Trogophloeus segnis, Erichson, 1840

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1067-1806 : 1067-1806

publication ID

https://doi.org/ 10.1206/0003-0090.265.1.1

persistent identifier

https://treatment.plazi.org/id/03E64505-FD10-5D90-FF0E-A287FDAE96C9

treatment provided by

Felipe

scientific name

Trogophloeus segnis
status

 

segnis Erichson, 1840 View in CoL , see: Thinodromus .

semilividus Bernhauer, 1943: 73 ( Trogophloeus ; subgenus Boopinus ; Type locality: Arabien: Ahwar).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: Saudi Arabia.

semiopacus Cameron, 1928: 13 ( Trogophloeus ; Type locality: Belgian Congo: Bikoro; Irebu). — Scheerpeltz, 1933: 1088 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: Congo.

semirudis Bernhauer, 1942c: 349 ( Trogophloeus ; subgenus Taenosoma ; Type locality:

Mont Cameroun, 1800–2000 m, versant Sud Est).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: Cameroon.

senex Fauvel, 1867 , see: Teropalpus .

senilis Sharp, 1880: 51 ( Trogophloeus ; Type locality: Oahu).

— Blackburn and Sharp, 1885: 224 ( Trogophloeus ; habitat; Hawaii).

— Fauvel, 1901: 71 ( Trogophloeus ; synonym of fulvipes ).

— Sharp, 1908: 543 ( Trogophloeus ; notes; Hawaii).

— Bernhauer and Schubert, 1911: 104 ( Trogophloeus ; catalog).

— Blackwelder, 1943: 62 ( Carpelimus ; synonym of fulvipes ).

— Herman, 1970: 393 ( Carpelimus ).

— Moore, 1975c: 105 ( Carpelimus ; Hawaii).

- DISTRIBUTION: Hawaii.

sericatus Sharp, 1889 , see: Thinodromus .

sericeipennis Bernhauer, 1938d: 326 ( Trogophloeus ; Type locality: Belgisches Kongo:

Pied du Ruwenzori).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: Congo.

sericeus Cameron, 1923: 392 ( Trogophloeus ; subgenus Boopinus ; Type locality: Jamaica).

— Scheerpeltz, 1933: 1088 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Blackwelder, 1943: 63 ( Carpelimus ; characters; Cuba; Jamaica; St. Croix).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: West Indies.

sextagonalis Bierig, 1935: 10 ( Trogophloeus ; Type locality: Caimito del Guayabal; Laguna La Canoa; Sierra del Rosario , cercania de Cayajabos, Aspiro; Rio Almendares).

— Blackwelder, 1943: 63 ( Carpelimus ; synonym of sericeus ).

sextagonalis Bierig, 1935 , see: sericeus Cameron, 1923 .

seydeli Cameron, 1952a: 324 ( Trogophloeus ; subgenus Troginus Type locality: Belgian Congo: Elisabethville).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: Congo.

sharpi Bernhauer, 1909c: 230 ( Trogophloeus ; replacement name for punctipennis Sharp ).

— Bernhauer, 1910b: 355 ( Trogophloeus ; cited again as a new replacement name for punctipennis Sharp and without reference to the earlier replacement).

— Bernhauer and Schubert, 1911: 104 ( Trogophloeus ; catalog).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: Guatemala.

punctipennis Sharp, 1887: 701 ( Trogophloeus ; [preoccupied]; Type locality: Guatemala, San Geronimo ).

— Bernhauer, 1909c: 230 ( Trogophloeus ; synonym of sharpi ).

— Bernhauer and Schubert, 1911: 104 ( Trogophloeus ; synonym of sharpi ). sharpianus Cameron, 1933a: 169 ( Trogophloeus ; subgenus Boopinus ; Type locality: Japan: Kobe, Harada).

— Cameron, 1949b: 460 ( Trogophloeus ; Japan).

— Herman, 1970: 393 ( Carpelimus ).

— Watanabe, 1998: 313 ( Carpelimus ; Japan).

- DISTRIBUTION: Japan.

siamensis Fauvel, 1886b: 144 ( Trogophloeus ; Type locality: Manille; Siam, Bangkok; Sumatra: Kaju Tanam , Singkara ; Java : Batavia ).

— Fauvel, 1907: 14 ( Trogophloeus ; Madagascar).

— Bernhauer and Schubert, 1911: 104 ( Trogophloeus ; catalog).

— Bernhauer, 1920: 5 ( Trogophloeus ; Australia).

— Cameron, 1927a: 22 ( Trogophloeus ; Samoa).

— Cameron, 1928d: 99 ( Trogophloeus ; Sumatra).

— Bernhauer, 1928: 5 ( Trogophloeus ; Sumatra).

— Cameron, 1930: 187 ( Trogophloeus ; characters; Bihar).

— Scheerpeltz, 1933: 1088 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Cameron, 1936: 4 ( Trogophloeus ; Society Islands).

— Cameron, 1949b: 461 ( Trogophloeus ; Japan).

— Scheerpeltz, 1954: 110 ( Trogophloeus ; subgenus Boopinus ; Ceylon).

— Scheerpeltz, 1957c: 224 ( Trogophloeus ; subgenus Boopinus ; Sumba).

— Herman, 1970: 393 ( Carpelimus ).

— Watanabe and Shibata, 1972: 60 ( Carpelimus ; Yaku-shima Island).

— Coiffait, 1976: 220 ( Trogophloeus ; Tahiti).

- DISTRIBUTION: Madagascar, India, Thailand, Indonesia, Australia, Samoa, Society Islands, Japan.

siwalikensis Cameron, 1930: 187 ( Trogophloeus ; variety of siamensis ; Type locality: Siwaliks).

— Scheerpeltz, 1933: 1088 ( Trogophloeus ; synonym of siamensis ).

siculus Mulsant and Rey, 1878: 733 ( Trogophloeus ; Type locality: Sicile).

— Klima, 1904: 48, 60 ( Trogophloeus ; subgenus Taenosoma ; characters; Sicily; Corfu).

— Bernhauer and Schubert, 1911: 104 ( Trogophloeus ; catalog).

— Herman, 1970: 393 ( Carpelimus ).

— Binaghi, 1974: 50 ( Trogophloeus ; characters; ecological notes).

— Sparacio, 1995: 144 ( Trogophloeus ; characters; notes; Sicily).

— Ciceroni and Zanetti, 1995: 11 ( Carpelimus ; Italy).

- DISTRIBUTION: Sicily, Corfu.

signatoides Scheerpeltz, 1972: 69 ( Trogophloeus ; subgenus Trogophloeus ; Type locality: Argentinien: Rio Negro, El Bolson, Mt. Piltriquitron, 720 m).

- DISTRIBUTION: Argentina.

signatus Erichson, 1834 , see: Thinodromus .

silvestris Cameron, 1918: 61 ( Trogophloeus ; Type locality: Singapore: Bukit Timah ; Mandai; Sembawang).

— Cameron, 1921a: 364, 400 ( Trogophloeus ; characters; Singapore).

— Scheerpeltz, 1933: 1088 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Scheerpeltz, 1957c: 224 ( Trogophloeus ; subgenus Taenosoma ; Sumba).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: Singapore, Indonesia.

similis Smetana, 1967e: 310 ( Trogophloeus ; subgenus Trogophloeus ; Type locality: Ost-Slowakei, Zboj. Also cited from Turkey).

— Herman, 1970: 393 ( Carpelimus ).

— Lohse, 1978: 9, 10 ( Carpelimus ; characters).

— Lohse and Lucht, 1989: 131 ( Carpelimus ; characters; Turkey; Italy; France; Czechoslovakia;

Switzerland; Germany).

— Boháč, 1993: 42 ( Carpelimus ; subgenus Boopinus ; Slovakia; Czech Republic).

— Allen, 1994b: 115 ( Carpelimus ; notes; England).

— Hodge and Jones, 1995: 32 ( Carpelimus ; Britain).

— Gildenkov, 1995: 11 ( Carpelimus ; characters; Russia).

— Ciceroni and Zanetti, 1995: 11 ( Carpelimus ; may be synonym of augustae ; Italy).

— Gildenkov, 1997b: 555, 557 ( Carpelimus ; characters; Russia).

