25) mm, narrowly elliptic or lanceolate, rarely ovate or elliptic. These two varieties have roughly different distribution areas (although there are even populations composed of individuals of different morphologies); whereas var. angustifolia occurs mainly in Western Europe (C and E France, W Switzerland and N Italy), var. teucrium extends widely towards the east. Although the epithet “vahlii ” [ Veronica teucrium [II] vahlii Gaudin (1828: 22) ; V. austriaca subsp. vahlii (Gaudin) D.A.Webb in Walters & Webb (1972: 267)] has frequently been used to designate those plants corresponding with V. teucrium var. angustifolia , (e.g., Hartl 1966; Walters & Webb 1972), the epithet “angustifolia ” prevails at the varietal level (Martínez-Ortega & Rico, 1999).

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V. teucrium var. angustifolia has frequently been confused with V. orsiniana in SE France. See note under V. orsiniana for details.

Specimens Examined: — AUSTRIA. Carinthia: Ferlach, Karawanten, 14 June 1937, Scheneider s.n. (WU); Gailtal, am Fuss des Dobratsch auf einem trokenen Hügel auf einer Weide in der Schütt gg. Federaun, 5 & 11 June 1972, Melzer s.n. (WU); Hermagor: in Gebüschen bei Passriach, 29 June 1990, Krebs s.n. (E); Rabenstein bei St. Paul, kalk, buschiger Hang unterhalb der Ruine, ca. 680 m, 18 June 1983, Polatschek s.n. (W); S-Kärnten, Rosental, am Bahnkörper zwischen Ferlach uns Unterbegen, 20 June 1918, Pokorny s.n. (W); Sankt Paul im Lavanttal, Rabenstein Burgruine, borde de prado sobre calizas, 656 m, 46º41'23.7''N, 14º52'27.0''E, 25 July 2011, X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés BR191 (SALA149406); Villach, Oberschütt, en la salida del pueblo hacia Unterschütt, borde bosque y prado sobre calizas, 532 m, 46º34'03.5''N, 13º45'39.5''E, 26 July 2011, X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés MO6042-A (SALA149418); Villach, Oberschütt, en la salida del pueblo hacia Unterschütt, borde bosque y prado sobre calizas, 532 m, 46º34'03.5''N, 13º45'39.5''E, 26 July 2011, X. Giráldez , M.M. Martínez Ortega & B. Rojas Andrés MO6042-B (SALA149419); Laxenburg: Wiesen bei Laxenburg nächst Wien, 18 June, Halacsy s.n. (WU); Lower Austria: Südostliche des Leopoldsburges bei Wien, 7 June 1902, Keller s.n. (W); Kahlenberg bei Wien, 5 June 1866, unknown collector (W); Vienna, between Kalenberg and Leopoldsberg, north of Vienna, grassy clearings in wood, local, September 1982, Bowen s.n. (RNG); Wienerwald, an grasig-(illegible word) Orten zwischen (illegible group of words), 18 June 1924, Korb s.n. (W); Niederösterreichische Kalkalpen, Türnitzer Höger bei Hohenberg, im oberen Dachsgraben, Kalk, in einer Wiesenhecke, ca. 600 m, 16 July 1978, Polatschek s.n. (W); zwischen Sommerein und Mannersdorf, 3 June 1963, Krendl s.n. (W); Wienerwald, Gebüschrand bei Alland, 20 June 1964, M. A. Fischer s.n. (WU); Huundsheimer Berg, W-Seite, Böschung am Rande eines Weingartens, bei Deutsh-Altenburg, 23 May 1968, M. A. Fischer s.n. (WU); in graminosis ad Laxenburg prope Vindobonam, 200 m, Woloszczak s.n. (B, BM, PR); in pratis planitiei ad Laxenburg prope Viennam, June 1884, E. Halacsy s.n. (MA112458); Böhk, Weg von Hinternberg nach Lanzendorf, 12 July 1975, Kaisergruber s.n. (WU); Tullner Feld, ca. 5–6 Km S von Kirchberg am Wagram, 24 July 1933, Fischer s.n. (WU); westlich von Neustadt, auf einer Wiese bei Weikersdorf uns auf der Zweierwiese bei Fischau, 1862, Sonklar s.n. (WU); Thermenlinie, Graslauterriegel 1, 5 km SW Gumpoldskirchen, trockenrasen,

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Gebüschsäme, 280–320 m, 6 July 1993, Till s.n. (WU); Laxenburg, nächst Wien, 2 June 1878, Heine s.n. (W); Moosbrunn, 31 May 1908, Zemann s.n. (WU); Austria inferior, in graminosis ad Laxenburg prope Vindobonam, 200 m, unknown date, Woloszak s.n. (B); (illegible word), 20 June 1881, Franz Ostermeyer s.n. (B100327733); Puchberg, 47º47'N, 15º54'E, 10 June 2000, D. Albach DA285 (WU0023357); entre Pottenstein y Weissenbach a. d. T., 28 June 2001, M. Martínez Ortega & G. Fischer MO1451 (SALA124602); Salzburg: ¿Mönchsberg?, 500 m, Eysn s.n. (WU); Rocks ???berg, Salzburg, 20 June 65, unknown collector (K); Styria: Judenburg, Karnerkreuz ob Reifling, Kalk, 900 m, 25 June 1932, Schallauf s.n. (B100327729, W, WU); Pegga?, 1 July 1855, Pittoni s.n. (K); Weiz, Eibisberg, 10 September 1954, Neumann s.n. (W); Tirol: In Tiroliae australis collinis, pr. Nickolsdorf, solo calc., 2000 ft., 31 May 1865, Auserdorfer s.n. (K); Lienz, in collib. ad dumenta, June 1870, Garder s.n. (K); Oberinntal, linkes Inn-Ufer zwischen Mötz und Magerbach buschiger Hang beim Birgele, kalk, 700 m, 9–19 July 1975, Polatschek s.n. (BM, W); Ötztaler Alpen, am Abstieg Stables-Nauders, silikate, in einer Rosenhecke, ca. 1500 m, 15 July & 9 August 1979, Polatschek s.n. (WU); Karwendel, Innsbruck, Höttinger Graben, buschigsteiniger Hang, c. 900 m, 15 August 1980, Polatschek s.n. (BM); ¿Tiesser? Thal, Tirol, 46º47'0.13"N, 12º39'57.86"E, June 1922, unknown collector (BM); Osttirol, Oberes Drautal, Wiesenhang am W-Fuss der Trögerwand bei Nörsach, ca. 700 m, 27 August 1973, Polatschek s.n. (W, WU); Innsbruck, Hungerburg, Höttinger Graben, entre Gramart y Höttinger Bild, borde de bosque, junto al camino, 846 m, 47º17'10.9''N, 11º22'22.1''E, 13 July 2011, X. Giráldez & B. Rojas Andrés BR170 (SALA149401); Oberinntal, am Weg Pfunds-Gh “Eggele” (Frudiger Kopf), silikat, 1100 m, 15 July & 9 July 1979, Polatschek s.n. (WU); Nordtirol, Innsbruck, Mühlau- Klamm, Kalk, 27 July 1895, Fischer s.n. (W); Fliess cerca de Landeck, prados sobre silicatos, 1044 m, 47º07'06.7''N, 10º38'28.4''E, 14 July 2011, X. Giráldez & B. Rojas Andrés BR171 (SALA149402); Upper Austria: Pfenningberg bei Linz, unter Gebüsch, 24 June 83, Strobl s.n. (K); bei Obernberg, 8 June 1909, Ronniger s.n. (W); Steyr. Hänge a. d. Steyr, June 52, Baschant s.n. (B100327728); Schwertberg. Oberöstr., June 1874, Keck s.n. (B); Trockene Wiesen b. Steyr., June 48, Baschant s.n. (B100327726); Steyr. Hügel a. d. Ems, June 52, Baschant s.n. (B100327725); Tirolis, collina prope Linz, 20 June 1864, unknown collector (BM); Schwetherg, Oberöstr., June 1874, Keck s.n. (B); Hügel, Ufer und Gräben bei Linz ¿a/Ahn?, May 1882, Melsheimer s.n. (K); Voralberg: Klostertal, Dalaas-Mutten bis Innerbraz, 800– 700 m, 15 June & 3 July 1971, Polatschek s.n. (WU); Walgau, ruine Ramschwag b. Nenzing Burghühgel u. angrenzendes Gebiet Wiesen und Waldgürtel um die Ruine, 18 June 1971, Seipka s.n. (W); Vienna: Roter Berg, Wien, 6 June 1908, Watzl s.n. (WU); Wien, 2 July 1899, Hübl s.n. (BM); Wien, Prater, 26 May 1877, Müllner s.n. (W); Perchtoldsdorf, Kalksburg, 28 June 2001, M. Martínez Ortega & G. Fischer MO1453 (SALA124565); Mödling, Eichkogel, 200–250 m, 13 July 2000, D. Albach, L. Delgado , M. Martínez Ortega, E. Rico & J. A. Sánchez Agudo MO 965 & LD533 (SALA118381); Kalksburg, en la subida desde la calle "In der Klausen", prados herbosos sobre calizas, 322 m, 48º08'36.7''N, 16º14'57.2''E, 24 July 2011, X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés MO6039 (SALA149417); unknown or dubious state: (illegible locality), June 52, Baschant s.n. (B); St. Peter, in pratis, 29 June 1902, unknown collector (PR); Kaenigsberg: Moosbude, 7 June 72, Ralnitz s.n. (K); Austria, 22 June 11, unknown collector (E).

