Celes Saussure, 1884

Storozhenko, S. Yu. & Benediktov, A. A., 2020, AN ANNOTATED CHECK-LIST OF ORTHOPTERA OF TUVA AND ADJACENT REGIONS. PART 3. SUBRODER CAELIFERA (ACRIDIDAE: GOMPHOCERINAE: GOMPHOCERINI; LOCUSTINAE), Far Eastern Entomologist 402, pp. 1-36 : 20-21

Genus Celes Saussure, 1884

22(86). Celes skalozubovi skalozubovi Adelung, 1906

Celes skalozubovi: Miram, 1907: 7–8 ; Berezhkov, 1951: 20–21; Sergeev, 1986: 212; Sergeev et al.,

1995: 96–97; Benediktov, 1997: 117.

Celes skalozubovi skalozubovi: Berezhkov, 1956: 158–159 .

MATERIAL. Tuva: W Sayan Mts., Alash Plateau, Ak-Sug River, 51°23ʹN, 90°28ʹE, 1208 m,

southern slope, stony steppe, 13.VIII 2016, 1 larva ( MS); W Sayan Mts. , Alash Plateau, Ak-Sug River ,

51°25ʹN, 91°04ʹE, 800–810 m, southern slope and upper terrace, semi-deserts, 13.VII 2016, 8 ♂ ( MS) ;

W Sayan Mts. , Alash Plateau, Ak-Sug River, 51°25ʹN, 91°04ʹE, 810 m, lower terrace and upper floodplain, meadows, 13.VIII 2016, 1 ♂ ( MS); Turan-Uyuk Intermountain Basin, Begreda River, 51°59ʹN ,

94°18ʹE, 829 m, terrace, meadow, 12.VIII 2018, 1 ♂ ( MS); E Tuva, Academician Obruchev Range,

southern slope, Koptu (Khapto) River, 51°44ʹN, 95°26ʹE, 1006 m, southern slope, stony steppe, 11.VIII

2018, 2 ♂, 1 ♀ ( MS); E Tuva, Academician Obruchev Range, southern slope, Koptu (Khapto) River ,

51°37ʹN, 95°21ʹE, 788–843 m, southern slope and upper terraces, steppes, 11.VIII 2018, 2 ♂, 1 ♀ ( MS) ;

Chaa-Khol River , 51°34′N, 92°23′E, 570–600 m, dry steppes, 14–18.VII 1962, 1 ♂, 3 larvae ( IS); 14 km

SW Shagonar City, Shagonar River, 51°28ʹN, 92°44ʹE, 615–629 m, slopes and piedmont plain, semideserts, 21, 23. VI 2017 , 11 larvae ( MS); 32 km SW Kyzyl City, Elegest River , 51°29ʹN, 94°10ʹE, steppe ,

22.VII 2014, 1 ♀ ( SS); Ulug-Khem Intermountain Basin , Elegest River, middle part, 51°22ʹN, 94°04ʹE ,

695 m, upper terrace, dry steppe, 16.VI 2017, a specimen observed ( MS); Kaa (Malyj Yenissei) River ,

right side, near Boyarovka settlement, 51°32ʹN, 95°21ʹE, 703 m, plain, dry steppe, 12.VIII2018, 3 ♂,

1 ♀ ( MS); 6 km SE Baj-Haak settlement, Sosnovka settlement, 51°08ʹN, 94°32ʹE, slope of balka with steppe, 20–21.VII 2014 , 4 ♂ ( SS); SE Ulug-Khem Intermountain Basin, Uzun-Kharagan River , near

Balgazyn settlement, 50°57ʹN, 95°16ʹE, 940 m, southeastern slope, stony steppe, 8.VII 1978, 22 larvae

( MS); SE Ulug-Khem Intermountain Basin, Uzun-Kharagan River, near Balgazyn settlement, 50°57ʹN,

95°16ʹE, 940 m, upper terrace, meadow with Caragana bushes, 8.VII 1978, 6 larvae ( MS); SE Ulug-Khem

Intermountain Basin, Shuurmak River, 25 km S Balgazyn settlement, near Kuran settlement, 50°47ʹN,

95°17ʹE, 1030–1050 m, southern slope, steppe and meadow, 10.VII 1978, 7 larvae ( MS); W Tannu-Ola

Mts., 5 km N Khandagajty settlement, 50°48ʹN, 92°05ʹE, 1300–1350 m, piedmont plain, stony steppes with Caragana , 16.VIII 1985, 1 ♂ ( MS); E Tannu-Ola Mts. , Shuurmak River, 25 km S Balgazyn settlement, near Kuran settlement, 50°47ʹN, 95°17ʹE, 1041 m, southern slope, mountain steppe, 10.VII