— Gildenkov, 1998a: 53 ( Carpelimus ; Ukraine).

— Bruge, 1999: 21 ( Carpelimus ; subgenus Carpelimus ; notes; Turkey; Italy; Czech Republic; Slovakia; Germany; Switzerland; France; Belgium).

- DISTRIBUTION: Europe, Russia, Turkey.

simplarius LeConte, 1877: 244 ( Trogophloeus ; Type locality: Coney Island, near New York). — Casey, 1889: 373 ( Trogophloeus ; notes).

— Casey, 1895: 450 ( Trogophloeus ; characters).

— Bernhauer and Schubert, 1911: 104 ( Trogophloeus ; catalog).

— Herman, 1970: 393 ( Carpelimus ).

— Downie and Arnett, 1996: 445 ( Carpelimus ; characters; USA).

- DISTRIBUTION: USA.

simplex Motschulsky, 1857a: 505 ( Trogophloeus ; Type locality: Indes orientales).

— Kraatz, 1859: 180 ( Trogophloeus ; characters; Ceylon; India orientalis).

— Fauvel, 1878g: 490 ( Trogophloeus ; characters; Australia; Celebes; East Indies; Ceylon).

— Fauvel, 1886b: 145 ( Trogophloeus ; Philippines; Ceylon; East Indies; Annam; Java; Celebes; Australia).

— Bernhauer, 1902b: 44 ( Trogophloeus ; Ceylon).

— Bernhauer and Schubert, 1911: 104 ( Trogophloeus ; catalog).

— Cameron, 1928d: 99 ( Trogophloeus ; Sumatra).

— Cameron, 1930: 189 ( Trogophloeus ; characters; Ceylon).

— Scheerpeltz, 1933: 1088 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Scheerpeltz, 1960: 2 ( Trogophloeus ; subgenus Paraboopinus ; Pakistan; Iran; Afghanistan).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: Iran, Afghanistan, Pakistan, Sri Lanka, Vietnam, Indonesia, Philippines Australia.

simplicicollis Wollaston, 1857 , see: halophilus Kiesenwetter, 1844 .

singaporensis Herman, 2001: 42 ( Carpelimus ; replacement name for littoralis Cameron ).

- DISTRIBUTION: Singapore.

littoralis Cameron, 1918: 63 ( Trogophloeus ; subgenus Taenosoma ; [preoccupied]; Type locality: Singapore: Pasir Panjang).

— Cameron, 1921a: 365, 400 ( Trogophloeus ; characters; Singapore).

— Scheerpeltz, 1933: 1085 ( Trogophloeus ; subgenus Trogophloeus ; cited as litoralis ; catalog). — Herman, 1970: 392 ( Carpelimus ).

— Herman, 2001: 42 ( Carpelimus ; synonym of singaporensis ).

singularis Fauvel, 1907 , see: Apocellagria .

siwalikensis Cameron, 1930 , see: siamensis Fauvel, 1886 .

skottsbergii Bernhauer, 1921 , see: Teropalpus .

smithi Bernhauer, 1909 , see: Thinodromus .

smithi Cameron, 1913 , see: Thinodromus .

smithianus Scheerpeltz, 1933, see: Thinodromus .

sobrinus Fairmaire and Germain, 1861 , see: bilineatus Stephens, 1834 .

socius Bernhauer, 1904 , see: Thinodromus .

sodalis Bernhauer, 1933e: 329 ( Trogophloeus ; Type locality: Argentinien: Iliar. La Rioja ).

— Herman, 1970: 393 ( Carpelimus ).

sordidus Cameron, 1923: 394 ( Trogophloeus ; subgenus Taenosoma ; Type locality:

Grand Cayman).

— Scheerpeltz, 1933: 1088 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Blackwelder, 1943: 84 ( Carpelimus ; characters; Grand Cayman).

— Blackwelder, 1947: 118 ( Carpelimus ; Cayman Islands).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: West Indies.

spectabilis Bernhauer, 1934 , see: Thinodromus .

spectatus Casey, 1889 , see: obesus Kiesenwetter, 1844 .

spinicollis Rye, 1870 , see: Thinodromus .

splendidus Bernhauer, 1915 , see: Thinodromus .

spretus Casey, 1889 , see: rivularis Motschulsky, 1860 .

strandi Bernhauer, 1934d: 148 ( Trogophloeus ; Type locality: Brasilien: Matto Grosso , Corumba).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: Brazil.

strandi Scheerpeltz, 1950 , see: subtilicornis Roubal, 1946 .

stricticollis Bernhauer, 1923 , see: richteri Bernhauer, 1927 .

stricticollis Fairmaire and Germain, 1861: 449 ( Trogophloeus ; Type locality: Forêts subandines de Chillan).

— Fauvel, 1867a: 34 [= 1868: 39] ( Trogophloeus ; characters; Chile).

— Bernhauer and Schubert, 1911: 104 ( Trogophloeus ; catalog).

— Coiffait and Saiz, 1968: 444 ( Trogophloeus ; subgenus Trogophloeus ; characters; Chile).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: Chile.

subaequus Mulsant and Rey, 1878, see: rivularis Motschulsky, 1860 .

subangulatus Jarrige, 1964 , see: halophilus Kiesenwetter, 1844 .

subdenticulatus Bernhauer, 1908d: 289 ( Trogophloeus ; Type locality: Paraguay).

— Bernhauer and Schubert, 1911: 104 ( Trogophloeus ; catalog).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: Paraguay.

subpolitus Coiffait, 1982: 158 ( Trogophloeus ; subgenus Boopinus ; Type locality: Nepal: Siklis, 2150 m).

- DISTRIBUTION: Nepal.

subrugulosus Rey, 1878, see: halophilus Kiesenwetter, 1844 .

subterraneus Smetana, 1960a: 154 ( Trogophloeus ; subgenus Myopinus ; Type locality: Nordost-Böhmen, Klášter n. Děd.).

— Horion, 1963: 213 ( Trogophloeus ; subgenus Myopinus ; Czech Republic).

— Lohse, 1964: 78 ( Trogophloeus ; subgenus Myopinus ; characters; central Europe).

— Herman, 1970: 393 ( Carpelimus ; cited erroneously as terraneus on page 394).

— Lucht, 1987: 92 ( Trogophloeus ; checklist; Germany; Czechoslovakia).

— Boháč, 1993: 42 ( Carpelimus ; subgenus Myopinus ; Czech Republic).

- DISTRIBUTION: Czechoslovakia.

subtilepunctatus Scheerpeltz, 1972: 86 ( Trogophloeus ; subgenus Boopinus ; Type locality: Argentinien: Buenos Aires: Ezeiza).

- DISTRIBUTION: Argentina.

subtilicornis Roubal, 1946: 71 ( Trogophloeus ; Type locality: Hercegovina: Jablanica).

— Steel, 1956c: 265 ( Carpelimus ; characters; England; Scotland).

— Palm, 1961a: 21 ( Trogophloeus ; characters; Sweden; Norway; Finland).

— Horion, 1963: 202 ( Trogophloeus ; subgenus Taenosoma ; Britain; Norway; Sweden; Finland; — Lohse, 1964: 80 ( Trogophloeus ; subgenus Taenosoma ; characters; central Europe).

— Herman, 1970: 393 ( Carpelimus ).

— Pope, 1977: 25 ( Carpelimus ; Britain).

— Zanetti, 1980c: 93, 95 ( Carpelimus ; habitat notes; Italy).

— Frank, 1982: 13 ( Carpelimus ; list of parasites).

— Lucht, 1987: 92 ( Trogophloeus ; checklist; Germany; Czechoslovakia; Switzerland; Benelux).

— Schülke and Uhlig, 1988: 4 ( Carpelimus ; notes; Germany).

— Boháč, 1993: 42 ( Carpelimus ; subgenus Trogophloeus ; Czech Republic).

— Hodge and Jones, 1995: 32 ( Carpelimus ; characters; Britain).

— Gildenkov, 1995: 14 ( Carpelimus ; characters; Russia).

— Ciceroni and Zanetti, 1995: 11 ( Carpelimus ; Italy).

— Gildenkov, 1997b: 555 ( Carpelimus ; Russia).

