Tylopus baenzigeri Golovatch & Enghoff, 1993

Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak, 2016, The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina, European Journal of Taxonomy 195, pp. 1-47 : 32-34

publication ID	https://doi.org/10.5852/ejt.2016.195
DOI	https://doi.org/10.5281/zenodo.3852400
persistent identifier	https://treatment.plazi.org/id/03EB5A00-FFE0-7D49-AF9D-D56EFDAE9AF7
treatment provided by	Valdenar (2020-05-21 14:25:40, last updated by Carolina 2025-01-20 12:59:16)
scientific name	Tylopus baenzigeri Golovatch & Enghoff, 1993
status

Treatment

Tylopus baenzigeri Golovatch & Enghoff, 1993 View in CoL

Figs 23–26 View Fig View Fig View Fig View Fig

Tylopus baenzigeri Golovatch & Enghoff, 1993: 101 View in CoL , figs 50–51 (Type locality, Doi Suthep National Park).

Tylopus baenzigeri View in CoL – Enghoff 2005: 99. — Likhitrakarn et al. 2010: 25; 2014a: 65.

Material examined

THAILAND: ♂, Royal Agricultural Station Angkhang, Fang, Chiang Mai, 19°54’05”N, 99° 2’25”E, ca 1420 m a.s.l., 12 August 2014, leg. N. Likhitrakarn ( CUMZ).

Descriptive notes

MEASUREMENTS AND COLOUR. Length 43.8 mm, width of midbody pro- and metazonae 3.51 and 5.04 mm, respectively. Coloration of live animal, as well as of alcohol material black-brown ( Fig. 23 View Fig A–G), head and venter dark to light brown, legs and antennae yellowish brown to light yellowish.

BODY. Antennae rather short ( Fig. 23B View Fig ), extending behind body segment 3 (♂) when stretched dorsally. In width, segment 3 <4 <5 <collum <segment 2 <6–17; thereafter body gently and gradually tapering. Collum with three transverse rows of setae: 4+4 anterior, 1+1 intermediate and 4+4 posterior; a very faint incision laterally in posterior 1/3; caudal corner of paraterga very broadly rounded, not drawn behind rear tergal margin ( Fig. 23B – C View Fig ). Paraterga 2 well-developed ( Fig. 23B – C View Fig ), horizontal, lateral edge with two minute incisions in anterior half, caudal corner very narrowly rounded. Paraterga 3 and 4 each nearly pointed, expanded laterally, with two minute incisions in the middle. Sterna densely setose and finely microtuberculate ( Fig. 24D View Fig ); with a large, subtrapeziform, ventrally slightly concave, sternal lobe between ♂ coxae 4 ( Fig. 23I – J View Fig ). Legs rather long and slender, midbody ones ca 1.2–1.3 (♂) as long as body height; ♂ legs without tarsal brushes; ♂ postfemora and tibiae each with an adenostyle (tubercle) on segments 8–17; ♂ coxae, prefemora, femora and postfemora microtuberculate ventrally ( Fig. 24C – D View Fig ).

Remarks

This new specimen agrees in most details with the original description ( Golovatch & Enghoff 1993), especially in gonopod structure which shows a huge spine z serrate on both sides ( Figs 24A – B View Fig , 3 View Fig ), and in a large body. Thus, to document its identity the new specimen has been illustrated in detail for comparative purposes ( Figs 23–25 View Fig View Fig View Fig ). The type locality of T. baenzigeri, Doi Suthep National Park, is located quite far away (about 120 km) from this new place.

Checklist

Table 1 View Table 1 presents a checklist of the known or new Tylopus species, all arranged in alphabetic order. Data on their distribution are also provided.

References

Enghoff H. 2005. The millipedes of Thailand (Diplopoda). Steenstrupia 29 (1): 87 - 103.

Golovatch S. I. & Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125.

Likhitrakarn N., Golovatch S. I., Prateepasen R. & Panha S. 2010. Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 72: 23 - 68. http: // dx. doi. org / 10.3897 / zookeys. 72.744

Likhitrakarn N., Golovatch S. I. & Panha S. 2014 a. Three new species of the millipede genus Tylopus Jeekel, 1968 from Thailand, with additional notes on the species described by Attems (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 435: 63 - 91. http: // dx. doi. org / 10.3897 / zookeys. 435.8286

Figures

Fig. 23. Tylopus baenzigeri Golovatch & Enghoff, 1993, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively.

Fig. 24. Tylopus baenzigeri Golovatch & Enghoff, 1993, Ƌ. A–B. Right gonopod, lateral and mesal views, respectively. C. Leg 8. D. Legs of segment 11. Scale bars = 0.5 mm. Abbreviations are explained in the text.

Fig. 25. Tylopus baenzigeri Golovatch & Enghoff, 1993, Ƌ, right gonopod. A–B. Mesal and lateral views, respectively. C–F. Distal part, submesal, sublateral, subcaudal and suboral views, respectively. Scale bar = 0.2 mm.

Fig. 26. Distribution of the new eight species of Tylopus. Filled triangle. T. punctus sp. nov. Filled circle. T. acuminatus sp. nov. Crossed circle. T. acuminatus sp. nov., T. dorsalis sp. nov. Crossed square. T. acuminatus sp. nov. T. retusus sp. nov. Open circle. T. retusus sp. nov. Open triangle. T. baenzigeri Golovatch & Enghoff, 1993. Filled square. T. hongkhraiensis sp. nov. Open diamond. T. thunghaihin sp. nov. Inverted filled triangle. T. flavolineatus sp. nov. Open square. T. moniliformis sp. nov.

Fig. 3. Tylopus flavolineatus sp. nov., holotype, Ƌ, right gonopod. A–B. Mesal and lateral views, respectively. C–F. Distal part, subcaudal, subcaudal, suboral and sublateral views, respectively. Scale bars = 0.2 mm.