Exiligada monochroma, Criscione & Law & Köhler, 2012

Criscione, Francesco, Law, Margot Louisa & Köhler, Frank, 2012, Land snail diversity in the monsoon tropics of Northern Australia: revision of the genus Exiligada Iredale, 1939 (Mollusca: Pulmonata: Camaenidae), with description of 13 new species, Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 166 (4), pp. 689-722 : 705

publication ID	https://doi.org/ 10.1111/j.1096-3642.2012.00863.x
publication LSID	lsid:zoobank.org:pub:AC086999-76A8-4F93-9A51-5B5B213A50E6
DOI	https://doi.org/10.5281/zenodo.10544648
persistent identifier	https://treatment.plazi.org/id/4C0EBAD8-878D-43A4-9E12-9AE34E06B58B
taxon LSID	lsid:zoobank.org:act:4C0EBAD8-878D-43A4-9E12-9AE34E06B58B
treatment provided by	Marcus
scientific name	Exiligada monochroma
status	sp. nov.

Treatment

EXILIGADA MONOCHROMA View in CoL SP. NOV.

( FIGS 4F View Figure 4 , 5H View Figure 5 , 14D–F View Figure 14 , 16 View Figure 16 )

Holotype: AUSTRALIA, NT, VRD, 46 km south-west of VRD Station H /S, limestone outcrop, 16°41 ′ 22 ′′ S, 130°42 ′ 12 ′′ E (coll. V. Kessner, 16.vii.2008) ( AM C.475768). GoogleMaps

large, open and very rugged limestone formation, open limestone formation, in talus, under spinifex (coll. V. Kessner, R. Hokkanen, 2.ix.2009), one wet ( WAM S83170) .

Etymology: In reference to the remarkable length of the penis, derived from ‘longus’ (Latin = long) and ‘cauda’ (Latin = tail, penis), adjective of feminine gender.

Description: Shell ( Figs 5G View Figure 5 , 14A–C View Figure 14 ). Large, moderately elevated, moderately thick; protoconch and teleoconch whorls moderately strong radial growth lines, micropustulations absent. Shell background colour yellowish horn fading to whitish towards shell base, with few brown-reddish spiral bands, equally conspicuous on entire shell, covering whorl periphery to shell base, often interrupted radially; no maculations.

Genitalia ( Fig. 15 View Figure 15 ). Epiphallus one third as long as penis. Vas deferens entering penial sheath halfway up. Penis 12 times longer than wide, heavily coiled inside sheath, posterior part of inner wall supporting many irregular tubercles; anterior part with narrow, regularly shaped and spaced longitudinal pilasters. Vagina almost as long as penis and more than two times as long as free oviduct. Bursa copulatrix with short, rounded, well-differentiated end.

Paratypes: AUSTRALIA, NT, VRD, same data as for holotype, 31 dry, two wet ( AM C.462772) GoogleMaps .

Etymology: In reference to the coloration, derived from ‘monos’ (Greek = unique) and ‘chroma’ (Greek = colour), adjective of feminine gender.

Description: Shell ( Figs 4F View Figure 4 , 5H View Figure 5 , 14D–F View Figure 14 ). Moderately large, moderately elevated, thin; micropustulations absent. Shell colour brown horn fading to whitish towards shell base; no maculations.

Genitalia ( Fig. 16 View Figure 16 ). Epiphallus as long as penis. Vas deferens entering penial sheath halfway up. Penis five times longer than wide, not coiled inside sheath, with inner wall supporting three moderately large, regularly spaced longitudinal pilasters running along its entire length. Vagina slightly longer than penis and one and a half times longer than free oviduct, bursa copulatrix with short, rounded well-differentiated end.

Remarks: Teleoconch microsculpture differing from that of E. montejinni by absence of micropustulations. Spiral bands almost always absent; readily distinguished from E. montejinni by its darker brownish horn background colour.

Penis length relative to width (pl/pw) average, longer than in E. montejinni ; epiphallus length relative to penis (el/pl) average and equal to E. montejinni . Inner penial wall sculpture similar to that of E. montejinni , but with more organized pattern. Vagina length relative to penis (vl/pl) and vagina length relative to free oviduct (vl/ol) larger than average and E. montejinni .