Ilex subrotunda Doweld, 2019

Doweld, Alexander B., 2019, New names for Ilex and Ilexpollenites (Aquifoliaceae), extant and fossil. Notulae Systematicae ad Palaeofloram Europaeam spectantes II. Aquifoliaceae, Phytotaxa 388 (2), pp. 179-191 : 184

Ilex subrotunda Doweld , sp. nov.

≡ Ilex ohashii Huzioka (1963: 208) , nom. inval. (Art. 40.1 of the ICN).

Type:—[fossil leaves] Yamakayakusa, Ani-machi, Kita-Akita-gun Akita Prefecture, Japan (holotype, AKMG-3132, Institute of Applied Geology, Mining College, Akita University, Akita, Japan-illustrated by Huzioka 1963: pl. 35, fig. 10).

Description:—Leaves elliptical, 4.5–5 cm long, 1.9–2.5 cm wide, acute at both basis and apex, entire, long-petiolate (6 mm long); venation of 7 pairs of secondary veinlets leave the midrib at angles of 45–40, camptodrome, tertiary venation reticulate or percurrent in the inter-secondary spaces (after Huzioka 1963, modified).

Occurrence:—Middle Miocene (Utto Formation); Eurasia (Japanese Archipelago).

Eponymy:—Similar to extant Ilex rotunda Thunberg (in Murray 1784: 168).

IFPNI:— 9DFB6C2F-7C4D-0AAB-DDD6-71C69F0A13C4.

Notes:—The fossil-species Ilex ohashii Huzioka (1963: 208) was not validly published since the type was not designated (Art. 40.1 of the ICN) [two specimens from two locations were incorrectly cited as “ holotypes ”]. Huzioka (1963) related it to extant Ilex rotunda Thunberg , therefore, a new name is proposed to replace invalid I. ohashii .