Beebea guglielmi Schaus

Bernard Landry, 2016, Taxonomic revision of the Spilomelinae (Lepidoptera, Pyralidae s. l.) of the Galápagos Islands, Ecuador, Revue suisse de Zoologie 123 (2), pp. 315-399 : 341

Key to the brown Galápagos species of Spilomelinae with darker brown markings, excluding Beebea guglielmi Schaus

1 Forewing ground colour golden brown, with distinct darker brown markings as complete basal, antemedial, and postmedial fasciae, and orbicular and reniform stigmata at base and end of cell respectively; hindwing linear markings also distinct, without other spots; wingspan usually above 22.5 mm (20-26 mm); male antenna evenly thin ( Fig. 48 ).................................................................................................................. Psara chathamalis (Schaus)

1’ Forewing ground colour greyish brown to dark brown, with darker brown markings often indistinct, without apparent basal line, usually with only reniform stigma apparent; hindwing markings as poorly defined fascia(e), with or without antemedian spot or short dash, or two spots; wingspan below 22.5 mm; male antenna thickened at base in one species.................................................................................................................................................... 2

2 Forewing reniform stigma contrasting, comma shaped; hindwing with shorter scales of fringe dark greyish brown and longer scales white; wingspan 18-22 mm; male antenna thickened at base and hindwing anal sector enlarged ( Figs 49, 50 )...................................................................................................... Rhectocraspeda periusalis (Walker)

2’ Forewing reniform stigma more or less contrasting, rarely comma shaped in one species; hindwing never with longer scales of fringe white, although sometimes paler than basal scales; wingspan 16-22.5 mm; male antenna not thickened at base and hindwing anal sector not enlarged......................................................................................... 3

3 Wingspan 18-22.5 mm; forewing reniform stigma a small spot or short dash; hindwing with one small contrasting spot or short dash in cell ( Figs 20, 21 ) ...................................................... Herpetogramma phaeopteralis (Guenée)

3’ Wingspan 15-20 mm; forewing reniform stigma large, variable, sometimes with paler centre or with second encircling ring; hindwing either with two clearly contrasting spots or an indistinct line ........................................ 4

4 Dark brown; forewing reniform stigma squarish, with paler centre; hindwing with indistinct fascia(e); termen with contrasting dark spots on veins of both wings; sexes dissimilar with male sporting distinctly elongate abdomen and forewing fovea ( Figs 44, 45 ) .......................................................................................... Penestola bufalis (Guenée)

4’ Drab to dark greyish brown; forewing reniform stigma variable, often with thin dark line encircling other marking, ‘U’ shaped to ‘8’ shaped; hindwing with two distinct spots in cell and postmedian fascia; termen with contrasting dark spots on veins of both wings sometimes slightly darker between veins; sexes not overly dissimilar in abdominal length ( Figs 69, 70 )............................................................................................................. Udea galapagensis sp. n.

Remarks: I have identified Galápagos specimens by comparing them with two syntypes from Cayenne, French Guiana and ‘Canad?’ without abdomen in the BMNH. The species was described from 15 specimens and I concur with Munroe (1995: 167) to refrain from designating a lectotype as better syntypes may turn up. The species is known as the Tropical sod webworm ( Heppner, 2003).

Figures

Figs 41 - 48. Neotropical specimens of Spilomelinae, size not to scale. (41, 42) Omiodes indicata. (41) ♂, Galápagos, Tagus Cove, emerged ex pupa on Desmodium glabrum, 20 - 25. v. 1992, MHNG. (42) ♀, Galápagos, Pinta, ± 50 m, 20. iii. 1992, MHNG. (43) Patania silicalis ♀, Galápagos, Alcedo, NE slope, 292 m, 30. iii. 2004, MHNG. (44, 45) Penestola bufalis. (44) ♂, Galápagos, Santiago, Cerro Inn, 28. iii. 1992, MHNG. (45) ♀, Galápagos, Santa Cruz, CDRS, wall of Inverts. lab., 11 m, 6. iv. 2004, MHNG. (46, 47) Pilocrocis ramentalis. (46) ♂, Venezuela, Aragua, PN H. - Pittier, Estacion Rancho Grande, 1100 m, 15. vii. 2009, MHNG. (47) ♀, Galápagos, Santa Cruz, E. C. C. D., 6. iii. 1992, MHNG. (48) Psara chathamalis ♀, Galápagos, Santa Cruz, Bahía Conway, 14. iv. 1992, MHNG.