BELARUS. Grodno: Ad Nowogródek ejusd. distr. Na miejscach trawiastych, 14 July 1893, W . Dybowski s.n. ( B100327791 , BM) .

BELGIUM. Hainaut: An Hecken bei Obourg , 5 June 1864, Martinis s.n. ( B, K) ; Obourg, pelouses des bois, calcaire, 9 June & 25 July 1866, Martinis s.n. ( BM, K) ; Obourg province de Hainart, 4 June 1864, ¿Martim? s.n. ( B100327715 ) ; Vlaams-Brabant: Tirlemont, June 1861, illegible collector ( WU) ; unknown province: Lejeune , unknown collector ( BM) .

BULGARIA: Montana: Balkan foothills, Montana, S of Prevala , in grassy place under trees along the street, 43º28'N, 22º51'E, 21 June 2001, D. Albach DA544 ( SALA121498 ); Pernik: Znepole region , Tran , east of Tran , rich meadow, 790 m, 42º50'16''N, 22º41'27''E, 15 June 2001, D. Albach DA501 ( SALA124597 ); Pernik: alrededores de Tran, en la carretera de Pernik a Tran, a 3.2 Km en línea recta al Este de la ciudad, 757 m, 42º50'16.6''N, 22º41'33.8''E, 14 June 2009, X. Giráldez , M. M . Martínez Ortega, B. Rojas Andrés & M. Santos Vicente MO4574 ( SALA149044 ) ; Sofia-city: Sofia, en la subida a Vitosha , junto a la carretera, 1021 m, 42º37'39.4''N, 23º17'26.8''E, 15 June 2009, X. Giráldez , M. M . Martínez Ortega, B. Rojas Andrés & M. Santos Vicente BR19 ( SALA149405 ); Mt. Vitosa, 22 July 1933, Thompson s.n. ( K); Vitosa Mountain , Simeonovo , base station of gondola, meadow, 750 m, 42º36'56''N, 23º20'51''E, 17 June 2001, D. Albach DA526 ( SALA124578 ); Vitosa Mountain , Ljulimin, meadow beneath ski slope, meadow, 20 June 2001, D. Albach DA530 ( SALA121494 ) ; Sofia-province: Bulgaria occid., Montes Vracansky kras, in pratis montanis carsticis ad viam publ. inter oppidum

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Phytotaxa 285 (1) © 2016 Magnolia Press • 81 Vraca et cavernam Ledenika, 1000 m, 23 June 1986, Fr. Cernoch s.n. ( SEV121494 ); Vratsa: In pratis Bulgaria occidentalis prope urbem Vraza, 23 June 1949, B . Kitanoff s.n. ( MA182270 ); unknown province: In graminosis siccis ad Pesteza, 1810, Urumoff s.n. ( WU) .

CROATIA. Primorje-Gorski: Istrien, südwestlich exponierten Abhang oberhalb der Mulde Mlaka zwischen Klana und Hermsburg [=Gomance], kalk, 700–900 m, Grinzbergen & Janchen s.n. ( WU) .

CULTIVATED. E horto Berolinense, 1817, Bergemann s.n. (B100327891); Ex horto bot. Jenensi, unknown collector (B100327889); Jardín Botánico de la Universidad Babes-Bolyai (Cluj-Napoca), cultivada, 29 June 2000, X. Giráldez , M. Martínez Ortega & J. A. Sánchez Agudo MO899 (SALA121511); Jardín Botánico de la Universidad Babes-Bolyai (Cluj-Napoca), cultivada, 29 June 2000, X. Giráldez , M. Martínez Ortega & J. A. Sánchez Agudo MO901 (SALA121510).

CZECH REPUBLIC. Central Bohemia: In rupibus calcareis in nemore in valle rivuli Studený prope opp. Dobrichovice , 280 m, 30 June 1937, Stearn s.n. ( K) ; Olomouc: Olmütz [= Olomouc], Steppen abh.?, ¿ Sckubolus?, May 1934, Taus s.n. ( MA112449 ); Olmültz [= Olomouc], (two illegible words), June 1935, Laus s.n. ( B100327751 ); Prague: Bei Prag, unknown date, Bergler s.n. ( W) ; South Moravia: Entre Mikulov y Klentnice , prados calizos secos, en lugares con vegetación panónica, 24 June 2001, M . Martínez Ortega MO1446 ( SALA124568 ); Knezdub: Meadows in the Certoryje National Natural Reserve , ca 3.5 km SSE of the village, 410 m, 48º51'29.0"N, 17º24'51.0"E, 2 June 2012, J . Tesitel s.n. ( SALA148182 ) ; South Moravia: Entre Klentnice y Mikulov, Tabulová , prados herbosos, 354 m, 48º50'09.8''N, 16º38'19.7''E, 23 July 2011, X. Giráldez , M. M . Martínez Ortega & B . Rojas Andrés BR188 ( SALA149404 ) ; Zlín: Moravia orient. Valasske Meziricí, Choryn, in monte Stráz , 370 m, June 1934, Rican s.n. ( K) ; Böhmen. Prag. Gebüsche bei Podbaba, Schiefer , 260 m, 9 June 1886, J . Freyn s.n. ( B100327750 ) .

FRANCE. Ain: On exposed mountain slope, Challex, Jura mountains, Savoy, 14 June 1936, Moldenke & Moldenke s.n. (BM); Alpes-Maritimes: Caussols, sur le plateau de Caussols au sud de la route D.12, à mi-chemin de l’Embut et du col de l’Ecre, praire rocailleuse sur terrain calcaire, 1100 m, 1 June 1990, B. de Retz 89961 (BIO14481, SALA 93486, VIT37163); Grasse, cerca de Caussols, en la carretera D12 hacia el col de l'Êcre, prados sobre calizas, 1080 m, 43º44'33.5''N, 06º55'03.6''E, 10 June 2012, X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés BR211 (SALA149409); Alsace: Weissenburg, Alsatia, 5 June 1860, F. Schultz s.n. (B, PRC); Prope Weissenburg, Alsatia, 5 June 1860, Schultz s.n. (WU); Prope Weissenburg, Alsatia, Schultz s.n. (W); Bas Rhin: Strasburg, Buchinger s.n. (B); In ¿tumulis celticis? ad Benfeld Alsatiae, Buchiger s.n. (K); ¿Kauslerberg? bei Bergheim, Ober-Elsass, 5 June 1931, Issler s.n. (W); Prés secs près de la Moder à Haguenau, 16 June 1847, Billot fils. s.n. (BM); Im carices humilis der (illegible group of words) zwischen ¿Wettolheime? und ¿Winzerheim?, Ober-Elsass, 26 June 1910, Issler s.n. (W); Mundolsheim près de Strasbourg, June 1852, Diomede s.n. (K); Weissenburg, Alsatia, 14 June 1869, F. Schultz s.n. (MA112467); a Mundolsheim près de Strasbourg (Bas-Rhin), June 1852, Billot s.n. (BM, K, PRC); Strasburg (two illegible words), 20 April 1945, Krumbholz s.n. (B); Calvados: Dunes a Merville près l'embouchure de l'Orne, 27 August 1841, unknown collector (P00208683); Calr. Dunes de Deauville, 4 June 1847, DD. s.n. (P00208675); Doubs: SW of Remont between ¿Morteau? and ¿Montbenort?, on steep slope by road with Bromus erectus, Poterium, Ophrys , Listera ovata , 770 m, 15 June 1961, illegible collector (K); Essone: Valpuiseaux, La Lieu, prados semiáridos dominados por Brachypodium al lado de pinar, 90 m, 48º23'32''N, 2º18'33"E, 4 July 2014, N. Padilla García & A. Torres NPG3bis (SALA154238); Valpuiseaux, La Lieu, prados semiáridos dominados por Brachypodium al lado de pinar, 110 m, 24 May 2014, N. Padilla García & V. di Donato NPG3 (SALA154237); Eure-et-Loir: S. of Thiville, SE os Chateaudun, in rough grassland with Euphorbia, Erodium , Bellis perennis , etc., 10 May 1968, illegible collector (K); Pelouses sèches, coteaux xc. A. C. Châteaux dun, 17 June 1847, Juillard s.n. (P00208679); Châteaudun, Thiville, prados en mancha relativamente conservada de orla de bosque-espinar con Juniperus , Quercus, Ligustrum , Rosa , etc., 127 m, 48º00'09.3''N, 01º23'21.5''E, 5 June 2011, A. Abad de Blas, N. López González , M. M. Martínez Ortega & B. Rojas Andrés MO6022 (SALA149413); Hautes-Alpes: Hautes Alpes, Mt. Chabriere, p. Chorges, 14 July 1865, Borel s.n. (K); Chorges, cerca de Gap. Monte Chabrières, en la subida desde Les Andrieux, borde de camino, 1393 m, 44º34'35.3''N, 06º16'35.3''E, 10 July 2011, X. Giráldez , M.M. Martínez Ortega & B. Rojas Andrés BR167 (SALA149398); Gap, Col du Noyer, entre Le Noyer y L'Enclus, prados húmedos sobre calizas, 1671 m, 44º41'34.7''N, 05º59'13.2''E, 14 June 2012, X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés BR217 (SALA149410); Haute-Savoie: Alta Sabaudia, Belleveaux, Chablais , 4 August 1859, Puget s.n. (BM); Gèneve, Salève, in pascuis siccis, 1500–2000’, June 1839, Nägeli s.n. (E); Salève, 4000 ft., 8 July 1888, Clarke s.n. (K);

