Anhoraeomorphus globosus (Franz) Jałoszyński, 2024

Jałoszyński, Paweł, 2024, The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 5505 (1), pp. 1-96 : 33-35

Anhoraeomorphus globosus (Franz) , comb. n.

( Figs 119–126 )

Horaeomorphus globosus Franz, 1986b: 169 .

Type material studied. Holotype (Madagascar): ♂, five labels: “Madagascar-Centre / Forét Imaitso / Anjavidilava 2030m / Andingitra-Ambalavao / 19.1. 58 P. Grivaud” [white, printed], “INSTITUT / SCIENTIFIQUE / MADAGASCAR ” [bluish, printed], “Hinterhüften / schmal getrennt {i.e., hind coxae narrowly separated}” [yellow, handwritten], “ Horaeomorphus / globosus m. / det. H.Franz ” [yellowish, handwritten and printed], “Typus” [red, handwritten] ( NHMW).

Revised diagnosis. Body ( Fig. 119 ) slender; head ( Figs 120–121 ) oval, with vertex posteriorly broadly rounded; pronotum ( Fig. 119 ) bell-shaped, broadest near anterior third, with one pair of large and deep antebasal pits; antennomeres 3–6 ( Fig. 122 ) each elongate, 7 about as long as broad, 8–10 each slightly transverse; profemora slightly thicker than meso- and metafemora; hind tibiae in male unmodified; aedeagus ( Figs 123–126 ) in ventral view broadest in sub-basal third, apical region of dorsal wall in lateral view abruptly and strongly bent dorsally, distolateral endophallic sclerites broadly separated, each elongate and with blunt subtriangular apex, each paramere with group of several short apical setae.

Redescription. Body of male ( Fig. 119 ) strongly convex, dark brown, tarsi and palps slightly lighter; setae light brown; BL 2.53 mm.

Head ( Fig. 120 ) in anterodorsal view oval, broadest at eyes, HL 0.50 mm, HW 0.45 mm; temple in lateral view ( Fig. 121 ) slightly more than twice as long as eye; vertex and frons confluent, together weakly convex and slightly elongate; vertex posteriorly broadly rounded; each eye conspicuously small and weakly convex, nearly circular, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna ( Fig. 122 ) slender but short, as long as half BL, AnL 1.25 mm, antennomeres 1–6 each elongate, 7 about as long as broad, 8–10 each slightly transverse, 11 as long as 9 and 10 combined, slightly wider than 10, about 1.8 times as long as broad.

Pronotum bell-shaped, broadest near anterior third; PL 0.70 mm, PW 0.58 mm. Anterior margin strongly rounded, lateral margins rounded in anterior half, weakly sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of conspicuously large and deep pits, sublateral carinae short but distinct. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.

Elytra together oval, broadest slightly in front of middle; EL 1.33 mm, EW 1.03 mm, EI 1.29; humeral calli not marked, basal elytral foveae indiscernible. Punctures dense but superficial and diffuse, inconspicuous; setae sparse, short and nearly recumbent.

Legs moderately long, slender, unmodified except tibiae: protibiae with distal region curved mesally and with large distomesal concave patch of dense adhesive setae; mesotibiae with distal region curved mesally; and metatibiae indistinctly recurved.

Aedeagus ( Figs 123–126 ) stout, AeL 0.45 mm; in ventral view pear-shaped, broadest in sub-basal third, distally gradually narrowing to truncate apex with slightly arcuate distal margin; in lateral view apical region of dorsal wall strongly bent dorsally; endophallus with asymmetrical and large distolateral sclerites and with strongly asymmetrical assemblage of median sclerotized structures, distolateral sclerites broadly separated, each with subtriangular and blunt apical region; parameres broad and their apices not projecting beyond aedeagal apex, each paramere with apical group of 4–5 short setae.

Female. Unknown.

Distribution. Southern Madagascar (Matsiatra Ambony Region).

Remarks. Franz (1986b) misidentified the holotype male as a female.

Anhoraeomorphus globosus can be easily distinguished from all remaining congeners by the body shape, tempora nearly twice as long as small eyes, slender legs with unmodified metatibiae, and unique structures of the aedeagus.

Figures

FIGURES 119–126. Anhoraeomorphus globosus (Franz), holotype. Habitus in dorsal view (119); head in anterodorsal view (120); head and prothorax in lateral view (only bristles on head are shown) (121); antenna (122); aedeagus in ventral (123, 125) and lateral (124, 126) views.