- DISTRIBUTION: Europe, Russia.

strandi Scheerpeltz, 1950a: 77 ( Trogophloeus ; subgenus Taenosoma ; [preoccupied]; Type locality: Målselv-Gebit Nord-Nordwegens: Rundhaug, Framnes, Slovang, usw.).

— Steel, 1956c: 265 ( Carpelimus ; synonym of subtilicornis ).

— Pope, 1977: 25 ( Carpelimus ; synonym of subtilicornis ).

kerstensi Scheerpeltz, 1954a: 7 View in CoL ([Note: Nomen nudum, published without description (Article 13.1.1)]).

— Steel, 1956c: 265 ( Carpelimus View in CoL ; synonym of subtilicornis View in CoL ).

subtilior Cameron, 1923: 396 ( Trogophloeus ; subgenus Taenosoma ; Type locality: Grenada).

— Scheerpeltz, 1933: 1088 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Blackwelder, 1943: 78 ( Carpelimus ; characters; Jamaica; Grenada).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: West Indies.

subtilis Erichson, 1839a: 606 ( Trogophloeus ; Type locality: Mark Brandenburg).

— Erichson, 1840: 812 ( Trogophloeus ; characters; Germany).

— Redtenbacher, 1849: 828 ( Trogophloeus ; characters; Germany).

— Fairmaire and Laboulbène, 1856: 618 ( Trogophloeus ; characters; France).

— Redtenbacher, 1857: 236 ( Trogophloeus ; characters; Germany).

— Kraatz, 1857d: 881 ( Trogophloeus ; characters; Germany).

— Crotch, 1867a: 385 ( Trogophloeus ; Azores).

— Fauvel, 1871a: 186 [= 1872: 160] ( Trogophloeus ; characters; France; Britain; Germany).

— Redtenbacher, 1874: 258 ( Trogophloeus ; characters).

— LeConte, 1877: 246 ( Trogophloeus ; southern and western States).

— Mulsant and Rey, 1878: 756 ( Trogophloeus ; subgenus Taenosoma ; characters; France).

— Fowler, 1888: 390 ( Trogophloeus ; characters; Britain).

— Ganglbauer, 1895: 657 ( Trogophloeus ; subgenus Taenosoma ; characters; middle Europe).

— Klima, 1904: 49, 65 ( Trogophloeus ; subgenus Taenosoma ; characters; central Europe).

— Reitter, 1909: 174 ( Trogophloeus ; subgenus Taenosoma ; characters; Germany).

— Bernhauer and Schubert, 1911: 104 ( Trogophloeus ; catalog).

— Cameron, 1912a: 50 ( Trogophloeus ; subgenus Taenosoma ; characters in key).

— Johansen, 1914: 578 ( Trogophloeus ; subgenus Taenosoma ; characters; Denmark).

— Notman, 1920a: 180 ( Trogophloeus ; New York).

— Portevin, 1929: 422 ( Trogophloeus ; characters; France).

— Scheerpeltz, 1933: 1088 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Scheerpeltz, 1937: 117 ( Trogophloeus ; subgenus Myopinus ; characters in key; central Europe; North America).

— Bernhauer, 1940c: 2 ( Trogophloeus ; Azores).

— Smetana, 1960a: 157 ( Trogophloeus ; subgenus Myopinus ; characters; Czechoslovakia).

— Horion, 1963: 213 ( Trogophloeus ; subgenus Myopinus ; Finland; Sweden; Norway; Britain; Denmark; Holland; Belgium; France; Switzerland; Italy; Dalmatia; Bosnia; Romania; Czech Republic; Slovakia; Germany; Austria).

— Lohse, 1964: 78 ( Trogophloeus ; subgenus Myopinus ; characters; central Europe).

— Herman, 1970: 393 ( Carpelimus ).

— Smetana 1975c: 164 ( Trogophloeus ; Mongolia).

— Pope, 1977: 25 ( Carpelimus ; Britain).

— Lucht, 1987: 92 ( Trogophloeus ; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux; Denmark; Sweden).

— Kashcheev, 1989: 284 ( Trogophloeus ; subgenus Myopinus ; Mongolia).

— Lohse and Lucht, 1989: 131 ( Carpelimus ; subgenus Taenosoma ; characters).

Borges, 1990: Table II ( Carpelimus ; Azores) .

— Boháč, 1993: 42 ( Carpelimus ; subgenus Myopinus ; Czech Republic; Slovakia).

— Ciceroni and Zanetti, 1995: 11 ( Carpelimus ; Italy).

— Downie and Arnett, 1996: 445 ( Carpelimus ; characters; USA).

— Gildenkov, 1997b: 555 ( Carpelimus ; Russia).

— Eyre, Luff, and Lott, 2000: 32 ( Carpelimus ; Britain).

- DISTRIBUTION: Azores, Europe, Mongolia, USA.

indigens Casey, 1889: 362 ( Trogophloeus ; Type locality: Rhode Island: Newport).

— Bernhauer and Schubert, 1911: 104 ( Trogophloeus ; synonym of subtilis ).

sudanensis Scheerpeltz, 1974c: 5 ( Trogophloeus ; subgenus Carpelimus , cited as Carpalimus ; Type locality: Sudan: Bahr el Ghazal, Wau).

- DISTRIBUTION: Sudan.

suffusus Sharp, 1887: 698 ( Trogophloeus ; Type locality: Guatemala, Paso Antonio ).

— Bernhauer and Schubert, 1911: 104 ( Trogophloeus ; catalog).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: Guatemala.

sulcifrons Coiffait and Saiz, 1968: 437 ( Trogophloeus ; subgenus Paracarpalimus ; Type locality: no locality cited, presumably from Chile).

- DISTRIBUTION: Chile.

sumatrensis Bernhauer, 1915 , see: Thinodromus .

sumbaensis Scheerpeltz, 1957c: 224 , 241 ( Trogophloeus ; subgenus Boopinus ; Type locality: Sumba: Baing).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: Indonesia.

suspectus Mulsant and Rey, 1878 , see: bilineatus Stephens, 1834 .

tagus Herman, 1970: 393 ( Carpelimus ; replacement name for pallidicornis Cameron ).

- DISTRIBUTION: Indonesia.

pallidicornis Cameron, 1945a: 143 ( Trogophloeus ; subgenus Boopinus ; [preoccupied]; Type locality: Java: Tji Solak).

— Herman, 1970: 393 ( Carpelimus ; synonym of tagus ).

tahitiensis Coiffait, 1976: 234 ( Taenosoma ; Type locality: Tahiti: Papenoo).

— Coiffait, 1977a: 171 ( Trogophloeus ; Tahiti).

- DISTRIBUTION: Society Islands.

tantillus Casey, 1889 , see: pusillus Gravenhorst, 1802 .

taprobanae Walker, 1859: 52 ( Trogophloeus ; Type locality: Ceylon).

— Bernhauer and Schubert, 1911: 106 ( Trogophloeus ; subgenus Troginus ; catalog).

— Cameron, 1930: 195 ( Trogophloeus ; characters; Ceylon).

— Scheerpeltz, 1933: 1088 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Herman, 1970: 393 ( Carpelimus ).

- DISTRIBUTION: Sri Lanka.

tarsalis Hochhuth, 1849 , see: obesus Kiesenwetter, 1844 .

tauricus Gildenkov, 1998 , see: troglodytes Erichson, 1840 .

tautus Herman, 1970: 393 ( Carpelimus ; replacement name for rufipennis Eppelsheim ). rufipennis Eppelsheim, 1878a: 422 ( Trogophloeus ; [preoccupied]; Type locality: Griechenland).

— Fauvel, 1902b: 63 ( Trogophloeus ; Algeria; Tunisia).

— Klima, 1904: 48, 62 ( Trogophloeus ; subgenus Taenosoma ; characters; Syria; North Africa).

— Bernhauer and Schubert, 1911: 104 ( Trogophloeus ; catalog).

— Cameron, 1912a: 50 ( Trogophloeus ; subgenus Taenosoma ; characters in key).

— Scheerpeltz, 1933: 1088 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— C. Koch, 1934: 40 ( Trogophloeus ; characters; Egypt).

— C. Koch, 1936b: 136 ( Trogophloeus ; Egypt).