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Mont Salève, Jura, 13 July 1856, Ducomun s.n. (BM, PRC); (illegible locality), 6 July 76, ???eseglise s.n. (E); Monte Salève cerca de Ginebra, en los prados calizos de la cresta de la cumbre, 1302 m, 46º08'06.4''N, 06º10'57.5''E, 10 July 2011, X. Giráldez , M.M. Martínez Ortega & B. Rojas Andrés BR168 (SALA149399); Haut-Rhin: Rufacher Berg, Ober-Elsass, 28 May 1932, Issler s.n. (W); Auf Rheinkies bei Fisch??? bei ¿Blosheim?, Ober Elsass, 7 July 1907, Issler s.n. (W); Bergheim, prados junto al cementerio alemán en Grasberg, prados secos, 338 m, 48º12'45.6''N, 07º21'10.8''E, 11 June 2011, A. Abad de Blas, N. López González , M.M. Martínez Ortega & B. Rojas Andrés MO6029 (SALA149415); Isère: Anjou, Chalonnes, etc., T. Bastard s.n. (P00208674); Montes de Grenoble, June 1839, Haden s.n. (E); Maine-et-Loire: pelouses calc. de Liré, 14 May 1870, unknown collector (BM); Montjeaun, rochers du Chateaup???, 26 May 1878, Borel s.n. (K); Saumur, ¿champiny?, 10 May 1864, illegible collector (BM); Moselle: Ippling, Wiese in der Bachaue S des Ortes (südlich Wald), 27 May 1995, R. Hand & Steffen Caspari 87 (B100327740); Nièvre: Varzy, 20 May 1863, Pin s.n. (K); Varzy, prados, 253 m, 47º21'44.3''N, 03º23'48.6''E, 4 June 2011, A. Abad de Blas, N. López González , M. M. Martínez Ortega & B. Rojas Andrés BR157 (SALA149397); Pas de Calais: sand dunes by the sea, 15 June 68, Verdcourt s.n. (K); Roadside near Ambleteuse, Pas de Calais, 19 May 78, unknown collector (K); Rhône: Vallon de Beaunant, près de Lyon, Rhône, 12 & 28 May 1871, Boullu s.n. (K, WU); Arnas, May 1898, Gandoger s.n. (W); Sarthe: La Mans, champs près de la ¿Boviniere?, 29 April 1934, Gentil s.n. (BM); Savoie: roadside, Val d’Isère, Savoie, 6 July 1976, unknown collector (E); (illegible group of words), 13 August 1932, Heard s.n. (BM); unknown locality, 9 June 1892, Pin s.n. (K); Seine-et-Marne: Sous de Cessens, Exemplaire cueilli a Fontaineblau par M. Desfontaines, et donné par lui en 1815 sous le nom de V. teucrium , herb. J. Gay s.n. (K); Somme: very fine and abundant on chalky (illegible word) near Bovelles, 9 June 1956, Lousley s.n. (BM); Ruine Boves bei Amiens, March 1871, Wirtgen s.n. (BHU); Val d'Oise: Route des Cretes, above La Roche Guyon, roadside on chalk, 11 May 67, Hunt s.n. (K); Yvelines: Bois de Beynes nr. Beynes c. 13 Km WNW of Versailles, at side of path beside wood on chalky clayey soil, Quercus pubescens , 11 May 1964, Hunt s.n. (K); unknown or dubious department: Doubs, Monttier, Chavelle de St. Foi, 5 June 1930, Williams s.n. (BM); Durbon, Col du Pensa au dessus de la Chartreuse, 27 July 1866, Borel s.n. (K); Peronne, 14 June 1782, Cavanilles s.n. (MA112445).

GERMANY. Baden-Württemberg: Alb. Hofbühl bei Metzingen, Steppenheide , 16 June 49, Ema Walter s.n. ( B100327902 ); Alb. Randecker Maar , Felsen im Gebüsch , 8 June 47, E . Walter & E . Mayer s.n. ( B100327901 ); Entre Fridingen an der Donau y Beuron , prados junto a la carretera, 4 May 2001 / 29 July 2001, M . Martínez Ortega MO1083 & MO1506 ( SALA120889 & 121493); Freiburg, June , unknown collector ( B); Heidelberg, (illegible word), June 1903, Pringsheim s.n. ( PRC) ; Lichter Hang mit wenig Gebüsch zwischen Unterkochen und Ebnat, 12 June 1948, Ema Walter s.n. ( B100327904 ); Schwäbische Alb , Hausen im Tal , Geröllhang , Hochstaudenflur , 26 September 85, Ketelhut-Hoffman s.n. ( B); Ulm, entre Bermaringen y Asch, prados y repisas de rocas calizas, 9 May 2001, M . Martínez Ortega MO1096 ( SALA120888 & 124577); Ulm , entre Bermaringen y Asch, prados y repisas de rocas calizas, 29 July 2001, M . Martínez Ortega MO1505 ( SALA124570 ); Wall od vineyard, near Riegel , Kaiserstuhl , 15 July 1906, Groves s.n. ( BM); Württemberg, Urach, Jura, 30 July 1953, Woolhouse s.n. ( RNG); Wollmatinger Ried, Diechselrain (mager), 4 June 1981, R . Heckmann & U . Rudolph s.n. ( B100327905 ); Bavaria: Augsburg: von (three illegible words), 09 June 1893, Weinhart s.n. ( B100327855 ); Fränkischer Jura : Treuchtlingen , Wiese mit Gebüsch auf südexponiertem Hang des Weinberges , 420 m., 6 June 66, H . Scholz & P . Hiepko 125 ( B100327789 ); Fränkische Schweiz, bei Bronn südöstlich Pottenstein , trockenrasen am Waldrand , 18 September 1960, Wagenitz s.n. ( B); Fränk-Schweiz, Wissenttal, 24 July 1916, Bornmüller s.n. ( B); Geisenbrunn bei München, 20 June 1914, Obern ??? s.n. ( BM); Jura, 400 m, 21 June 1917, Fr. Kudorfer s.n. ( PR); Obere Donau, Falkenstein, 25 June 02, Ernst H. L . Krause s.n. ( B100327814 ); Pommelsbrunn Jura, 49º30'10.61"N, 11º30'45.37"E, August 1907, Meister s.n. ( PR); Riedenburg , Jura , 400 m, 21 June 1917, Fr. Kudorfer s.n. ( MA185322 ); Stadtsteinach, buschige Abhänge, Raine bei Untersteinach, vorzüglich auf Muschelkalk, 320–340 m, 8 June 1918, W . Putchler s.n. ( MA185323 , PR); Wisenttal i.d. Fränk. Schweiz , 22 July 1880, J . Bornmüller s.n. ( B100327816 ); Wisenttal, bei Stempfermühle , 19 July 1916, J . Bornmüller s.n. ( B100327824 ); Berlin: Bredower Forst, 6 June 1886, Scheppig s.n. ( BM); Brandenburg: Friesack, June 1913, Roman Schulz s.n. ( B100327899 ); Kreis Fürstenwalde: Trockenrasen am Ortsausgang Briesen Richtung Kersdorf , 14 June 1974, I . Heinrich s.n. ( B100327693 ); Lkr. Havelland , S . Elstal (Bra 283), Krautfluren , Strassenrand , 38 m, 52º31'59''N, 12º59'54''E, 30 May 2002, R. & E . Willing s.n. ( B100104498 ); Nauen: In Bredower Forst süd-westlich vom Forsthause , 17 June 1894, V . et R. Schulz s.n. ( B100327898 , B100327900 ); Bötzow, June 1918, Roman Schulz s.n. ( B); Krs. Strasburg: (illegible word); ¿ Bonicke?, 27 June 1945, illegible collector ( B100327819 ); Augermunde: an den westl .

