Gomphocerus Thunberg, 1815

Storozhenko, S. Yu. & Benediktov, A. A., 2020, AN ANNOTATED CHECK-LIST OF ORTHOPTERA OF TUVA AND ADJACENT REGIONS. PART 3. SUBRODER CAELIFERA (ACRIDIDAE: GOMPHOCERINAE: GOMPHOCERINI; LOCUSTINAE), Far Eastern Entomologist 402, pp. 1-36 : 2-4

Genus Gomphocerus Thunberg, 1815

2(66). Gomphocerus sibiricus (Linnaeus, 1767)

Gomphocerus sibiricus: Miram, 1907: 6 ; Ivanova, 1967: 133.

Gomphocerus sibiricus sibiricus: Berezhkov, 1956: 100 .

Aeropus sibiricus: Berezhkov, 1951: 19–20 ; Sergeev et al., 1995: 96–98; Benediktov, 1997: 117;

Bukhvalova & Vedenina, 1998: 118.

Aeropus sibirikus (sic!): Pavlov, 2004: 65.

MATERIAL. Tuva: W Sayan Mts., Sayanskij Pass, 50°42ʹN, 89°53ʹE, 2230–2240 m, mountain tundra, 12.VII 2012, 1 larvae ( MS); W Sayan Mts., Alash Plateau, Ak-Sug River , 51°37ʹN, 90°05ʹE ,

1450 m, terraces and flood-plain, short meadows, 14.VIII 2016, 1 ♂ ( MS); Alash Plateau, Kara-Khol ,

25. VI 2003, 1 ♀ (M. Zasypkina); Chaa-Khol River , 51°34′N, 92°23′E, 570–600 m, dry steppes, 14–18.VII

1962, 2 ♂, 1 ♀ ( IS); 6 km SE Baj-Haak settlement, Sosnovka settlement, 51°08ʹN, 94°32ʹE, plakor, mixed steppe, 20–21.VII 2014 , 1 ♂, 2 ♀ ( SS); SE Ulug-Khem Intermountain Basin, Uzun-Kharagan River , near

Balgazyn settlement, 50°57ʹN, 95°16ʹE, 940 m, southeastern slope, stony steppe, 8.VII 1978, 3 ♀ ( MS) ;

SE Ulug-Khem Intermountain Basin, Shuurmak River , 25 km S Balgazyn settlement, near Kuran settlement, 50°47ʹN, 95°17ʹE, 1000–1039 m, terraces, steppe and meadow, 9.VII 1978, 1 ♂, 9 ♀ ( MS) ;

W Tannu-Ola Mts., 9 km N Khandagajty settlement, 50°53ʹN, 92°04ʹE, 2250 m, mountain tundra,

19.VIII 1985, some specimens observed ( MS); W Tannu-Ola Mts. , 9 km N Khandagajty settlement ,

50°52ʹN, 92°04ʹE, 2060 m, alpine steppe and meadow, 19.VIII 1985, 10 ♂, 13 ♀, 6 larvae ( MS); W

Tannu-Ola Mts., 9 km N Khandagajty settlement, 50°51ʹN, 92°04ʹE, 1700–1900 m, mountain steppes,

19.VIII 1985, 31 ♂, 26 ♀, 6 larvae ( MS); W Tannu-Ola Mts. , NW Khandagajty settlement, near Solchur settlement, 50°46ʹN, 92°01ʹE, 1150–1200 m, agricultural fields including abandoned and plots along canal and roads , 21.VIII 1978, 1 ♀ ( MS); W Tannu-Ola Mts. , 5 km W Torgalyg settlement, Khandybai

Mt., near timber-line, meadows, 23.VI 1962, 5 ♂, 4 ♀, 6 larvae ( IS); W Tannu-Ola Mts. , 5 km W

Torgalyg settlement, Khandybai Mt. , mountain steppe, 23. VI 1962, 2 ♂, 12 ♀, 10 larvae ( IS); E TannuOla Mts. , Shivelig River , 50°45ʹN, 94°34ʹE, 1330–1340 m, lower terrace, steppe and short meadow between shrubs and Larix trees, 17.VII 1978, 8 ♂, 25 ♀ ( MS); the same locality, southern slope and upper terrace, stony steppes, 17.VII 1978, 3 ♀ ( MS); E Tannu-Ola Mts., Shivelig River, 50°45ʹN, 94°34ʹE, 1330 –