— Herman, 1970: 393 ( Carpelimus ; synonym of tautus ).

temporalis Casey, 1889: 357 ( Trogophloeus ; Type locality: New York; Massachusetts).

— Bernhauer and Schubert, 1911: 104 ( Trogophloeus ; catalog).

— Herman, 1970: 393 ( Carpelimus ).

— Downie and Arnett, 1996: 444 ( Carpelimus ; characters; USA).

- DISTRIBUTION: USA.

tenellus Erichson, 1839 , see: gracilis Mannerheim, 1830 .

tener Bernhauer, 1902a: 249 ( Trogophloeus ; Type locality: Aulie Ata, Turkestan).

— Klima, 1904: 47, 58 ( Trogophloeus ; subgenus Boopinus ; characters; Turkestan).

— Bernhauer and Schubert, 1911: 105 ( Trogophloeus ; catalog).

— C. Koch, 1934: 38 ( Trogophloeus ; characters; Egypt).

— C. Koch, 1936b: 135 ( Trogophloeus ; Egypt).

— Scheerpeltz, 1963b: 426 ( Trogophloeus ; subgenus Boopinus ; Egypt).

— Herman, 1970: 393 ( Carpelimus ).

— Coiffait, 1970d: 144 ( Trogophloeus ; Uzbekistan).

— Jarrige, 1971: 499 ( Trogophloeus ; Iran).

— Smetana 1975c: 163 ( Trogophloeus ; Mongolia).

— Kashcheev, 1989: 284 ( Trogophloeus ; subgenus Boopinus ; Mongolia).

- DISTRIBUTION: Egypt, Iran, Kazakhstan, Uzbekistan, Mongolia.

tenerepunctus Gildenkov, 1994: 123 [= 1996: 84] ( Carpelimus ; subgenus Paraboopinus ; Type locality: Rossiia: Samarskaia Obl., 130 km iuzhnee g. Samara, r. S’ezzhaia u pos. Alekseevka).

— Gildenkov, 1997a: 615 ( Carpelimus ; subgenus Paraboopinus ; characters).

— Gildenkov, 1998a: 54 ( Carpelimus ; Ukraine).

- DISTRIBUTION: Russia.

tenuipunctus Bernhauer, 1912d: 31 ( Trogophloeus ; Type locality: Argentinien:

Prov Buenos Aires).

— Scheerpeltz, 1933: 1089 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Herman, 1970: 393 ( Carpelimus ).

— Frank, 1982: 13 ( Carpelimus ; list of parasites).

- DISTRIBUTION: Argentina.

teres Fall, 1926 , see: Thinobius .

terminalis Cameron, 1945a: 142 ( Trogophloeus ; Type locality: Java: Mt. Gede , altitude

8000 feet).

— Herman, 1970: 394 ( Carpelimus ).

- DISTRIBUTION: Indonesia.

testaceipennis Cameron, 1923: 395 ( Trogophloeus ; subgenus Taenosoma ; Type locality: Haiti).

— Scheerpeltz, 1933: 1089 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Blackwelder, 1943: 84 ( Carpelimus ; characters; Haiti).

— Herman, 1970: 394 ( Carpelimus ).

- DISTRIBUTION: West Indies.

texanus Casey, 1889 , see: fulvipes Erichson, 1840 .

— Herman, 1970: 394 ( Carpelimus ).

- DISTRIBUTION: Greece.

tibialis Fauvel, 1907 , see: Thinodromus .

topali Scheerpeltz, 1972: 81 ( Trogophloeus ; subgenus Trogophloeus ; Type locality: Argentinien: Rio Negro, El Bolson, Mt. Piltriquitron, 1000 m).

- DISTRIBUTION: Argentina.

torrentum Cameron, 1930: 196 ( Trogophloeus ; Type locality: Mussoorie district: Arni Gad). — Scheerpeltz, 1933: 1089 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Herman, 1970: 394 ( Carpelimus ).

— Abdullah and Qadri, 1970: 123 ( Trogophloeus ; characters; West Pakistan).

— Watanabe and Shibata, 1976: 329 ( Trogophloeus ; Malaysia).

- DISTRIBUTION: Pakistan, India, Indonesia.

traegardhi Scheerpeltz, 1974: 63 ( Trogophloeus ; subgenus Taenosoma ; Type locality: Natal: Sweet-Waters).

- DISTRIBUTION: South Africa.

transmarinus Fauvel, 1907: 14 ( Trogophloeus ; Type locality: Mombasa; baie de Kavirondo; Madagascar: Diego Suarez).

— Bernhauer and Schubert, 1911: 105 ( Trogophloeus ; catalog).

— Herman, 1970: 394 ( Carpelimus ).

- DISTRIBUTION: Kenya, Madagascar.

transversalis Wollaston, 1857 , see: Thinodromus .

transversicollis Scheerpeltz, 1947: 279 ( Trogophloeus ; subgenus Taenosoma ; Type locality: Österreich: Neusiedler See).

— Horion, 1963: 210 ( Trogophloeus ; subgenus Taenosoma ; Austria).

— Lohse, 1964: 80 ( Trogophloeus ; subgenus Taenosoma ; characters; central Europe).

— Herman, 1970: 394 ( Carpelimus ).

— Lucht, 1987: 92 ( Trogophloeus ; checklist; Austria).

- DISTRIBUTION: Austria.

travei Coiffait, 1982: 158 ( Trogophloeus ; subgenus Paraboopinus ; Type locality: Nepal: environs de Pokhara, 2200 m).

- DISTRIBUTION: Nepal.

tremolerasi Bernhauer, 1912d: 32 ( Trogophloeus ; Type locality: Uruguay: Montevideo).

— Scheerpeltz, 1933: 1089 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Herman, 1970: 394 ( Carpelimus ).

- DISTRIBUTION: Uruguay.

triangulum Sharp, 1887 , see: Thinodromus .

tripunctulatus Coiffait, 1981 g: 304 ( Trogophloeus ; subgenus Boopinus ?; Type locality:

Ile Galapagos: Ile Santa Cruz, Station Darwin).

- DISTRIBUTION: Galapagos Islands.

trivialis Cameron, 1930: 197 ( Trogophloeus ; Type locality: Central Provinces : Rahatgaon, Hoshangabad).

— Scheerpeltz, 1933: 1089 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Herman, 1970: 394 ( Carpelimus ).

- DISTRIBUTION: India.

troglodytes Erichson, 1840: 810 ( Trogophloeus ; Type locality: Sardinia).

— Fauvel, 1878: 93 [= 1878a: 13] ( Trogophloeus ; Algeria).

— Fauvel, 1886: 20 [= 1886a: 12] ( Trogophloeus ; Algeria).

— Fauvel, 1902b: 62 ( Trogophloeus ; characters; Barbary; Egypt; Canaries).

— Klima, 1904: 48, 61 ( Trogophloeus ; subgenus Taenosoma ; characters; Sardinia; North Africa; Atlantic Islands).

— Bernhauer and Schubert, 1911: 105 ( Trogophloeus ; catalog). — Scheerpeltz, 1961: 2 ( Trogophloeus ; subgenus Taenosoma ; Iran).

— Herman, 1970: 394 ( Carpelimus ).

— Kashcheev, 1989: 284 ( Trogophloeus ; subgenus Taenosoma ; Mongolia).

— Ciceroni and Zanetti, 1995: 11 ( Carpelimus ; Italy).

- DISTRIBUTION: Madeira, Canary Islands, Morocco, Algeria, Tunisia, Egypt, Italy, Sardinia, Iran, Mongolia.

ruficollis Wollaston, 1864: 601 ( Trogophloeus ; Type locality: Fuerteventura : La Antigua. Teneriffe: Puerto Orotava).

— Wollaston, 1865: 520 ( Trogophloeus ; Madeira Islands; Canary Islands; Morocco).

— Fauvel, 1878: 93 [= 1878a: 13] ( Trogophloeus ; Algeria; Egypt).

— Fauvel, 1886: 20 [= 1886a: 12] ( Trogophloeus ; Morocco; Algeria; Tunisia; Egypt).

— Fauvel, 1902b: 62 ( Trogophloeus ; variety of troglodytes ).

— Bernhauer and Schubert, 1911: 105 ( Trogophloeus ; variety of troglodytes ).