TAXONOMIC REVISION OF VERONICA SUBSECTION PENTASEPALAE

Phytotaxa 285 (1) © 2016 Magnolia Press • 83 Abhängen des Krähenbergs, 16 July 1914, Roman Schulz s.n. (B100327894); Angermünde: zwischen Stolzenhagen am Odertal und dem Forsthaus Buchsmühle an den westlichen Abhängen des Krähenbergs, 16 July 1914, Roman Schulz s.n. (B100327895); Angermünde: Auf den Hügeln beim Forsthaus Buchsmühle nach Stolzenhagen hin., 5 August 1911, Roman Schulz s.n. (B100327699); Krs. Strasburg: Bandelow, ¿Poradies?, 20 June 1945, K s.n. (B100327818); Kreis Nauen: NSG Grosser Jahnberg, Südrand, 20 June 1984, I. Heinrich s.n. (B100327677); Lübben, alte Hirschland, Ziegelgrube an der Luckauer Str., 6 June 49, Bialucha s.n. (B100327675); Mtb. 3452-1 Seelow, Steilhang beim Bahnhof, 27 June 1974, Fritz Hans s.n. (B100327676); Kreis Rathenow: Nennhausen, Waldrand zwischen Dammer Strasse und Kotzen, 30 June 1982, W. Jaschke s.n. (B100327678); Frankfurt a. O.: Buschmühle, 9 June 1895, G. Hirte s.n. (B100327680); Flora der Mark. Templin: (two illegible words); Alt-Placht: (illegible word); Lychen: And der (three illegible words) ¿Stargardter? (illegible word), June 69, ¿G. L. Heiland? s.n. (B100327686); Lychen: Mull an der (three illegible words) Alt-Placht, unknown date, ¿G. L. Heiland? s.n. (B100327685); Nauen bei Berlin, June 1884, J. Bornmüller s.n. (B100327689); Paulinenaue, ¿Jasuberge?, 18 June 39, Joh. Günther s.n. (B100327691); Kreis Seelow: Feldrand nördlich des Krummen Sees südwestlich von Dolgelin, 15 June 1974, U. Müller s.n. (B100327692); Kreis Seelow: Oderhänge nördlich Bahnhof Seelow, 17 June 1974, M. Bässler s.n. (B100327694); Kreis Eisenhüttenstadt: "Steile Wand" bei Lossow, 18 June 1974, D. Benkert s.n. (B100327698); Hesse: Frankfurt a. M., Bagge s.n. (PR); Frankfurt, 1830, Hafsel s.n. (BM); Lkr. Bergstrasse, Gross-Rohrheim, Hammer Aue, Sandwert, bei Fluss-km 457,724, Auenwiese (leicht gedüngte, noch magere Glatthafer-Wiese); vereinzelt, 89 m, 5 June 2003, Karl Peter Buttler & Klaus Hemm K. P. Buttler 33450 (B100122214); Lower Saxony: Braunschweig, coteaux herbeux, June 1890, Krummel s.n. (BM); Mecklenburg-Vorpommern: Datzenberg bei Neubrandenburg, 26 July 1957, U. Dettmann s.n. (B100327809); Rügen, Göhren Nordpehrd, July 35, Bothe s.n. (B100327706); Höft/Mecklenburg, am Reusenplatz, 17 August 1952, A. Buhl 872 (B100327822); North Rhine-Westphalia: Entre Iversheim y Arloff, prados sobre calizas, 258 m, 50º35'29.7''N, 06º46'45.4''E, 9 June 2011, A. Abad de Blas, N. López González , M.M. Martínez Ortega & B. Rojas Andrés MO6025 (SALA149414); Rhenania, Nord-Eifel, Iversheim, sonnige kalktriffen an der Stasse nach Arloff, S. u. SW-Exposition, 270–315 m, 28 May 1930, Andres s.n. (B); Rhineland-Palatinate: an Wegrändern bei Coblenz, June 1865, Writger s.n. (BM); between Leubsdorf and Linz, on alcalline or basic soil, Rubutcher s.n. (K); Pentes del collines exposées au soleil près de Coblence, 20 May & 24 July 1855, Wirtgen s.n. (BM); Flora of Westerwald, between Leubsdorf and Linz, alcalline or basic soil,?ubutcher s.n. (K); env. de Landau, 20 May 1929, unknown collector (B, MA112464); Env. de Landau, 29 May 1929, F. Schultz s.n. (B100327815); Flora der Region Trier, Traben-Trarbach, ca. 150 m. ESE Parkplatz Heller Sass in HTR-Rest. TK 6008/2, 23 May 1998, Ralf Hand 2466a (B100121774); Rheingebiet, St. Goarshausen, June 20, Baschant s.n. (B100327827); Flora der Pfalz, Bezirk Ludwigshafen, neben der Mundenheimer Strasse, Alluvium, ca. 95 m, unknown date, Dr. Hermann Poeverlein s.n. (B100327903); Rheinpreussen, Mittelrhein, Ruine Hammerstein b. Andernach, Thonschiefer, 13 June 94, F. Wirtgen s.n. (B100327801); Trockene Wiesen b. Coblenz, May–June, F. Wirtgen s.n. (B100327803); Wasserliesch, Moselwiesen SW Albachmündung; TK 6205/3, 27 May 1990, Ralf Hand s.n. (B100121564); Palatinat, environs de Landau, 20 June 1929, D’Alleizette s.n. (BM); Kuppe des Hüttbergs bei Nittel-Köllig, May 1996, Hand HA745 (Herb. Hand 745); Saarland: Rheinpreussen, Saargebiet, Merzig, am Wege nach Mondorf, unknown date, ¿Schunen? s.n. (B100327804); Saxony: Coschütz bei Dresden, 1900, Ulich s.n. (B); Dresden, dürrer Hang von Steinbruch, unknown date, Herbert Beger s.n. (B100327643); Bienitz bei Leipzig, June 1845, unknown collector (B100327845); Lommatzsch: ¿belonschte? Hänge im Ketzerbachtal, 3 June 1911, H. Beger s.n. (B100327647); Bienitz bei Leipzig, June 1845, Herbar Andrae s.n. (B100327842); Guttau, Im kontinentalen trockenrasen mit Phleum boehmeri auf dem Eisenberg (Mesobrometum) , 5 July 1960, W. Hempel 2065 (B100327823); Bienitz bei Leipzig, 4 July 1881, (1885 fr), unknown collector (B100327649); Dresden: dürrer Hang des Sanberges gegen den Garnisonsmühlen, steinbruch, unknown date, Herbert Beger s.n. (B100327644); Auf grasigen Anhöhen bei Dresden, July, Sehlmeyer s.n. (B100327884); Hgr. Sachsen: Coschütz bei Dresden, 1900, illegible collector (Herb. Fr. Ulich) (B100327653); Meissen: b. Niederau, 10 June 11, Herbert Beger s.n. (B100327645); Hainewalde, am Spitzberg zwischen Hainewalde und Scheibe, 16 June 1953, A. Buhl 1961 (B100327810); Auf grasigen Anhöhen bei Dresden, July, unknown collector (K); Auf grasigen Anhöhen bei Dresden, July, unknown collector (K); Bieniz bei Leipzig, 4 June 1904, Pringsheim s.n. ( PRC); Bienitz bei Leipzig, 4 July 1881; 1885 fr., J. Bornmüller s.n. (B100327649); Saxony-Anhalt: Südharz, aus Stolberg zu Steigerthal, Gyps, 20 June 90, F. Wirtgen s.n. (B100327800); Waldwegrand (trockner Kalkgerölldamm) bei der Rudelberg-Saale, 15 July 1957, Sigurd FrÖhner 1824 (B100327795); Flora des Harzes, Elbingerode, an der Bocksbergklippen bei Rotenhütte, 10 July 1913,