1340 m, upper terrace, steppe and short meadow with Caragana , 17.VII 1978, 6 ♂, 13 ♀, 2 larvae ( MS) ;

the same locality, upper flood-plain, meadow, 17.VII 1978, 2 ♂ ( MS); the same locality, lower terrace ,

steppe, 12.VIII 1978 5 ♂, 7 ♀ ( MS) ; the same locality, upper terrace, steppe, 12.VIII 1978, 3 ♀ ( MS) ; the same locality, upper flood-plain, meadow, 12.VIII 1978, 1 ♀ ( MS) ; the same locality, upper terrace, short meadow, 12.VIII 1978, 2 ♀ ( MS) ; E Tannu-Ola Mts. , Shivelig River, 50°45ʹN, 94°34ʹE, 1330–1340 m ,

flood-plain and terraces, steppe and short meadow between shrubs and Larix trees, 28.VI 1978, 15 ♂, 30 ♀,

20 larvae ( MS); E Tannu-Ola Mts. , Shuurmak River, 25 km S Balgazyn settlement, near Kuran settlement, 50°47ʹN, 95°17ʹE, 1041 m, piedmont plain, grazed meadow, 10.VII 1978 , 1♀ ( MS); E

Tannu-Ola Mts. , Shuurmak River , 5 km N Shuurmak settlement, 50°41ʹN, 95°19ʹE, 1173 m, local southern slope and piedmont plain, steppe, 10.VII 1978, 1 ♂, 1 ♀ ( MS); E Tannu-Ola Mts., Shuurmak

River, 5 km N Shuurmak settlement, 50°42ʹN, 95°20ʹE, 1106 m, southern slope, mountain steppe, 11.VII

1978, 2 ♂, 3 ♀ ( MS); E Tannu-Ola Mts. , Shuurmak Pass, 50°37ʹN, 95°11ʹE, 1495 m, southern slope and upper terrace, mountain steppe, 12.VII 1978 , 17 ♂, 26 ♀ ( MS); E Tannu-Ola Mts. , Shuurmak Pass ,

50°38ʹN, 95°11ʹE, 1492 m, short meadow, 5.VII 2017, 4 ♂, 12 ♀ ( MS); E Tannu-Ola Mts. , 13 km NEE

Samagaltaj settlement, Kaldak-Khamar (Shuurmak) Pass, 50°38ʹN, 95°11ʹE, 1500 m, rangeland, 19.VII

2014, 3 ♂, 4 ♀ ( SS); NE Uvs-Nuur Intermountain Basin, Shivelig River , 50°43ʹN, 94°33ʹE, 1200–1300

m, southern slopes and piedmont plain, steppes with Caragana , 24.VII 2003, a male observed ( MS); NE

Uvs-Nuur Intermountain Basin, Shivelig River, 50°45ʹN, 94°34ʹE, 1280–1301 m, terraces, dry meadows,

8.VII 2017, some specimens observed ( MS); NE Uvs-Nuur Intermountain Basin, Shivelig River ,

50°43ʹN – 50°45ʹN, 94°33ʹE – 94°34ʹE, 1400–1650 m, mountain slopes, 24. VI –12.VIII 1978,>10

specimens, including larvae ( T. Myagkaya, A. Lee); Uvs-Nuur Intermountain Basin, near Shara Lake,

50°14ʹN, 94°31ʹE, 894 m, flood-plains of lake and small stream, short meadows with halophytes, 6.VII

2017, 1 ♀ ( MS); Uvs-Nuur Intermountain Basin, S Tore Lake , 50°01′N, 95°04′E, 1154-1166 m, 07.VII

2017, sand semi-desert and upper terraces with birch forest, 2 ♀ ( MS); Uvs-Nuur Intermountain Basin, S

Tore Lake, 50°05′N, 95°09′E, 1174 m, sandy semi-desert with Caragana bungei , 7.VII 2017, some specimens observed ( MS); 31 km NEE Erzin settlement, Erzin River, 50°21ʹN, 95°34ʹE, 1300 m, northern slope, balka with Caragana , 18.VII 2014, 2 ♀ ( SS). Krasnoyarsk Region: southern part, West Sayan

Mts., Us Intermountain Basin, near abandoned Idzhim settlement, 52°16′N, 93°07′E, 680–700 m,

terraces and southern slope, steppes, 22–23.VII 1995, 1 ♂, 9 ♀ ( MS) .

DISTRIBUTION (nominotypical subspecies). Tuva: WSW, UKH, KKH, MT, WTO,

ETO, UVS, SAN. – N, NE Europe, Siberia (except the extreme North); N Kazakhstan, N

Mongolia, NE China. Other subspecies are in the mountains of S Europe, Asia Minor, Caucasus, Tien Shan, Pamiro-Alay, SE Tibet, and Himalayas.

ECOLOGY. In Tuva, the species inhabits a wide variety of habitats: from the mountain tundra to the sandy semi-deserts. However, it prefers the mountain steppes with some grasses and forbs.