Figs 49 - 56. Neotropical specimens of Spilomelinae, size not to scale. (49, 50) Rhectocraspeda periusalis. (49) ♂, Galápagos, Santiago, Central, 700 m, 9. iv. 1992, MHNG. (50) ♀, Galápagos, Isabela, ± 15 km N Pto Villamil, 25. v. 1992, MHNG. (51, 52) Salbia haemorrhoidalis. (51) ♂, Galápagos, San Cristóbal, antiguo botadero, ca. 4 km SE Pto Baquerizo, 169 m, 22. ii. 2005, MHNG. (52) ♂, Ecuador, Pichincha, Septimo Paraiso Reserve, ± 1300 m, 8. v. 2002, MHNG. (53, 54) Samea castellalis. (53) ♀, Galápagos, Santiago, close to Caseta, 686 m, 6. iii. 2005, MHNG. (54) ♂, Galápagos, San Cristóbal, pampa zone, 15. ii. 1989, CNC. (55, 56) Samea coffea. (55) ♀ holotype, CNC. (56) ♂ paratype, Galápagos, San Cristóbal, 4 km SE Pto Baquerizo, 12. ii. 1989, CNC.

Figs 17 - 24. Neotropical specimens of Spilomelinae, size not to scale. (17) Diaphania nitidalis ♀, Brazil, Bahía, Camacan, Res. Serra Bonita, 800 m, 8. iv. 2011, MHNG. (18) Ercta vittata ♂, Galápagos, Marchena, 12. iii. 1992, MHNG. (19) Herpetogramma bipunctalis ♂, Galápagos, Santa Cruz, CDRS, Barranco, 20 m, 30. iv. 2002, MHNG. (20, 21) Herpetogramma phaeopteralis. (20) ♂, Galápagos, Isabela, NE slope Alcedo, Los Guayabillos camp, 869 m, 2. iv. 2004, MHNG. (21) ♀, Galápagos, San Cristóbal, near La Loberia, sea level, 24. ii. 2005, MHNG. (22, 23) Hymenia perspectalis. (22) ♀, Galápagos, Santa Cruz, Los Gemelos, 4. v. 2002, MHNG. (23) ♀, same locality, 27. v. 1992, MHNG. (24) Lineodes integra ♂, Pinta, Cabo Ibbetson, 8 m, MHNG.

Figs 65 - 72. Neotropical specimens of Spilomelinae, size not to scale. (65, 66) Synclera jarbusalis. (65) ♂, Brazil, Bahía, Camacan, Res. Serra Bonita, 800 m, 8. iv. 2011, MHNG. (66) ♀, Venezuela, Aragua, PN H. - Pittier, Paso Portachuelo, 1136 m, 21. vii. 2009, MHNG. (67, 68) Terastia meticulosalis. (67) ♀, Galápagos, Santa Cruz, casa L. Roque-Albelo & V. Cruz, 137 m, 27. ii. 2005, MHNG. (68) ♂, Brazil, Mato Grosso, environs de Dourados, iii. 1966, MHNG. (69, 70) Udea galapagensis. (69) ♂ holotype, MHNG. (70) paratype ♀, Galápagos, San Cristóbal, El Junco, east side, 654 m, 25. ii. 2005, MHNG. (71, 72) Udea sideralis. (71) holotype ♂, MHNG. (72) paratype ♀, Santiago, N side, 527 m, 5. iii. 2005, MHNG.