84 • Phytotaxa 285 (1) © 2016 Magnolia Press

ROJAS-ANDRÉS & MARTÍNEZ-ORTEGA

Roman Schulz s.n. (B100327650); Havelufer bei Schmokenberg, 12 June 1921, Becker s.n. ( B100327700 ); Harz. Sch ??? b. Thale, (illegible word) Schieferboden , 300 m, 24 June 1955, U . Dettman s.n. ( B100327808 ); Kreis Naumburg: Wegrand oberhalb Rossbach , 17 June 1977, P . Brückner , C . Gonnermann s.n. ( B100327796 ); Kreis Naumburg: Wegrand zwischen Rudelsburg und Bad Kösen, 27 June 79, Birgit Mory s.n. ( B100327812 ); Kreis Stendal : Böschungshang an der Elbe nördlich Arneburg , 24 June 1977, D. Benkert s.n. ( B100327695 ); Flora des Harzes. Elbingerode : An der Ruine Königsburg ( Bodfeld ) bei Königshof , 8 July 1913, Roman Schulz s.n. ( B100327651 ); Kreis Stendal : Strassenrand nördlich Tangermünde rechts der Strasse nach Hämerten , 10 July 1977, D. Benkert s.n. ( B100327696 ); Hangwald im NSG Arneburger Hang bei Arneburg, Kreis Stendal, 8 July 77, M . Bässler s.n. ( B100327826 ); Thuringia: mons Hausberg prope Jena , locis herbosis, 30 June 1911, H . Bessel Hagen s.n. ( MA112446 ); Rossbach, Wiesenberg , 17 June 1977, Strohr s.n. ( B); ???ersleben, 3 May 1894, Kappel s.n. ( E); Kreis Sonneberg, Wiesen am Galgenberg bei Schalkau, 17 June 1975, Müller s.n. ( B); Kunitz bei Jena , im Karstwald auf dem Rücken des Gleissberges, mit Elsbeere und Schlehe, 4 August 1954, W . Hempel 873 ( B100327817 ); zwischen Jena und Wöllnitz, 4 June 1950, Launert s.n. ( BM); Weg zum Eingefallenen Berg Werra Tal, 16 June 1962, Vent s.n. ( B); Saalfeld , gemein, 18 June 1905, J . Bornmüller s.n. ( B100327661 , B100327662 ); Weimar: Ettersberg , Jägerhütte , Kalk , 30 June 1927, J . Bornmüller s.n. ( B100327666 , B100327667 ); Weimar: Ziegental bei Öttern , September 1932, J . Bornmüller s.n. ( B100327663 ); Weimar: Berka , bei Bergern , 3 June 1945, J . Bornmüller s.n. ( B100327665 ); Martinsroda bei Orlamünde , 7 July 1965, W . Schleper s.n. ( B100327806 ); Hänge am Leutratal ( Jena ), 18 June 1958, U . Dettman s.n. ( B100327807 ); Bei Mellingen , 27 June 1927, J . Bornmüller s.n. ( B100327664 ); Flora des Harzes. Nordhausen: Trockene Gipsfelsen zwischen Steigertal und Grasmühle, 28 May 1920, Fiedler s.n. ( B100327652 ); Numburg am Kyffhäuser , July 1884, J . Bornmüller s.n. ( B100327656 ); an Greifenstein in Thrg. , 29 June 1884, J . Bornmüller s.n. ( B100327660 ); Fl. v. Weimar. Wach ?olderberg bei Berka , Kalk , July 1944, J . Bornmüller s.n. ( B100327659 ); ¿ Burgmenden ?, 15 May 1930, J . Bornmüller s.n. ( B100327658 ); Jena: Isserstedter Forst , 1 July 1908, J . Bornmüller s.n. ( B100327657 ); Leutratal bei Jena , Auf trock. Kalkhügel , 16 June 58, M . Bässler s.n. ( B100327825 ); Jena , 1907, A . Hase s.n. ( B100327670 ); Laubwald bei Jena , 12 June 1965, C . Beck 7579 ( B100327799 ); Wiese am Weg von Weimar nach N . Grünsteck , July 1909, J . Bornmüller s.n. ( B100327672 ); Rossbach / Thüringen: Wiesenweg , 17 June 1977, G . Stohr s.n. ( B100327820 ); Kreis Eisenberg: Am Wegrand auf Kalk bei Taupadel südwestlich von Jena, 25 May 1966, E . Peige s.n. ( B100327811 ); Bad Kösen / Saale , Kalkgeröllreicher wegrand am Weg zu Rudelsberg , 28 May 1961, Sigurd FrÖhner 1825 ( B100327797 ); Weimar : Legefeld , Hengstbachtal , 28 June 1922, J . Bornmüller s.n. ( B100327668 , B100327669 ); Kreis Sonneberg: Wiesen am Galgenberg bei Schalkau , 17 June 1975, U . Müller s.n. ( B100327821 ); In Thuringia, unknown collector ( K); Trockene Wiesen b. Weimar , July 22, Baschant s.n. ( B100327673 ); Weimar , Gelmeroda , felden, July 1920, J . Bornmüller s.n. ( B100327655 ); unknown or dubious state: Gebüsch bei Türnberg , 25 May 1902, C . Semler s.n. ( MA112447 ); Habitat in Germania, unknown collector ( K); Prov. Sachsen, Südharz, alter Molberg zu ¿ Rosselrode?, Gyps, 20 June 90, F . Wirtgen s.n. ( B100327805 ) .

HUNGARY. Borsod-Abaúj-Zemplén: Hungaria bor-or., Jósvafo, situ or. a Aggtelek et Domica, in mt. Nagyoldal, in pratis, solo calcareo, 601 m, 13 June 1961, Zertová s.n. (PR); Budapest: Hungaria centralis, in graminosis ad Budam, solo calc., 200 m, Steinitz s.n. (B100327761, BM, K, PR, WU); Budapest, W??? hat bei orfen, June, W. Sveinitz s.n. (MA112466); Budapest, Altofner Wiesen, 6 June 1886, V. Szépligeti s.n. (B100327752); Fejér: in exstirp. silvae fauces vallis Fáni-völgy prope Vérteskozma, 250 m, 27 May 1934, Boros s.n. (PR); Heves: Hungaria borealis, collis Nagyeged versus boreo-orientem ab oppido Eger, 250–450 m, 24 August 1970, Vasak s.n. (PR); Eger, entre Eger y Noszvaj, borde de pista forestal en margen de bosque de Quercus , sobre calizas, 410 m, 24 July 2014, X. Giráldez , M. M. Martínez Ortega, B. Rojas Andrés & N. López González MO 6110 (SALA155871); Nógrád: N Salgó, N von Salgótarján, ca. 600 m, Felsheiden, Felsen, Basalt, 9 July 1973, Krendl s.n. (W); Salgótarján, Salgó, en la subida al castillo, borde del camino, sobre basaltos, 571 m, 48º08'39.4''N, 19º50'50.1''E, 27 July 2014, X. Giráldez , M. M. Martinez Ortega, B. Rojas Andrés & N. López González MO 6111bis (SALA155869); Salgótarján, Somoskö (en dirección Vadaspark), prados herbosos en base de robledal, 451 m, 48º10'12.3''N, 19º50'58.0''E, 27 July 2014, X. Giráldez , M. M. Martínez Ortega, B. Rojas Andrés & N. López González MO 6111 (SALA155868); Pest: in valle Stara-Vod prope Szent-Endre, 4 June 1916, J. Andrsovszky s.n. (MA333222); in dumosis montis ¿Kakukhegy?, supra opp. Budapest, 430 m, 4 June 1936, Kápáti s.n. (PR); Visegrad Feketehegy, 2 October 75, Sag s.n. (BM); Vallée de Kataline près de Veröce, 22 June 80, unknown collector (BM); ca. 2 Km S of Domsod, grassland on calcareous alluvial soil of R. Danube, 30 May 1970, Th. J. Visser s.n. (MA522899); In declivibus graminosis montis Köhegy ad. pag. Pomáz com. Pest, 200–250 m, 21 May

TAXONOMIC REVISION OF VERONICA SUBSECTION PENTASEPALAE

Phytotaxa 285 (1) © 2016 Magnolia Press • 85 1936, A . Bartha s.n. ( B100327749 ); Pilisszántó , junto a la carretera hacia Piliisvörösvár, prados herbosos, 191 m, 47º39'13.0''N, 18º54'08.2''E, 28 July 2014, X. Giráldez , M. M . Martínez Ortega, B. Rojas Andrés & N. López González BR 278 ( SALA155870 ); unknown or dubious county: Ungarn, Malenica , June 189?, Wiemann s.n. ( WU) .

