Exiligada longicauda, Criscione & Law & Köhler, 2012

Criscione, Francesco, Law, Margot Louisa & Köhler, Frank, 2012, Land snail diversity in the monsoon tropics of Northern Australia: revision of the genus Exiligada Iredale, 1939 (Mollusca: Pulmonata: Camaenidae), with description of 13 new species, Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 166 (4), pp. 689-722 : 704-705

EXILIGADA LONGICAUDA SP. NOV.

( FIGS 5G , 14A–C , 15 )

Holotype: AUSTRALIA, NT, VRD, 7 km east of Kirkimbie Station H /S, top of the cliffs along the stream, well-developed spinifex, 17°43 ′ 34 ′′ S, 129°18 ′ 36 ′′ E (coll. V. Kessner, 2.ix.2009; WA-84 /09) ( WAM S83169).

Paratypes: AUSTRALIA, NT, VRD, same data as for holotype, one wet ( WAM S49228 ); east of Duncan Hwy , 37.7 km north-west of Kirkimbie Station H /S ,

Remarks: Only two shells examined. Shell colour pattern very similar to shells of four congeners; to be distinguished from E. nodulicauda and E. negriensis by reduced number of spiral bands, from E. pallida by darker background, and from E. punctata by absence of maculations. Along with E. nodulicauda , only species with a coiled penis.

Penis length relative to width (pl/pw) almost two times the average and largest amongst congeners; epiphallus length relative to penis (el/pl) one third of average and smallest amongst congeners. Inner penial wall sculpture unique by consisting exclusively of tubercles. Vagina length relative to penis (vl/pl) larger than average and almost as large as in E. nodulicauda ; vagina length relative to free oviduct (vl/ ol) largest amongst congeners.

Figures

Figure 5. Shells of Exiligada species. A, E. negriensis (holotype AM C.64865). B, E. brabyi sp. nov. (paratype C.443279). C, E. calciphila sp. nov. (paratype C.462723). D, E. floraevallis sp. nov. (paratype WAM S49204). E, E. gregoriana sp. nov. (paratype AM C.462820). F, E. limbunya sp. nov. (paratype AM C.470205). G, E. longicauda sp. nov. (paratype WAM S49228). H, E. monochroma sp. nov. (paratype AM C.469772).

Figure 14. SEM micrographs of Exiligada shells. A-C, E. longicauda sp. nov. (paratype WAM S 49228). A, Apical view showing protoconch and first whorl. B, Close-up from above of sculpture across entire shell. C, Microsculpture on teleoconch whorl. D-F, E. monochroma sp. nov. (paratype AM C.462772). D, Apical view showing protoconch and first whorl. E, Close-up from above of sculpture across entire shell. F, Microsculpture on teleoconch whorl. Scale bars: A-B, D-E = 1 mm, C, F = 0.5 mm.

Figure 15. Penial anatomy of E. longicauda sp. nov. (holotype WAM S83169). A. Detail of apical portion of inner penial wall. B. Detail of basal portion of inner penial wall. Scale bars: 5 mm; A, B = 1 mm.