Malgadrymus pameroides Zámbó & Kondorosy, 2022
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publication ID |
https://doi.org/ 10.37520/aemnp.2022.016 |
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publication LSID |
lsid:zoobank.org:pub:31583440-DFEE-4C2A-AF77-2654A660E6F3 |
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persistent identifier |
https://treatment.plazi.org/id/03EA87F4-FFE7-3B57-FCED-FBFC560EFE52 |
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treatment provided by |
Felipe |
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scientific name |
Malgadrymus pameroides Zámbó & Kondorosy |
| status |
sp. nov. |
Malgadrymus pameroides Zámbó & Kondorosy , sp. nov.
( Figs 1 View Figs 1–2 , 5, 9 View Figs 3–10 , 12 View Fig )
Type locality. Central Madagascar, Ranomafana National Park,Antseranana district, Marovato village (S14°24′18.18″ E 47°42′50.80″).
Type material. Hඈඅඈඍඒඉൾ: J, Madagascar C / 6-10. I. 1998, Ranomafana / (pr. Fianarantsoa) / P. Pacholátko leg. // HOLOTYPUS [pr] / Malgadrymus / pameroides [hw] / det.Zámbó & Kondorosy [pr, red label] ( NHMW). Pൺඋൺඍඒඉൾඌ: N. Madagascar / Antseranana distr. / Sambirana riv. / Marovato vill., 2001 / D. Hauck leg. 5.-12.12. (1 J, EHIA); N Madagascar, 7.-16. i. 2015 / Mt. de Ambre N. P., 1042m / upper camp near Ambohitra / S12°31′37.7″ E49°10′19.1″ / M.Trýzna leg. (1 ♀, MMBC); Coll. Mus. Tervuren / N. E. Madagascar: / Fizono IX. 1959 / J. Vadon (1 ♀, RMCA); Coll. Mus. Tervuren / Madagascar: Fampanambo / 1962 / J. Vadon (1 ♀, RMCA); Coll. Mus. Tervuren / Madagascar: Fampanambo / 1961 / J. Vadon (1 ♀, RMCA); Coll. Mus. Tervuren / Madagascar: Fampanambo / 1960 / J. Vadon (1 J, RMCA); E Madagascar / Tamatave distr. / Andasibe, 17.-30.12. / D. Hauck leg. 2001 (1 ♀, EHIA); Madagascar East / distr. Mananara-N. / Antanambe VIII / Vadon et Peyrieras (1 ♀, EKKH); Madagascar-Est / Distr. Mananara-N / Seranombe / Vadon-Peyrieras / VII.1965 [hw] (1 ♀, MNHN); RNF / Jan 2017 FIT / E Madagascar 2017 / Ranomafana N.P.; 10.-14.i. / S21°15′46″ E47°25′17″, 981m / Flight Interc.Trap, P. Baňař lgt. (1J, MMBC); Madagascar 1995 / Fianarantsoa Distr. / Ranomafana en., Nov. 29 / Dec. 1., J. Stolarczyk leg. // 5977/23631 / coll. Upper Silesian Museum / (USMB) Bytom, Poland (1 ♀, USMB); Mus. Roy.Afr. Centr. / Madagascar Est: / Ambodivoangy [correctly Ambodivoahangy] I. 1960 / J. Vadon (1 J, RMCA); MADAG.: Ste. Marie / Manandriana-Fluss / 15.- 26.11.1993 / leg. Madl (1 ♀, NHMW); Madagaskar: Andasibe (Peri / net) E von Moramanga Berg- / regenwald am Flussufer, 920m / 18°55’40”S / 48°24’56”E / leg. U. Göllner, 28.-29. x. 2003 [blue label] / Zool. Mus. Berlin [green label] (1 ♀, ZMHB); CE Madagascar, 2.-4. ii. 2016 / Mantadia N. P., “circ. Eulophia” / S18°48′03.5″ E48°25′44.5″ (2JJ, MMBC); CE Madagascar, 5. ii. 2016 / Andasibe N. P., “circ. Indri 2” / S18°56′22.2″ E48°25′08.4″, M. Trýzna leg. (1 ♀, MMBC); Madagascar, 1000m / Ranomafana N. P., circuit / Talatakely, 8.-12. i. 2019 / S21°15′47.2″ E47°25′20.7″ / primary forest, M.Trýzna leg.