ITALY. Aosta: Valle di la Thuile, oberhalb La Thuile an der Strasse zum Petit Saint Bernard (Itinerarnummer 190), 1580 m, 45º42'32''N, 6º56'26''E, 23 July 2001, G. Schneeweiss, P. SchÖnswetter & A. Tribsch AT6522 (SALA124600); N-W Italien, Valle d’Aosta: Entreves bei Courmayeur, 22 July 1970, Polatschek s.n. (W); La Thuile, en la carretera SS26 hacia el Col du Petit St. Bernard, prados sobre sustrato ácido junto a un aparcamiento, 1615 m, 45º42'44.3''N, 06º55'58.5''E, 29 July 2011, X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés BR194 (SALA149408); Bolzano: Alp near St. Jacob [S. Giacomo] in Grödenthal [Val Gardena], South Tirol, 13 July 1874, Churchill s.n. (K); St. Christina de Gardena, pasado el pueblo en dirección Bolzano, 1400 m, 19 July 2000, L. Delgado , M. Martínez Ortega E. Rico & J. A. Sánchez Agudo JA 154 (SALA118205); Coi junto a Santa Cristina Valgardena , prados herbosos entre las casas, sobre calizas, 1349 m, 46º34'03.7''N, 11º41'32.6''E, 26 July 2011, X. Giráldez , M.M. Martínez Ortega & B. Rojas Andrés MO6043 (SALA149420); Padova: Colli Euganei, in nemore montis Castello supra Este, solo siliceo, 120 m, May 1912, Béguinot s.n. (K); Parma: In agro fidentino, N alle siepi, F. in Giugnu, Luglio, unknown collector (K); Pordenone: in loc. rupestr. pr. Erto, sol. calc., 2000’, 8 July 1873, Huter & Porta s.n. (BM, PRC); Sondrio: Lombardia, luoghi dumetoso-aprici presso Bormio, substr. vario, 1200–1600 m, 2–19 July 1909, Longa s.n. (RNG, UPA); Lombardei, Bormio, Kalk, Buschige Wiesenränder am nördl. Ortsrand, ca. 1250 m, 16 July 1980, Polatschek s.n. (W); Torino: Cafasse, boschi sotto il Mte. Basso, Valle di Lauzo, 18 May 1906, E. Ferrari s.n. (BM, MA112448); Trento: N slope of a (illegible word), on N sude of Sasso della Padella, ¿Primieso?, south Tirol, 5 July 1872, Churchill s.n. (K); Trieste: Ca??? tergestino, in dumetis prope Opcina, 13 June 1905, Evers s.n. (WU); M. Bocusso, June 1884, Marchesetti s.n. ( PRC); Udine: Weissenfels-Krain, July 1922, Meebold s.n. (K); Weissenfels-Krain, June 1921, Meebold s.n. (K); Verona: Monte Baldo, en la subida hacia la Malga de Valfreda, desde la Malga Ime, prados herbosos sobre calizas, 1482 m, 45º39'16.7''N, 10º48'48.1''E, 27 July 2011, X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés BR192 (SALA149407); Monte Baldo, en la subida a la Malga de Valfreda desde la Malga de Ime, prados secos sobre calizas, 1351 m, 45º39'18.5''N, 10º49'08.7''E, 20 June 2014, X. Giráldez , M. M. Martínez Ortega, N. Padilla García & N. López González NLG116 (SALA155884); Venetia, in montosis prope Mtem. Pastello, alla Majore, solo. calcar., 1000 m, 15 June 83, Rigo s.n. (K); unknown or dubious province: Cana?ei, Val di Lassa, 9 August 1925, Barton s.n. (BM).

LITHUANIA. Vilna: d Antokol, distr. Vilna, Po miejscach trawiastych., 22 June 1893, T . Symonowiczówna s.n. ( B100327790 , BM) .

POLAND. Greater Poland: Goranger Berge bei Czarnikau, June 1905, Bothe s.n. ( B100327782 , B100327785 ) ; Bentschen [= Zbąszyń]: Gebüsch links von der Chausee nach Tirschtiegel [= Trzciel], 4 July 1913, H. Bothe s.n. ( B100327786 ) ; Bentschen [= Zbąszyń] (illegible group of words), June 1904, illegible collector ( B100327781 ) ; Lesser Poland: Poloniae meridionalis districtus Chrzanów: vicus Dubie, in graminosis ad declive sub rupibus calcareis, 27 June 1974, Palkowa & J. Necka s.n. ( BM, RNG) ; Lower Silesia: Striegelmühl bei Zobten, 29 June 1890, G. Hieronymus s.n. ( B100327794 ) ; Podlaskie: E. Poland. Distr. Biatystok: slandy slope at the Bug river in the village Malozew near Siemiatycze , 27 June 1984, Adam Rostanski s.n. ( B100011361 ) ; Pomerania: Gross Watkowitz, August 1910, Luise Thiersch-Patzki s.n. ( B100327783 ) ; West Pomerania: Zehden [= Cedynia]: Bei Bellinchen [= Bielinek] an den Randhöhen des Odertals über der Ziegeleigrube , 30 May 1914, Roman Schulz s.n. ( B100327683 ) ; Zehden [= Cedynia]: Bei Bellinchen [= Bielinek] am Abhange des Odertals jeseils der Ziegeleigrube, 14 June 1914, Roman Schulz s.n. ( B100327682 ) ; Zehden [= Cedynia]: Bei Bellinchen [= Bielinek] an den Rand-hölen des Odertals über der Ziegeleigrube , 14 June 1914, Roman Schulz s.n. ( B100327684 , B100327893 ) ; Zehden [= Cedynia]: Bei Bellinchen [= Bielinek] am Abhange des Odertals jeseils der Ziegeleigrube, 4 July 1914, Roman Schulz s.n. ( B100327681 ) ; unknown or dubious province: ¿ Laubwaldwiesen? Dorf Wygoda bei Warschau , June 1866, illegible collector ( B100327780 ) ; West-Preusen, Pr. Friedland, sonnige Anhöhen, 12 June 1914, R. Gross s.n. ( B100327787 , B100327788 ) ; Josephinenberg bei Althof, June 60, R. Behnsch s.n. ( B100327709 ) .

ROMANIA. Alba: Transylvania, 3 Km N Mühlbach [= Sebes], ob des Szekesbaches, ca. 1–2 Km W der Strasse nach Daia, Gebüsch Trockenwiese tertiäre Sande, ca. 250–300 m, 6 June 1981, Krendl s.n. ( W) ; an dumetis: Langenthal [= Valea Lungă], 200 m, 16 June 1904, J . Barth s.n. ( B100327766 ) ; Brașov: Kronstadt [=

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Brașov]: son. Abh. o. Zinne [= Tâmpa], am Gipfel , Jura-Kalk, 800–950 m, 25 June 1912, J . Bornmüller s.n. ( B100327770 ); Zinne [= Tâmpa] bei Kronstadt [= Braşov], June 1885, illegible collector ( B100327771 ); Kronstadt [= Braşov]: am Gipfel der Zinne [= Tâmpa], 8–900 m, 27 June 1912, J . Bornmüller s.n. ( B100327769 ); Kronstadt [= Braşov]: Breitenberg bei Honigberg, 540 m, 3 July 1912, J . Bornmüller s.n. ( B100327767 ); Kronstadt [= Braşov], bei Honigberg [= Hărman], 540 m, 3 July 1912, J . Bornmüller s.n. ( B100327768 ); In monte "Zinn" [= Tâmpa] (961 m) prope Coronam [= Braşov] in Transsilvania, in calcareis, July 1890, J . RÖmer s.n. ( B100327764 ); In silvis prope Coronam [= Braşov] in Transsilvania, June 1889, J . RÖmer s.n. ( B100327763 ); Kronstadt [= Braşov]: Waldränder des "Breiten-Berg" bei Honigberg [= Hărman], c. 540 m, 3 July 1912, J . Bornmüller s.n. ( B100327772 ); Caraș-Severin: Banat, am Waldrande bei Drenkova a. d. Donau , 1 June 1909, Watzl s.n. ( WU); in (illegible word) m. Domogled, 6 July 1887, J . Bornmüller s.n. ( B100327745 ); Banat. M . Domogled , 6 July 1887, J . Bornmüller s.n. ( B100327746 ); Romania austr., distr. Orsova in valle riv. Cserna prope vicum Baile Herculanae, 160 m, 26 June 1970, Soják s.n. ( PR); Mehedinți: Banat, auf Wiesen vor dem Kasaanpasse, 3 June 1909, Watzl s.n. ( WU); Sibiu: In collibus: Talmesch [= Tălmaciu], 16 June 1908, J . Barth s.n. ( MA112456 ); Timiș: Banatus, distr. Timis , in herbidis ad margines silvarum prope pag. Banloc, solo humoso, ca. 91 m, 10 June 1948, Borza s.n. ( K) .