— Scheerpeltz, 1933: 1089 ( Trogophloeus ; aberration of troglodytes ).

— Maachi, 1991: 249 ( Trogophloeus ; variety of troglodytes ; Morocco).

tauricus Gildenkov, 1998a: 55 ( Carpelimus ; subspecies of troglodytes ; Type locality: Ukraina: Krym, Leninskii raion, s. Semenovka, bereg Aktashskogo ozera). tunapunae Blackwelder, 1943: 80 ( Carpelimus ; Type locality: Trinidad: foothills north of Tunapuna).

— Herman, 1970: 394 ( Carpelimus ).

- DISTRIBUTION: West Indies.

unicolor Sharp, 1900 , see: Teropalpus .

uniformis LeConte, 1877: 244 ( Trogophloeus ; Type locality: Pennyslvania ; Georgia).

— Casey, 1889: 381 ( Trogophloeus ; characters; notes).

— Bernhauer and Schubert, 1911: 105 ( Trogophloeus ; catalog).

— Herman, 1970: 394 ( Carpelimus ).

— Downie and Arnett, 1996: 445 ( Carpelimus ; characters; USA).

- DISTRIBUTION: USA.

unipustulatus Cameron, 1941 , see: Thinodromus .

usingeri Bernhauer, 1942a: 42 ( Trogophloeus ; subgenus Carpelimus , cited as Carpalimus ; Type locality: Guam: Inarajan).

— Herman, 1970: 394 ( Carpelimus ).

- DISTRIBUTION: Marianna Islands.

vagans Cameron, 1930: 198 ( Trogophloeus ; Type locality: Dehra Dun: Nun Nadi. Mussoorie: Ringal Gad. Almora: Dwarakhet , alt. 5000 to 6000 feet).

— Scheerpeltz, 1933: 1089 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Abdullah and Qadri, 1970: 123 ( Trogophloeus ; cited as vagens; characters; West Pakistan).

— Herman, 1970: 394 ( Carpelimus ).

— Coiffait, 1982: 43 ( Trogophloeus ; Nepal).

- DISTRIBUTION: Pakistan, India, Nepal.

vagus Sharp, 1889: 417 ( Trogophloeus ; Type locality: Yokohama, Nagasaki, Ogura Lake , Niigata).

— Bernhauer and Schubert, 1911: 105 ( Trogophloeus ; catalog).

— Cameron, 1949b: 461 ( Trogophloeus ; Formosa).

— Nakane, 1963a: 83 ( Carpelimus ; characters; habitus photograph; Japan).

— Herman, 1970: 394 ( Carpelimus ).

— Yuh, Paik, Kwon, and Lee, 1985: 253 ( Carpelimus , cited as Carperimus; Korea).

— J. Li, 1993: 20 ( Carpelimus ; China).

- DISTRIBUTION: China, Korea, Japan, Taiwan.

vancouverensis Hatch, 1957: 93 ( Carpelimus ; subgenus Troginus ; Type locality: Oregon: McMinnville. Also cited from British Columbia).

— Herman, 1970: 394 ( Carpelimus ).

varicornis Bernhauer, 1904: 24 ( Trogophloeus ; Type locality: Grenada, Vendome Est., Leeward side).

— Bernhauer and Schubert, 1911: 105 ( Trogophloeus ; catalog).

— Blackwelder, 1943: 73 ( Carpelimus ; characters; Cuba; Haiti; Grenada; Trinidad).

— Herman, 1970: 394 ( Carpelimus ).

- DISTRIBUTION: West Indies.

laetipennis Cameron, 1923: 395 ( Trogophloeus ; subgenus Taenosoma ; Type locality: Grenada).

— Scheerpeltz, 1933: 1085 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Blackwelder, 1943: 73 ( Carpelimus ; synonym of varicornis ).

— Herman, 1970: 392 ( Carpelimus ).

variegatus Cameron, 1944d: 312 ( Carpelimus ; subgenus Troginus ; Type locality: Mauritius: Port Louis).

— Herman, 1970: 394 ( Carpelimus ).

- DISTRIBUTION: Mascarene Islands.

velutinus Sharp, 1887: 700 ( Trogophloeus ; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet).

— Bernhauer and Schubert, 1911: 105 ( Trogophloeus ; catalog).

— Herman, 1970: 394 ( Carpelimus ).

- DISTRIBUTION: Panama.

venustulus Rosenhauer, 1856 , see: Ochthephilus .

vespertinus Notman, 1920a: 189 ( Trogophloeus ; Type locality: Windsor, Broome Co., N.Y.). — Scheerpeltz, 1933: 1089 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Herman, 1970: 394 ( Carpelimus ).

— Downie and Arnett, 1996: 446 ( Carpelimus ; characters; USA).

- DISTRIBUTION: USA.

vianai Bernhauer, 1939 h: 233 ( Trogophloeus ; subgenus Taenosoma ; Type locality: Argentinien: Buenos Aires: Tigre).

— Herman, 1970: 394 ( Carpelimus ).

- DISTRIBUTION: Argentina.

vicinus Sharp, 1876: 400 ( Trogophloeus ; Type locality: Amazons).

— Bernhauer and Schubert, 1911: 105 ( Trogophloeus ; catalog).

— Herman, 1970: 394 ( Carpelimus ).

- DISTRIBUTION: Brazil.

vitalei Bernhauer, 1935a: 40 ( Trogophloeus ; Type locality: Pogliarino , Sizilien).

— Herman, 1970: 394 ( Carpelimus ).

— Ciceroni and Zanetti, 1995: 11 ( Carpelimus ; Italy).

- DISTRIBUTION: Italy.

vittatus Sharp, 1887 , see: Thinodromus .

volans Notman, 1920a: 188 ( Trogophloeus ; Type locality: Windsor, Broome Co., N.Y.).

— Scheerpeltz, 1933: 1089 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Hatch, 1957: 93 ( Carpelimus ; subgenus Trogophloeus ; cited as volitans; characters;

British Columbia; Washington; Oregon).

— Herman, 1970: 394 ( Carpelimus ).

— Downie and Arnett, 1996: 446 ( Carpelimus ; characters; USA).

- DISTRIBUTION: Canada, USA.

vulneratus Bernhauer, 1926 , see: Thinodromus walkeri Bernhauer, 1933 , see: Thinodromus .

weberi Blackwelder, 1943: 68 ( Carpelimus ; Type locality: Trinidad, base of Galeota Point). — Herman, 1970: 394 ( Carpelimus ).

- DISTRIBUTION: West Indies.

weissi Notman, 1924a: 71 ( Trogophloeus ; Type locality: Middlesex Co., New Jersey).

— Scheerpeltz, 1933: 1089 ( Trogophloeus ; subgenus Trogophloeus ; catalog).

— Herman, 1970: 394 ( Carpelimus ).

— Downie and Arnett, 1996: 444 ( Carpelimus ; characters; USA).

- DISTRIBUTION: USA.

wendeleri Herman, 1970: 394 ( Carpelimus ; replacement name for oculatus Wendeler ; cited as wendleri [see Herman, 2001]).

— Herman, 2001: 12 ( Carpelimus ; spelling corrected).

- DISTRIBUTION: Brazil.

oculatus Wendeler, 1930: 183 ( Oxytrogus ; [preoccupied]; Type locality: Brasilien,

Porte Alegre).

— Scheerpeltz, 1933: 1092 ( Oxytrogus ; catalog).

— Herman, 1970: 394 ( Carpelimus ; synonym of wendeleri , cited as wendleri ).

wittmeri Coiffait, 1979 , see: Thinodromus .

yemenicus Coiffait, 1981: 20 ( Trogophloeus ; subgenus Paraboopinus ; Type locality: Yemen: Wadi Zabid).

— Gildenkov, 1997a: 615 ( Carpelimus ; subgenus Paraboopinus ; characters).

- DISTRIBUTION: Yemen.

zahiri Abdullah and Qadri, 1970: 123 , 130 ( Trogophloeus ; Type locality: W. Pakistan: Karachi ).

- DISTRIBUTION: Pakistan.

zealandicus Sharp, 1900: 233 ( Trogophloeus ; Type locality: New Zealand: Greymouth ).