1978, 5 larvae ( MS); E Tannu-Ola Mts. , Shivelig River, 50°45ʹN, 94°34ʹE, 1330–1340 m, upper terrace ,

steppe and short meadow with Caragana , 17.VII 1978, 1 larva ( MS); E Tannu-Ola Mts. , Shuurmak

River, 5 km N Shuurmak settlement, 50°41ʹN, 95°19ʹE, 1173 m, local southern slope and piedmont plain,

steppe, 10.VII 1978, 6 larvae ( MS); E Tannu-Ola Mts. , Shuurmak River, 5 km N Shuurmak settlement ,

50°42ʹN, 95°20ʹE, 1106 m, southern slope, mountain steppe, 11.VII 1978, 3 larvae ( MS); E Tannu-Ola

Mts., 7 km W Shuurmak settlement, 50°37ʹN, 95°14ʹE, 1400 m, local southern slope, stony steppe,

12.VII 1978, 6 larvae ( MS); E Tannu-Ola Mts. , Shuurmak Pass, 50°37ʹN, 95°11ʹE, 1495 m, southern slope and upper terrace, mountain steppe , 12.VII 1978, 9 larvae ( MS); N Uvs-Nuur Intermountain Basin ,

Iribitej River , 50°44′N, 93°08′E, 938–973 m , piedmont plain, stony semi-deserts with Nanophyton grubovii , 24.VI 1978, 5 larvae ( IS, MS); N Uvs-Nuur Intermountain Basin, Iribitej River , 50°44′N ,

93°08′E, 938 m, lower terrace and flood-plains, steppe, 30.VII 1978, 2 ♂ ( MS); NE Uvs-Nuur

Intermountain Basin , Shivelig River, 50°39ʹN, 94°28ʹE, 990–995 m, flood-plain and terraces, dry steppes with Caragana , 28.VI 1978, 3 larvae ( MS); the same locality and habitats, 18.VII 1978, 3 ♂, 10 larvae

( MS); the same locality and habitats, 10.VIII 1978, 9 ♂, 9 ♀ ( MS); NE Uvs-Nuur Intermountain Basin ,

Shivelig River , 50°43ʹN, 94°33ʹE, 1215 m, lower terrace, dry steppe, 15.VII 1978, 1 larvae ( MS); NE

Uvs-Nuur Intermountain Basin , Shivelig River, 50°43ʹN, 94°33ʹE, 1200–1300 m, southern slopes and piedmont plain, steppes with Caragana , 24.VII 2003, 2 ♀, 65 larvae ( MS); Uvs-Nuur Intermountain

Basin , Tes River, 50°33ʹN, 94°31ʹE, 932–934 m,terraces, dry meadows and steppes, 27.VI 1978, 2 larvae

( MS); Uvs-Nuur Intermountain Basin, Tes River, 50°32ʹN, 94°31ʹE, 930–932 m, upper flood-plain,

meadows, 13.VII 1978, 2 larvae ( MS); Uvs-Nuur Intermountain Basin , 7 km S Erzin settlement, Tes

River, 50°11ʹN, 95°12ʹE, 1097 m, flood-plains, meadows, 21.VII 1978, 4 larvae ( MS); Uvs-Nuur

Intermountain Basin, 7 km S Erzin settlement, Tes River, 50°11ʹN, 95°12ʹE, 1097 m, lower terrace, steppes,

23.VII 1978, 1 larva ( MS). Krasnoyarsk Region: southern part, W Sayan Mts. , Us Intermountain Basin ,

near abandoned Idzhim settlement, 52°21′N, 93°13′E, 780–800 m, southern slope, dry mountain meadow and steppe, 22.VII 1995, 6 larvae ( MS); the same locality, 52°20′N, 93°20′E, 700–750 m, low terrace ,

meadow, 22.VII 1995, 1 larva ( MS); the same locality, 52°16′N, 93°07′E, 680–700 m, terraces and southern slope, steppes, 22–23.VII 1995, 18 larvae ( MS) .

DISTRIBUTION (nominotypical subspecies). Tuva: WSW, WSE, ET, KHE, UKH, KKH,

ETO, UVS. – S Siberia; N Kazakhstan, Mongolia, N China. Celes skalozubovi akitanus

(Shiraki) occurs in S Russian Far East, E Mongolia, NE China, Korea, and Japan.

ECOLOGY. More or less common form of the dry steppes.

Tribe Bryodemini