RUSSIA. Mordovia: Рузаевский р-н, В 4 км юго-запабнее с. Болбово, Опушка широколиственного леса, 23 June 1984, B . Tikhomirov , L . Deistfeldt , T . Yanitskaya , B . Skvorov s.n. ( B100327778 ); Инсарский р-н. В з км сев.-зап. г. Инсар. Остепнẽнный склон на правом берегу р. Исса., 2 I / YI 1984 , T. A . Troitskaya , A. V . Shpotya , M. N . Yefremova s.n. ( B100327776 ); Republic of Tatarstan: Prope stationem viae ferreae Observatorija, 30 km ad occidentem ab oppido Kazan , in prato sicco, 10 June 89, Askerova T ., Chairullina R . s.n. ( B100327779 ) ; Stavropol Krai: Maschuk ost ¿ Abhang ?, 28 June 1866, illegible collector ( B100327773 ); unknown or dubious region: Koenigsberg: Moosbude, 7 June 72, C . ¿ Raenit ? s.n. ( B100327792 ) .

SERBIA. Bor: zwischen Zlot und Brestovacka Banja, Kalk , zwischen Gebüschen am Strassenhang, 10–24 June 1978, Polatschek s.n. (WU); South Bačka: Com. Bars. Kysác, Via Tukokensis, 9 July 1912, S. Kupcok s.n. (BM); Novi Sad, Fruska Gora, Crveni Čot, prados herbosos, 467 m, 45º09'28.2''N, 19º45'00.4''E, 28 July 2010, S. Andrés , X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés MO5571 (SALA149412); Kosovo: Sar Planina, zwischen der Passhöhe (S Doganovic) und Globocnica, kalk, Wiese im Buchenwald, 10–24 June 1978, Polatschek s.n. (W); Pomoravlje: Beljanica planina, zwischen Despotovac und Stejevac E Cuprija, Kalk, buschiger Hang, 10– 24 June 1978, Polatschek s.n. (WU); Raška: Serbia, Suva Planina, 1000 m, 26 July 22, Turrill s.n. (K).

SLOVAKIA. Banská Bystrica: Muráň, Muránska Planina, prado en sotobosque de Larix , Betula y Pinus , 649 m, 16 July 2014, X. Giráldez , M. M. Martínez Ortega, B. Rojas Andrés & N. López González MO 6098** (SALA155865); Bratislava: Szeleskut [= Sološnica], kl. Karpaten, 11 June 1916, Ronniger s.n. (W); Thebener Kogel, prados sobre calizas, 3 June 2001, M. Martínez Ortega MO1318 (SALA124599); entre Sološnica y Plavecký Mikuláš, Malé Karpaty (kleine Karpaten), borde de bosque, sobre calizas, 207m, 48º28'25.0''N, 17º15'15.7''E, 15 July 2014, X. Giráldez , M. M. Martínez Ortega, B. Rojas Andrés & N. López González NLG 126 (SALA155867); Košice: in declivibus dumetosis ad opp. Igló [= Spišská Nová Ves ], 28 June 1914, Filarszky s.n. (BM, K, WU); Zadiel, prados herbosos sobre calizas, 262m, 48º20'54.9''N, 20º50'07.4''E, 26 July 2014, X. Giráldez , M. M. Martínez Ortega, B. Rojas Andrés & N. López González NLG 134 (SALA155866); Dvorníky- Vceláre, Zádiel, prados secos sobre calizas, ca. 300 m, 48º36'46''N, 20º50'32''E, 3 July 2002, M. Martínez Ortega, X. Giráldez & Muñoz Centeno MO1551 (SALA124598); Žilina: 21 c. Velká Fatra: inter T. Stiavnicka et Sklabina / kota Basta, 24 June 1971, Bernátová s.n. (B100327762).

SLOVENIA. Upper Carniola: Bled, Slovenis, jugo Slarva, June 1926, Heard s.n. ( BM); Wocheiner Feistritz- Kamnje, 31 August 1913, Ronniger s.n. ( W); Slovenija, limestone grassland near the Bellevue hotel, Mali Grad, SE corner of Bohinjsko Jezero, 4 July 1972, Halliday s.n. ( RNG) ; Savinja: Dolenjska: Brunk S of Radece, 555 m, 46º02'12.0''N, 15º10'23.0''E, 24 September 2009, B . Frajman 12656 ( SALA149411 ) .

SWITZERLAND. Basel-Landschaft: Basilea, in nemorosis montanis, May 1841, Meissner s.n. ( E) ; Bern: Erguel, Jura bern, 21 August 59, unknown collector ( K) ;

Geneva: Near Geneva, 4 June 1863, unknown collector (K); Graubünden: Engadina Bassa, Guarda, herbazal en talud, gneis-esquisto, 1432 m, SC11460, 17 July 1990, S. Castroviejo & al. s.n. (MA491440, SALA54926, SALA93469); Jan???, Grisons, 14 July 1867, Joad s.n. (K); Zermett, Grisons, 17 July 86, unknown collector (K); Buschränder ob der Bahnlinie, S. exp., 930 m, June 1914, Herbert Berger s.n. (B100327735); Celerina, cerca de St. Moritz, siguiendo el camino hacia el Alp Saluver, prados herbosos, 1825 m, 46º30'58.8''N, 09º51'09.2''E, 16 July 2011, X. Giráldez & B. Rojas Andrés BR 174 (SALA149271); Oberengandin, La Resgia, entre St. Moritz y Zernez,

TAXONOMIC REVISION OF VERONICA SUBSECTION PENTASEPALAE

Phytotaxa 285 (1) © 2016 Magnolia Press • 87 carretera 27, talud en el borde de la carretera, 1664 m, 46º37'48.8''N, 10º00'31.0''E, 15 July 2011, X. Giráldez & B. Rojas Andrés BR172 (SALA149403); Savoguin, meadow, 1730 m, 46.613ºN, 9.592ºE, 4 July 1999, D. Albach DA199 (WU0025650); Pontresina, Grisons, July 1886, Murray s.n. (K); Haute Engadine, canton Grisons, August 1855, unknown collector (K); Zernetz, Grisons, 16 July 1886, Murray s.n. (K); Engadin valley, canton Grisons, 1862, Hooker s.n. (K); Jura: In pascuis saxosis bei ¿Bémont? (illegible word) Jura, 3400 m, Lerch s.n. (K); Neuchâtel: Jura vue des Alpes, summit of pass between La Chaux de Fonds and Neuchâtel, 1285 m, 23 July 1966, Brummit s.n. (K); ¿Conret?, July 1907, P. Holzer s.n. (B100327718); La Vue-des-Alpes, entre Neuchâtel y La Chaux-de-Fonds, bordes de camino y prados sobre calizas, 1273 m, 47º04'26.4''N, 06º52'21.4''E, 12 July 2011, X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés BR169 (SALA149400); Valais: Bister, canton du Valais, pré maigre (Mesobromion), à pente 10º, sur calcschistes, avec Brachypodium pinnatum , Bromus erectus , Poa pratensis , Plantago media , Ranunculus bulbosus , Centaurea scabiosa , Thymus pulegioides , Stachys recta , Carex caryophyllea , Galium verum , G. boreale …, 1060 m, TH5426, 17 June 1986, J. P. Theurillat s.n. (SALAF21854); Fully, wood, 19 May 1886, Hamilton s.n. (E); Berne, in montibus et vallibus Jura etiam supra leukerbad Vale, 1837, Bischoff s.n. (E); Bister, cerca de Brig, siguiendo la carretera, al final del pueblo, borde de prado sobre esquistos micáceos, 1075 m, 46º21'34.6''N, 08º03'46.4''E, 28 July 2011, X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés MO6046 (SALA149421); w. Frimentz, August 1898, Nicholson & Reed s.n. (RNG); Vouvry, canton du Valais, district de Monthey, Chablais valaisan: Allamont, sommité du massif frontière du Grammont, au NW du lac de Tanay, praire, 1900 m, 14 August 1979, A. Stork s.n. (B, MA333223, RNG, SALAF2121, SEV75147); Wood N?ully, Valais, 19 May 1916, Hamilton s.n. (E); Zinal, Valais, subalpine pastures, 10 August 1903, unknown collector (K); Foot of (illegible word) wand, Valais, 5600’, 5 August 1924, Barton s.n. (BM); Valais, Evolena, between Lion and Ven, 30 May 1906 & 5 July 1901, Gregor s.n. (K); Wallis, direkt auf dem Plateau bei St. Christophe, 1585 m, bei Verbier, 24 July 1968, T. Eckardt 1015 (B100327741); En la subida al lago de Tanay, taludes en el borde de la pista, sobre calizas, 1418 m, 46º20'36.0''N, 06º50'33.4''E, 11 July 2011, X. Giráldez , M. M. Martínez Ortega & B. Rojas Andrés MO6038 (SALA149416); Vaud: Mt. ¿Triphon?, Vaud, 13 July 1874, Joad s.n. (K); near (illegible word), Ormond dess???, 20 June 1903, Murray s.n. (BM); Vallais de Vaud, (illegible group of words), 69, Thomas s.n. (K); Signal de Chexbres, Vaud, 5 June 69, unknown collector (K); Above Villeneuve, Vaud, 23 May 1896, Murray s.n. (K); Vaud, près Lausanne, à Béthusy, Leresche s.n. (E); unknown or dubious canton: Subalpine helvet., Wist s.n. (K); Wolver ¿in Witerlluz?, July 1885, R. Keller s.n. (MA112465).