— Fauvel, 1900c: 183 ( Trogophloeus ; subgenus Teropalpus ; cited as zeelandicus).

— Bernhauer and Schubert, 1911: 106 ( Trogophloeus ; subgenus Troginus ; catalog).

— Herman, 1970: 394 ( Carpelimus ).

— Pope, 1977: 25 ( Carpelimus ; Britain).

— Allen, 1979: 98 ( Carpelimus ; collecting notes; Britain).

— Lohse, 1987a: 141 ( Carpelimus ; characters).

— Lohse and Lucht, 1989: 133 ( Carpelimus ; characters).

— Kuschel, 1990: 26 ( Carpelimus ; endemic; winged; habitat; New Zealand).

— Haghebaert, Bruge, and Drugmand, 1990: 37 ( Carpelimus ; Belgium).

— Schillhammer, 1995: 230 ( Carpelimus ; Austria).

— Hodge and Jones, 1995: 33 ( Carpelimus ; characters; Britain).

— Gildenkov, 1997a: 616 ( Carpelimus ; subgenus Troginus ; characters).

— Emberson, 1998: 35 ( Carpelimus ; collecting notes; Chatham Islands).

— Assing, Frisch, Kahlen, et al., 1998: 128 ( Carpelimus ; Germany; Austria).

— Whitehead, 1999a: 141 ( Carpelimus ; habitat notes; Britain).

— Bruge, 1999: 45 ( Carpelimus ; not in Belgium).

- DISTRIBUTION: New Zealand, Chatham Islands, Britain, Austria, Germany. incongruus Steel, 1969: 70 ( Carpelimus ; Type locality: Kent: Cliffe).

— Pope, 1977: 25 ( Carpelimus ; synonym of zealandicus ).

— Lohse, 1987a: 142 ( Carpelimus ; synonym of zealandicus ).

— Lohse and Lucht, 1989: 133 ( Carpelimus ; synonym of zealandicus ).

zellichi Bernhauer, 1903b: 189 ( Trogophloeus ; Type locality: Ragusa, Dalmatien).

— Klima, 1904: 48, 61 ( Trogophloeus ; subgenus Taenosoma ; characters; Dalmatia).

— Bernhauer and Schubert, 1911: 105 ( Trogophloeus ; catalog).

— Herman, 1970: 394 ( Carpelimus ).

- DISTRIBUTION: Croatia.

zerchei Gildenkov, 1998a: 57 ( Carpelimus ; subgenus Troginus ; cited as “in lit.”; [Note: Nomen nudum, without description]; cited from various sites in Ukraine).

zlobini Gildenkov, 1996a: 627 ( Carpelimus ; subgenus Paraboopinus ; Type locality: Mongoliia: Iuzhno-Gobiisk, aimak Bordzon-Gobi, 80 km iugo-iugo-vostochnee Nomgona). Crassodemus

[1 species; Neotropical Region]

Crassodemus Herman, 1968: 1 (species included: forminosus). TYPE SPECIES: Crassodemus foraminosus Herman , fixed by original designation.

— Herman, 1970: 372 (characters; distribution; notes; list of species).

foraminosus Herman, 1968: 5 ( Crassodemus ; Type locality: Brazil: São Paulo).

— Herman, 1970: 372 ( Crassodemus ).

- DISTRIBUTION: Brazil.

Eppelsheimius

[2 species; Palaearctic Region]

Eppelsheimius Bernhauer, 1915 j: 270 (replacement name for Oncophorus Eppelsheim ). TYPE SPECIES: Oncophorus pirazzolii Eppelsheim , fixed by objective synonymy with Oncophorus .

— Winkler, 1925: 338 (catalog for Palaearctic region).

— Scheerpeltz, 1933: 1077 (world catalog supplement).

— Normand, 1935: 358 (list of species of Tunisia).

— Blackwelder, 1952: 150 (type species).

— Kocher, 1958: 96 (checklist of species; Morocco).

— Herman, 1970: 369 (characters; distribution; notes; list of species).

— Herman, 1983a: 1–12 (characters; revision of species; sister group).

Oncophorus Eppelsheim, 1885a: 46 (species included: pirazzolii ; preoccupied by Glocker, 1850, and Rudow, 1870). TYPE SPECIES: Oncophorus pirazzolii Eppelshiem , fixed by monotypy.

— Bernhauer and Schubert, 1911: 90 (world catalog; 2 species).

— Scheerpeltz, 1933: 1077 (synonym of Eppelsheimius ).

— Blackwelder, 1952: 275 (type species).

— Herman, 1970: 369 (synonym of Eppelsheimius ).

— Herman, 1983a: 6 (synonym of Eppelsheimius ).

Oncogenys Champion, 1919b: 154 (replacement name for Oncophorus Eppelsheim ). TYPE SPECIES: Oncophorus pirazzolii Eppelsheim , fixed by objective synonymy with Oncophorus .

— Scheerpeltz, 1933: 1077 (synonym of Eppelsheimius ).

— Blackwelder, 1952: 274 (type species).

— Herman, 1970: 369 (synonym of Eppelsheimius ).

— Herman, 1983a: 6 (synonym of Eppelsheimius ).

Eugenius Kastcheev, 1988a: 781 [= 1989a: 45] (species included: transaralicus , zarudnyi , ferrugineus , monstrosus , kryshanovskii). TYPE SPECIES: Eugenius transaralicus Kashcheev , fixed by original designation.

— Gusarov and Petrenko, 1989: 610 [= 1990: 83] (synonym of Eppelsheimius ).

ferrugineus Kashcheev, 1988 , see: pirazzolii Eppelsheim, 1885 .

kryzhanovskii Kashcheev, 1988 , see: pirazzolii Eppelsheim, 1885 .

magniceps Reitter, 1902 , see: pirazzolii Eppelsheim, 1885 .

miricollis Fauvel, 1898: 96 ( Oncophorus ; Type locality: Province de Constantine ; Tunisie).

— Fauvel, 1897d: 263 ( Oncophorus ; Algeria; Tunisia).

— Fauvel, 1902b: 73 ( Oncophorus ; Algeria; Tunisia).

— Bernhauer and Schubert, 1911: 90 ( Oncophorus ; catalog).

— C. Koch, 1934: 35 ( Eppelsheimius ; characters; Egypt).

— C. Koch, 1936b: 134 ( Eppelsheimius ; Egypt).

— Scheerpeltz, 1958: 17 ( Eppelsheimius ; characters; Algeria; Tunisia; Egypt; northern Sudan).

— Herman, 1970: 370 ( Eppelsheimius ).

— Jarrige, 1971: 499 ( Eppelsheimius ; Iran).

— Gusarov and Petrenko, 1989: 610 [= 1990: 83] ( Eppelsheimius ; notes).

- DISTRIBUTION: Algeria, Tunisia, Egypt, Sudan, Djibouti, Saudi Arabia, Iran, Turkmenistan.

monstrosus Kashcheev, 1988a: 782 , 787 [= 1989a: 51] ( Eugenius ; Type locality: Turkmeniia, Kara-Bogaz v 40 km sev Kyzyl-Arvata).

— Gusarov and Petrenko, 1989: 610 [= 1990: 83] ( Eppelsheimius ; synonym of miricollis ). monstrosus Kashcheev, 1988 , see: miricollis Fauvel, 1898 .

persicus Scheerpeltz, 1958 , see: pirazzolii Eppelsheim, 1885 .

pirazzolii Eppelsheim, 1885a: 47 ( Oncophorus ; Type locality: Tunis).

— Fauvel, 1886: 17 [= 1886a: 9] ( Oncophorus ; Tunisia).

— Fauvel, 1897d: 263 ( Oncophorus ; Algeria; Tunisia).

— Fauvel, 1902b: 73 ( Oncophorus ; Algeria; Tunisia).

— Bernhauer and Schubert, 1911: 90 ( Oncophorus ; catalog).

— C. Koch, 1936b: 134 ( Eppelsheimius ; Egypt).

— Normand, 1947a: 5 ( Eppelsheimius ; Tunisia).

— Scheerpeltz, 1958: 16 ( Eppelsheimius ; characters; Algeria; Tunisia; Libya; Egypt; northern Sudan).

— Herman, 1970: 370 ( Eppelsheimius ).

