Verbascum faurei subsp. acanthifolium (Pau) Benedi & J.M.Monts., Lagascalia 20: 169. 1997.

Khamar, Hamid, Oualidi, Jalal El, Touhami, Amina Ouazzani & Civeyrel, Laure, 2023, Nomenclature and typification in Verbascum (Scrophulariaceae) from North Africa, PhytoKeys 225, pp. 115-152 : 115

Verbascum faurei subsp. acanthifolium (Pau) Benedi & J.M.Monts., Lagascalia 20: 169. 1997.

Celsia acanthifolia ≡ Celsia acanthifolia Pau in Font Quer. Iter Maroc. n° 566. 1927. Type: [Morocco]. pr. Badú (Atlante rhiphaeo); Hab. in saxosis, solo schistose, 6 July 1927, P. Font Quer, 566 (lectotype, selected by Benedí and Montserrat 1997, pg. 169, first step “type”, as " Font Quer 566-1927 "; second step, designated here: BC [BC43694]!; isolectotypes: LD [LD1222710]!, GDA [GDA39155]!, G [G00015117]!, MPU [MPU009627, MPU009628]!, BCN [BCN18029]!, BM [BM000930561]!). [image available at http://psimg.jstor.org/fsi/img/pdf/i0/10.5555/al.ap.specimen.bc43694_normal.pdf]

Celsia acanthifolia ≡ Celsia acanthifolia Pau, in Cavanillesia 1: 47. 1928, nom. nud.

Celsia faurei ≡ Celsia faurei var. acanthifolia (Pau) Maire in Jahand. & Maire. Cat. PI. Maroc.3: 669 (1934).

Notes.

Font Quer (1927) based his species on a collection of plants from the Rif region that he has gathered during his 1927 campaign in Morocco. Font Quer gave the following diagnosis "Folia Acanthi mollis L., superiora oblonga, sinuato-dentata, floribus longe pedunculatis, pedicellis glanduliferis, racemosis; bractae lanceolatae brevissimae, calycis laciniae lanceolatae, margine integro, capsula globosa, obtusa calyce triplo longior" with the collection locality "hab. in saxosis, pr. Badù (Atlante rhiphaeo), 1500 m. alt., solo schistoso, 6 julii". This protologue has been validly published by Gonzâlez-Bueno (1988). According to Nualart (2017), Benedí and Montserrat (1997) wrongly cited the collection number of the lectotype as “BC436934” instead of “BC43694”. So, in addition to the specimen kept at Herb. BC (BC43694), seven other specimens are in agreement with the description in the protologue. They are housed at Herbs LD (LD1222710), GDA (GDA39155), G (G00015117), MPU (MPU009627, MPU009628), BCN (BCN18029), and BM (BM000930561). According to Art. 9.6 of the ICN ( Turland et al. 2018), all of these specimens and those persevered at Herb. BC must be treated as syntypes. The designation by Benedí and Montserrat (1997) can be considered here as a first-step typification (see Art. 9.17 of the ICN, Turland et al. 2018). As the herbarium specimen BC43694 [mounted on three sheets]) kept at Herb. BC is the one showing the best quality of preservation of morphological features described in the protologue, we select it here as second-step lectotype of the name.

The taxonomic status of Celsia acanthifolia has been assessed by some authors. Maire (in Jahandiez and Maire 1934) regarded it as a variety under Ceslia faurei Murb. [ C. faurei var. acanthifolia (Pau) Maire]. Later, Benedi and Montserrat (1997) raised it to the rank of subspecies and they published it as a new combination, Verbascum faurei (Murb.) Hub.-Morath subsp. acanthifolium (Pau) Benedí & J.M.Monts.