(1 ♀, NMPC); Madagaskar: 2.5km NE von /Anara, SW von Iarintsena / ca. 975m NN, Buschtal / 21°51′03″S / 46°50′34″E / leg. U. Göllner, 6. xi. 2003 [blue label] / Zool. Mus. Berlin [green label] (1 ♀, ZMHB); Madagaskar: Ranomafana / NW von Infanadiana, 630m NN / sek. Bergregenwald / 21°15′40″S / 47°27′23″E / leg. U. Göllner, 2. xi. 2003 [blue label] / Zool. Mus. Berlin [green label] (1 ♀, ZMHB); SE Madagascar: 9km of NO of / Ft. Dauphin, 17.2.2004 / Foret de Mandena, 20m / 24°57′S 47°00′E, J. Janák lgt. (1 J, EHIA); Madagascar: Province / Fianarantsoa, Parc National / Ranomafana, radio tower / at forest edge, elev. 1130m / 21-28 January 2002 / 21°15.05′S, 47° 24.43′E / coll. M. Irwin, R. Harin’Hala / California Acad. of Sciences / malaise, mixed tropical / forest MA-02-09B-13 (in ethanol) (1 ♀, CAS); MADAGAS- CAR: Fianarantsoa Province, Parc National d’Isalo / Sahanafa River, 29.2 km 351° / N Ranohira 10-13 Feb 2003 / 22° 18′ 48″ S, 045° 17′ 30″ E / coll. Fisher, Griswold et al. / California Acad. of Sciences / general collecting gallery forest / elev 500m code: BLF7650 (in ethanol) (1 J, CAS); Madagascar: Fianarantsoa / Tsaranoro Massif, Catta / Camp 9 km SE Vohitsaoka / 22° 04′ 56″ S / 46° 46′ 32″, 950m / rivulet near Camp, shore / with bushes and trees / leaf litter sievings / 16.-18. iv. 2007, lg. M. Uhlig [blue label] / Museum für Naturkunde, Berlin (1 ♀, ZMHB); Madagascar: Fianarantsoa / Tsaranoro Massif, Catta / Camp 9 km SE Vohitsaoka / 22° 04′ 59″ S / 46° 46′ 34″, 950m / rivulet forest, shore / with bushes and trees / leaf litter sievings / 18. iv. 2007, lg. M. Uhlig [blue label] / Museum für Naturkunde, Berlin (1 J, ZMHB).
Description. Colour. Body ( Fig. 1 View Figs 1–2 ) fuscous (sometimes pale brown); proximal half (or more) of distiflagellum except for narrowly brown base, at least anterior half of clavus along corial margin and one spot at apex of clavus (sometimes entire clavus white and basal claval spot rather pale yellow), proximal half of corium and one spot on posterior part of lateral margin, meso- and metafemora except for more or less wide subapical ring and basal and inner margin of membrane widely and arcuately (except a fuscous subbasal reverse V-shaped spot) white; pedicel (dark ochraceous contrasting to other yellow body parts), labium except for fuscous apex of labiomere IV, one spot on each lateral margin of posterior pronotal lobe (often extending onto humeral angles but sometimes absent) and one inverse T-shaped spot in middle of posterior pronotal lobe (being often white), one relatively large subquadrangular subapical spot on corium close to claval commissure, often apex of profemora, tibiae and tarsi pale yellow; scape yellowish-brown (sometimes rather pale brown); apical and lateral margin of corium usually red (often fuscous); often posterior half of pronotum, dark parts of corium, metepimeroid and dorsal side of abdomen with connexiva paler brown.
Pilosity. Dorsal surface of head except for pair of long cephalic trichobothria with hardly visible decumbent pubescence, better visible on anterior part, antenniferous tubercles mostly bearing 1–2 somewhat elongate fine erect setae on ventral part, antennomeres densely covered with rather short fine decumbent and very sparsely with fine semierect setae (except for pedicel); head ventrally sparsely covered with short fine semidecumbent setae. Dorsal surface of body and sternum bare, abdominal venter densely covered with short fine decumbent setae. Legs sparsely covered with short fine decumbent setae, profemora bearing 4–6 elongate fine erect setae relatively far from each other in row ventrally; tibiae with short fine stiff semierect setae, tarsi dorsally with some elongate fine semierect setae as well.