— Jarrige, 1971: 498 ( Eppelsheimius ; Iran).

— Coiffait, 1973a: 270 ( Eppelsheimius ; Morocco).

— Coiffait, 1979a: 163 ( Eppelsheimius ; Saudi Arabia).

— Herman, 1983a: 12 ( Eppelsheimius ; characters; Algeria; Egypt; Iran; Iraq; Morocco;

Saudi Arabia; Tunisia).

— Gusarov, 1991: 4 ( Eppelsheimius ; notes).

- DISTRIBUTION: Morocco, Algeria, Tunisia, Egypt, Sudan, Saudi Arabia, Iraq Iran, Turkmenistan, Uzbekistan, Kazakhstan.

magniceps Reitter, 1902: 186 ( Bledius ; subgenus Belidus ; Type locality: Transcaspien: Askhabad).

— Bernhauer and Schubert, 1911: 132 ( Bledius ; catalog).

— Winkler, 1925: 347 (subgenus Belidus ).

— Herman, 1970: 378 ( Bledius ).

— Herman, 1986: 223, 316 ( Bledius ; angustus group; distribution; catalog).

— Kashcheev, 1988: 37 ( Bledius ; subgenus Belidus ; characters).

— Gusarov, 1997: 288 ( Eppelsheimius ; synonym of pirazzolii ).

persicus Scheerpeltz, 1958: 14 ( Eppelsheimius ; Type locality: Persia: Sultanabad ).

— Herman, 1970: 370 ( Eppelsheimius ).

— Herman, 1983a: 12 ( Eppelsheimius ; synonym of pirazzolii ).

— Gusarov and Petrenko, 1989: 608 [= 1990: 81] ( Eppelsheimius ; synonym of pirazzolii ). transaralicus Kashcheev, 1988a: 782 [= 1989a: 47] ( Eugenius ; Type locality: pustynia Kyzylkum, Karakalpakiia, pos. Chabankazgan).

— Gusarov and Petrenko, 1989: 608 [= 1990: 81] ( Eppelsheimius ; synonym of pirazzolii ). zarudnyi Kashcheev, 1988a: 782 , 784 [= 1989a: 48] ( Eugenius ; Type locality: Iran, okrestnosti Kermana).

— Gusarov and Petrenko, 1989: 608 [= 1990: 81] ( Eppelsheimius ; synonym of pirazzolii ). ferrugineus Kashcheev, 1988a: 782 , 786 [= 1989a: 49] ( Eugenius ; Type locality: Iran, okrestnosti, Kermana).

— Gusarov, 1991: 4 ( Eppelsheimius ; synonym of pirazzolii ).

— Gusarov and Petrenko, 1989: 608 [= 1990: 81] ( Eppelsheimius ; synonym of pirazzolii ). kryzhanovskii Kashcheev, 1988a: 782 , 788 [= 1989a: 52] ( Eugenius ; Type locality: poima r. Syrdar’i, pos. Tselinnoe).

— Gusarov and Petrenko, 1989: 608 [= 1990: 81] ( Eppelsheimius ; synonym of pirazzolii ). Manda

[3 species; Nearctic and Palaearctic Regions]

Manda Blackwelder, 1952: 230 (replacement name for Acrognathus Erichson ). TYPE SPECIES: Omalium mandibulare Gyllenhal , fixed by original designation.

— Tottenham, 1954: 39 (characters; British species).

— Moore, 1964: 57, 58 (characters; key to species).

— Herman, 1970: 370 (characters; distribution; notes; list of species).

— Moore and Legner, 1975: 244 (catalog; North America north of Mexico).

— Pope, 1977: 24 (list of British species).

— Burakowski, Mroczkowski, and Stefańska, 1979: 83 (catalog; Poland).

— Moore and Legner, 1979: 223 (characters; notes).

— Muona, 1979: 19 (list of Scandinavian species).

— Frank, 1986a: 366 (checklist of species of Florida).

— Lohse and Lucht, 1989: 129 (valid name).

— Silfverberg, 1992: 23 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Navarrete-Heredia and Márquez-Luna, 1993 (cave collections; Mexico).

— Boháč, 1993: 42 (in list of species of Czech Republic and Slovakia).

— Ciceroni and Zanetti, 1995: 10 (list of species of Italy).

— M. Hansen, 1996: 98 (list of Danish species).

— Lundgren, 1998: 47 (list of species from Florida; records for other States listed for each species).

— Schülke, 1998i: 125 (characters).

— Newton, Thayer, Ashe, and Chandler, 2000: 380 (1 Nearctic species; notes; characters in key).

Acrognathus Erichson, 1839a: 607 (species included: mandibularis , palpalis ; preoccupied by Agassiz, 1836). TYPE SPECIES: Omalium mandibulare Gyllenhal , fixed by subsequent designation by Duponchel, 1841: 57.

— Erichson, 1840: 816 (characters).

— Redtenbacher, 1849: 742 (characters).

— Schaum, 1852: 29 (list of species; Europe).

— Lacordaire, 1854: 121 (characters; notes; list of species).

— Fairmaire and Laboulbène, 1856: 620 (characters).

— Gistel, 1856: 389 (list of species).

— Jacquelin du Val, 1857: 59 (characters).

— Redtenbacher, 1857: 238 (characters; Austrian species).

— Kraatz, 1857d: 898 (characters).

— Thomson, 1858: 39 (characters).

— Thomson, 1859: 45 (characters; type species: mandibularis ).

— Schaum, 1859: 31 (catalog; European species).

— Thomson, 1861: 141 (characters).

— Wencker and Silbermann, 1866: 35 (list of species; collecting notes; France).

— Jacquelin du Val, 1868: 78 (catalog; European species).

— Fauvel, 1871a: 152 [= 1872: 126] (characters).

— Sharp, 1871d: 14 (list of British species).

— Redtenbacher, 1874: 261 (characters).

— Fauvel, 1875a:VIII [= 1875b: 210] (catalog).

— Mulsant and Rey, 1878: 677 (characters).

— Heyden, 1880: 81 (list of species of Siberian region).

— Fowler, 1888: 394 (notes).

— Ganglbauer, 1895: 674 (characters).

— Heyden, Reitter, and Weise, 1906: 138 (list of species of Europe and Caucasus).

— Reitter, 1909: 177 (characters).

— Bernhauer and Schubert, 1911: 90 (world catalog; 1 species).

— Petri, 1912: 56 (list of species of Siebenbürgen).

— Johansen, 1914: 568 (characters).

— Roubal, 1930: 322 (catalog; Slovakia).

— Joy, 1932: 139 (key to British species).

— Scheerpeltz, 1933: 1077 (world catalog supplement).

— Normand, 1935: 358 (list of species of Tunisia).

— Kloet and Hincks, 1945: 162 (list of British species).

Tottenham, 1949: 359, 408 (type species: mandibularis Gyllenhal ; list of British species). — Palm, 1961a: 8 (characters; species of Sweden) .

— Horion, 1963: 162 (list of central European species).

— Székessy, 1963: 28 (characters).

— Lohse, 1964: 73 (characters; key to central European species).

— Scheerpeltz, 1968a: 20 (catalog of Austrian species).

— Herman, 1970: 370 (synonym of Manda ).

— Tikhomirova, 1973a: 143 (checklist of species of USSR).

— Pope, 1977: 24 (synonym of Manda ).

africana Fairmaire, 1860: 338 ( Compsochilus ; Type locality: Constantine).

— Fauvel, 1878: 91 [= 1878a: 11] ( Planeustomus ; Algeria).

— Fauvel, 1886: 17 [= 1886a: 9] ( Planeustomus ; Algeria).

— Fauvel, 1902b: 57 ( Planeustomus ; Algeria; Tunisia).

— Bernhauer and Schubert, 1911: 91 ( Planeustomus ; catalog).

— Scheerpeltz, 1933: 1077 ( Acrognathus ; catalog; Morocco).

— Peyerimhoff, 1936: 235 ( Acrognathus ; characters in key).

— Moore, 1964: 58 ( Manda ; characters).

— Herman, 1970: 371 ( Manda ).

- DISTRIBUTION: Morocco, Algeria, Tunisia.

cephalotes Erichson, 1840 , see: Planeustomus .

mandibularis Gyllenhal, 1827: 468 ( Omalium ; Type locality: Suecia meridionalis).