UKRAINE. Dnipropetrovsk: (illegible word) bei Dnjepropetrovsk, June 1943, D. Grela s.n. ( B100327774 ) ; Donetsk: Slovyansk district, the town of Mayaki, the oak forest at the placor of the left bank the Sivirskiye Donets river , 9 June 1963, V . Tkachenko s.n. ( B100327777 ) ; Ivano-Frankovsk: Galich district, the outskirsts of Burshtin village, grassing slope of the mountain, named Kosova , 12 June 1984, O . Dubovik & L . Kritska s.n. ( B100327775 ) .

UNKNOWN OR DUBIOUS LOCATION. Switzerland?, about 1820, unknown collector ( K) ; Koenigsberg , Moosbade, unknown collector ( B) ; Transylvania , auf dem ¿ Gordiburg? bei (illegible word), July 1870, Schur s.n. ( K) ; Ex-Yugoslavia , ¿ Luva ? Planina , 1000 m, 26 July 22, Turrill s.n. ( K) ; Ex-Yugoslavia , in monte Bijelo (illegible word), prope Vardiste, 880 m, 24 June 1907, Maly s.n. ( WU) ; Wiesen bei Lomnitz, June 1882, Aur. Guil. Scherfel s.n. ( B100327759 ) ; Fl. Carinthiae, (three illegible words), unknown date, Herb. D. ¿ Pacherff? s.n. ( B100327723 ); (illegible word) Kalkschotter am Südhang d. ¿ Christianne? mare (Südkarpaten), July 1969, R. Bült ?? s.n. ( B100327721 ) ; illegible locality, 26 June 1836, L. v. Moor s.n. ( B100327731 ) ; illegible locality, June 43, illegible collector ( B100327717 ) ; unknown collector ( B100327736 ) ; unknown locality, unknown date, Schäde s.n. ( B100327674 ) ; ¿ Lied? du Saleve, June 1892, unknown collector s.n. ( B100327737 ); (illegible word) , 28 June 55, Krumbholz s.n. ( B100327798 ) ; two illegible words, 7 ¿ June? 49, Schäde s.n. ( B100327679 , B100327688 ) ; illegible group of words, unknown date, illegible collector ( B100327839 ) ; Lui ??????, 17 July 1971, Herbar Andrae s.n. ( B100327840 ) ; ????burg b. Breslau, unknown date, Nohr s.n. ( B100327712 ) ; 25 May 54, unknown collector s.n. ( B100327813 ) ; Flora von Siebenburgen, Ocesém, in subalpinis, 1400 m., 28 July 1897, Barth s.n. ( B100327765 ) ; In collib. calc. inter (illegible word) ¿ Stunjerthaln Hoh.?, 27 June 83, illegible collector ( B100327642 ) ; Laubwald, June 1905, Bothe s.n. ( B100327784 ) ; illegible word, 10 June 17, K. Osterwald s.n. ( B100327892 ) ; ¿ Lupamban ?, 22 May 1926, H. V. Berg s.n. ( B100327713 ) ; illegible word, 12 June 73, Erwin Koch s.n. ( B100327697 ) ; Blu. Bot. Garten, Gebirge ???südl. Europa. Bl. Blau, 27 October 42, Joh. Günther s.n. ( B100327888 ) ; Lausitzer Bergland : Spitzberg bei ¿ Scheibe?, Basalt., 880 m, 27 June 1919, O. Weder s.n. ( B100327654 ) ; Flora v. Westpreussen. Preuss. Friedland: Bergabhang südl. vom Seminar, 17 June 1890 & 10 June 1891, Rud. Grofs s.n. ( B100327711 ) ; two illegible words, 2 June 62, Golenz s.n. ( B100327690 ) ; illegible word, 18 ¿6/7? 18, Eisenstein s.n.

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(B100327883); unknown locality, unknown date, E. Loffia, J. Blume , s.n. ( B100327886 ) ; illegible word, June 1886, unknown collector ( B100327887 ) ; ¿ Pudlin ?, July 1886, unknown collector ( B100327760 ) ; Nant ???, 4000 ft., 25 June 1882, Clarke s.n. ( K); M. Jura supra (illegible word), 5 May 1834, unknown collector ( K) ; Flora carniolica, Grabovo um Wald ??????, 24 June 1890, ¿ Haperz? s.n. ( B100327744 ) ; ¿ Lagoso: Esinffog ?, 27 June 1943, ¿ Golenz? s.n. ( B100327687 ) ; Maroni? Deauville à Termèlle , August 1845, unknown collector ( P00208678 ) .

Material provisionally assigned to V. teucrium :— BOSNIA AND HERZEGOVINA. Municipality of Bosanski Petrovak: Bosnia, Petrovac, 19 June, Beck s.n. ( PRC); Sarajevo Canton: Bosnia, Vogosta, 7 July, Beck s.n. ( PRC); unknown canton/municipality: Bosnia, Krain, unknown date, Beck s.n. ( PRC).

GERMANY. Baden-Württemberg: Breisgau-Hochschwarzwald, Bitzenberg bei Achkarren , 25 May 1931, Sleimer s.n. ( W) .

ITALY. Cosenza: Calabria (Sila), c. 25 km W of San Giovanni in Fiore, lago Arvo c. 7 km W of Lorica , dry pastures on gravelly soils, c. 1300 m, Davis & Sutton s.n. ( BM) ; Potenza: Monte Pollino, Cóppola di Paola towards Col di Dragone , crevices in limestone pavement, 1600 m, 21 June 1979, Davis & Sutton s.n. ( BM) .

ROMANIA. Alba: Mănărade , predos secos en laderas cercanas a la carretera y a la vía de tren, 270m, 46º08'43.9''N, 23º58'41.3''E, 20 July 2014, X. Giráldez , M. M . Martínez Ortega, B. Rojas Andrés & N. López González MO 6107 ( SALA155837 ) .

SWITZERLAND. Neuchâtel: A La Brevine, Jura de Neuchâtel, 8 July 1853, Leresche s.n. ( W) ; La Brévine, Jura, Neuchat., July 61 & 72, Maret s.n. ( K) ; Chapelle de ¿ Béucoud?, Jura neuchateloise, July 1952, illegible collector ( MA112518 ) ; Montagnes du Jura, Neuchatel, chaille, 1809, unknown collector ( B).

Figures

FIGURE 25. Veronica teucrium var. teucrium (a–e, k. Bulgaria, Znepole region, Tran, D. Albach DA501, SALA 124597; g–h. Austria, Carinthia, Sankt Paul im Lavanttal, Rabenstein Burgruine, B. Rojas-Andrés et al. BR191, SALA149406; f, j. Slovakia, Košice, Zádiel, M. Martínez-Ortega et al. MO1551, SALA124598; l. Austria, Niederösterreich, Perchtoldsdorf, Kalksburg, M. Martínez-Ortega & G. Fischer, MO1453, SALA124565). Veronica teucrium var. angustifolia (i. France, Dep. Hautes-Alpes, Chorges, Mt. Chabrières, B. Rojas-Andrés et al. BR167, SALA149398): a, b. upper and lower part of a plant, respectively; c. Stem indument; d, f, g. Medium leaf (adaxial face); e. Detail of a medium leaf (adaxial face); h. Detail of a medium leaf (abaxial face); i, j. Medium leaf (adaxial face); k. Part of a flowering raceme; l. Part of a fruiting raceme, with a capsule. Illustration by Juan Castillo.

FIGURE 26. Distribution map of V. teucrium based on herbarium specimens (black circles) and collected material (open circles).