Leiodes koreana, Park & Ahn, 2007

Hoshina, Hideto, 2012, Review of the tribes Sogdini and Leiodini from Japan and North Chishima Islands. Part II. Genera Hydnobius and Leiodes (Coleoptera: Leiodidae), Acta Entomologica Musei Nationalis Pragae (suppl. 1) 52, pp. 1-168 : 49-50

Leiodes koreana species group

Species included. Leiodes koreana Park & Ahn, 2007 , L. masatsugui sp. nov., L. toyoshimai sp. nov.

Diagnosis. Elytra unicolor, with sparsely arranged large punctures between rows of punctures ( Figs. 27D , 30D , 33E ); mesoventrite with or without excavation between median carina and transverse carina ( Figs. 27E , 30E , 33F ); tarsomeres 2–4 of male protarsi expanded ( Figs. 28A , 31A , 34A ); mesotibiae without sexual dimorphism, simply square at interoapical corner ( L. koreana and L. masatsugui sp. nov.) or sexually dimorphic, with male mesotibiae more widening from base to apex than in female (in L. toyoshimai sp. nov, Figs. 34C, D ); dorsal posteroapical projection of metafemora small and not showing sexual dimorphism ( Figs. 28G, 28H , 31H, 31I , 34I, 34J ); pubescence of metaventrite without or with indistinct sexual dimorphism ( Figs. 33G, 33H ); aedeagus robust, bearing some large sclerites in inner sac ( Figs. 29A , 32A , 35A ); male abdominal sternite 8 weakly to strongly curved ( Figs. 29C , 32C , 35D ); female abdominal sternite 8 with a bifurcate projection ( Fig. 29D ) or two projections distant from each other at anterior margin ( Figs. 32D , 35E ).

Differential diagnosis. The Leiodes koreana species group can be separated from other species groups by having a robust aedeagus with large sclerites in the inner sac, and female abdominal sternite 8 with a bifurcate projection or two projections distant from each other at the anterior margin.

Note. Female abdominal sternite 8 of most of species of Palaearctic Leiodes has not been examined until now. Hence, I cannot exclude that additional Palaearctic species also having females with similar abdominal sternite 8’s and belonging to the L. koreana group.

Figures

Fig. 27. Leiodes koreana Park & Ahn, 2007.A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for E.

Fig.30. Leiodes masatsugui sp. nov.A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E.

Fig. 33. Leiodes toyoshimai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – basal margin of pronotum; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for F; IV: 0.5 mm for G and H.

Fig. 28. Leiodes koreana Park & Ahn, 2007. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male hind leg, ventral view; D – female hind leg, ventral view; E – male protibia, dorsal view; F – female protibia, dorsal view; G – male metafemur, dorsal view; H – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 0.2 mm for E and F; III: 0.2 mm for G and H.

Fig. 31. Leiodes masatsugui sp. nov.A – male fore leg, ventral view; B – female fore leg, ventral view; C – hind leg of male (body size: 4 mm), ventral view; D – ditto (body size: 3.2 mm); E – female hind leg, ventral view; F – male protibia, dorsal view; G – female protibia, dorsal view; H – male metafemur, dorsal view; I – female metafemur, dorsal view. Scale I: 1 mm for A–C and E; II: 1 mm for D; III: 0.2 mm for F and G; IV: 0.2 mm for H and I.

Fig. 34. Leiodes toyoshimai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J.

Fig. 29. Leiodes koreana Park & Ahn, 2007.A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D; III: 0.1 mm for E.

Fig. 32. Leiodes masatsugui sp. nov.A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D; III: 0.1 mm for E.

Fig. 35. Leiodes toyoshimai sp. nov.A – aedeagus, dorsal view; B – ditto, lateral view; C – apex of aedeagus, ventral view; D – male abdominal sternite 8; E – female abdominal sternite 8; F – coxite and stylus. Scale I: 0.5 mm for A–C; II: 0.2 mm for D and E; III: 0.1 mm for F.