Structure. Mostly shiny except for dull scutellum and sometimes parts of pronotum, sternum dull except for shiny mesosternum. Abdomen shiny. Scutellar row of punctures on clavus always very sparse, consisting of less than 10 punctures. Area between corial furrow and row along Cu with some sparse punctures proximally, apical part without punctures between rows (excluding apical row). Body 3.0–3.2 times (males: 3.1–3.2, females: 3.0–3.2) longer than wide. Eyes 2.1–2.4 times wider than length of rather short antenniferous tubercles. Widest part of pronotum 1.1–1.4 times (males: 1.1–1.2, females: 1.2–1.4) wider than entire length, anterior lobe of pronotum 1.1–1.4 times (males: 1.1–1.3, females: 1.1–1.4) longer than posterior one.
Male abdomen and genitalia. Abdomen more or less evenly flat. Pygophore: Subglobose ( Fig. 5 View Figs 3–10 ), dorsal aperture more or less round and large: entire length of pygophore about 1.5 times as long as dorsal aperture; both parts separated with large triangular projection (with slightly indented anterior side), distance between projections similar to length of each one; posterior margin with triangular incision ( Fig. 5 View Figs 3–10 ). Entire surface covered with numerous short fine decumbent setae, posterior part with long fine erect setae, as well. Paramere: Blade ( Fig. 9 View Figs 3–10 ) elongate triangular (length of blade about half entire length of paramere), about 1.4 times longer than widest part and ending in long and relatively narrow apex ( Fig. 9 View Figs 3–10 ), inner projection triangular and about as long as wide, outer projection more or less straight in anterior three quarters and slightly rounded in posterior one quarter (and hardly protruding as well), shank well developed.
Female abdomen. Abdominal sternite VI medially strongly narrowed (posterior margin forming “V”), sternite VII medially almost as long as sternites IV–VI together. Abdomen higher, 1.49–1.58 times longer than high, outline of sternite VII straight, connecting to sternite VI in angle of about 45°.
Measurements (in mm). Males (n = 9; holotype in parentheses). Total body length: 3.47–4.05 (3.99); head: length 0.51–0.66 (0.57), width 0.56–0.75 (0.59), interocular space 0.36–0.45 (0.42); length of antennomeres: I 0.30–0.53 (0.48), II 0.75–0.94 (0.91), III 0.62–0.87 (0.81), IV 0.61–0.72 (0.72); length of labiomeres: I 0.36–0.42 (0.42), II 0.39–0.51 (0.51), III 0.23–0.27 (0.27), IV 0.17–0.18 (0.18); pronotum: length 0.75–1.02 (1.02), width 0.96–1.14 (1.07); scutellum: length 0.51–0.63 (0.59), width 0.49–0.59 (0.57).
Females (n = 15). Total body length: 2.91–3.93; head: length 0.54–0.60, width 0.60–0.76, interocular space 0.39– 0.45; length of antennomeres: I 0.37–0.45, II 0.65–0.99, III 0.62–0.75, IV 0.62–0.71; length of labiomeres: I 0.39–0.43, II 0.41–0.57, III 0.23–0.30, IV 0.17–0.19; pronotum: length 0.81–0.96, width 0.90–1.17; scutellum: length 0.59–0.75, width 0.50–0.75.
Etymology. The scientific name “ pameroides ” refers to the resemblance to the shape of the pronotum of different Myodochini genera; Pamera Say, 1831 was the old collective name for them, the suffix - oides means ‘resembling, -like’. Collecting circumstances. Specimens of M. pameroides were collected by various collecting methods, namely (P. Baňař and M. Trýzna) by sifting of forest leaf litter, sweeping of vegetation, beating of trees and shrubs, by flight interception traps, and they were also attracted to light; a single female was also collected in a Malaise trap. According to various localities and habitats of the type series (see e.g. Fig. 15 View Fig ), as well as large spectrum of collecting methods this species seems to be ecologically rather flexible.
Distribution. Endemic to Madagascar ( Fig. 12 View Fig ).
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