— Erichson, 1839a: 608 ( Acrognathus ; characters; Germany).

— Erichson, 1840: 817 ( Acrognathus ; characters; Germany; France; Sweden).

— Kiesenwetter, 1844a: 376 ( Acrognathus ; notes; Germany).

— Redtenbacher, 1849: 742 ( Acrognathus ; characters; Austria).

— Kiesenwetter, 1851a: 433 ( Acrognathus ; France).

— Fairmaire and Laboulbène, 1856: 620 ( Acrognathus ; characters; France).

— Redtenbacher, 1857: 238 ( Acrognathus ; characters; Austria).

— Kraatz, 1857d: 900 ( Acrognathus ; characters; Germany).

— Thomson, 1859: 45 (type species of Acrognathus ).

— Thomson, 1861: 142 ( Acrognathus ; characters; Scandinavia).

— Hochhuth, 1862: 100 ( Acrognathus ; notes; Russia).

— Fauvel, 1871a: 153 [= 1872: 127] ( Acrognathus ; characters; France; Scandinavia; Britain; Germany; Austria; Italy; Russia; Algeria).

— Redtenbacher, 1874: 262 ( Acrognathus ; characters; Austria).

— Fauvel, 1876a: 54 [= 1876: 239] ( Acrognathus ; Siberia).

— Seidlitz, 1875: 248 ( Acrognathus ; characters; Baltic region).

— Fauvel, 1878: 91 [= 1878a: 11] ( Acrognathus ; Algeria).

— Mulsant and Rey, 1878: 679 ( Acrognathus ; characters; France).

— Fauvel, 1886: 16 [= 1886a: 8] ( Acrognathus ; Algeria).

— Fowler, 1888: 395 ( Acrognathus ; characters; Britain).

— Seidlitz, 1891: 356 ( Acrognathus ; characters; Europe).

— Seidlitz, 1891a: 380 ( Acrognathus ; characters; Europe).

— Ganglbauer, 1895: 675 ( Acrognathus ; characters; north and middle Europe; Mediterranean region).

— Fauvel, 1897d: 249 ( Acrognathus ; Algeria).

— Fauvel, 1902b: 57 ( Acrognathus ; Algeria).

— Sainte-Claire Deville, 1907: 57 ( Acrognathus ; characters; France).

— Reitter, 1909: 177 ( Acrognathus ; characters; Germany).

— Portevin, 1929: 427 ( Acrognathus ; characters; France).

— Scheerpeltz, 1933: 1077 ( Acrognathus ; catalog).

— Jarrige, 1946: 98 ( Acrognathus ; natural history notes).

— Palm, 1961a: 10 ( Acrognathus ; characters; Sweden; Denmark).

— Horion, 1963: 162 ( Acrognathus ; Russia; Bulgaria; Romania; Hungary; Slovakia; Czech Republic; Poland; Sweden; Denmark; Britain; Holland; Belgium; France; Switzerland; Italy; Greece; Corfu; Austria; Germany).

— Lohse, 1964: 73 ( Acrognathus ; characters; central Europe).

— Moore, 1964: 58 ( Manda ; characters).

— Herman, 1970: 371 ( Manda ).

— Pope, 1977: 24 ( Manda ; Britain).

— Frank, 1982: 13 ( Manda ; list of parasites).

— Segers, 1986: 23 ( Manda ; checklist of species; Belgium).

— Lucht, 1987: 91 ( Acrognathus ; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux; Denmark; Sweden).

— Ádám, 1987: 138 ( Manda ; collecting notes; Hungary).

— Schülke and Uhlig, 1988: 3 ( Manda ; Germany).

— Boháč, 1993: 42 ( Manda ; Slovakia; Czech Republic).

— Terlutter, 1995: 6 ( Manda ; Germany).

— Ciceroni and Zanetti, 1995: 10 ( Manda ; Italy).

— Cibuļskis, 1998: 67 ( Manda ; Latvia).

- DISTRIBUTION: Europe, Russia.

nearctica Moore, 1964: 58 ( Manda ; Type locality: Florida: Archbold Biological Station, Lake Placid ).

— Herman, 1970: 371 ( Manda ).

- DISTRIBUTION: USA.

palpalis Erichson, 1839 , see: Planeustomus .

Mimopaederus

[1 species; Oceanic Region]

Mimopaederus Cameron, 1936: 4 (species included: insularis ). TYPE SPECIES: Mimopaederus insularis Cameron , fixed by monotypy.

— Blackwelder, 1952: 248 (type species: insularis ).

— Herman, 1970: 382 (characters; distribution; notes; list of species).

insularis Cameron, 1936: 5 ( Mimopaederus ; Type locality: Rapa, northwest ridge, Mangaoa Peak, altitude 900 to 1200 feet; northeast ridge, altitude 1000 to 1100 feet; northwest slopes, Tautautu Peak, altitude 700 to 800 feet).

— Herman, 1970: 383 ( Mimopaederus ).

- DISTRIBUTION: Austral Islands.

Neoxus

[2 species; Neotropical and Nearctic Regions]

Neoxus Herman, 1970: 394 (species included: crassicornis , dissimilis ). TYPE SPECIES: Thinobius crassicornis Casey , fixed by original designation.

— Moore and Legner, 1975: 246 (catalog; North America north of Mexico).

— Moore and Legner, 1979: 226 (characters; notes).

— Newton, Thayer, Ashe, and Chandler, 2000: 380 (1 Nearctic species; notes; characters in key). crassicornis Casey, 1889a: 87 ( Thinobius ; Type locality: Texas, Austin).

— Bernhauer and Schubert, 1911: 138 ( Thinobius ; subgenus Thinophilus ; catalog).

— Herman, 1970: 394 ( Neoxus ).

- DISTRIBUTION: USA.

dissimilis Sharp, 1887: 707 ( Thinobius ; Type locality: Panama, Bugaba ).

— Bernhauer and Schubert, 1911: 138 ( Thinobius ; subgenus Thinophilus ; catalog).

Ochthephilus

[48 species; Nearctic, Palaearctic, and Oriental Regions]

Ochthephilus Mulsant and Rey, 1856: 1 [= 1856c: 1] (species included: flexuosus ). TYPE SPECIES: Ochthephilus flexuosus Mulsant and Rey , fixed by monotypy.

— Schaum, 1859: 31 (synonym of Ancyrophorus ).

— Gemminger and Harold, 1868: 655 (synonym of Ancyrophorus ).

— Fauvel, 1871a: 165 [= 1872: 139] (synonym of Ancyrophorus ).

— Fauvel, 1875a: IX [= 1875b: 211] (synonym of Ancyrophorus ).

— Mulsant and Rey, 1878: 794 (characters; key to French species).

— Seidlitz, 1891: 352 (cited as Ochtephilus; key to some European species).

— Seidlitz, 1891a: 375 (cited as Ochtephilus; key to some European species).

— Heyden, Reitter, and Weise, 1891: 117 (list of species of Europe and Caucasus).

— Bernhauer and Schubert, 1911: 92 (synonym of Ancyrophorus ).

— Cameron, 1930: 176 (cited as Octhephilus; synonym of Ancyrophorus ).

— Tottenham, 1949: 360 (synonym of Ancyrophorus , thought by Tottenham to be a junior homonym of Ochthephilum ).

— Blackwelder, 1952: 266 (type species: flexuosus ).

— Palm, 1961a: 11 (characters; key to species of Sweden).

— Ferreira, 1962a: 10 (list of Portuguese species).

— Székessy, 1963: 28 (characters).

— Herman, 1970: 383 (characters; distribution; notes; list of species).

— Moore and Legner, 1975: 247 (catalog; North America north of Mexico).

— Shibata, 1976: 141 (checklist of species of Japan).

— Pope, 1977: 24 (list of British species).

— Burakowski, Mroczkowski, and Stefańska, 1979: 85 (catalog; Poland).

— Moore and Legner, 1979: 227 (characters; notes).

— Muona, 1979: 19 (list of Scandinavian species).

— Muona and Viramo, 1986: 16 (list of species of northeastern Finland).

— Segers, 1986: 24 (checklist of species; Belgium).

— Lohse and Lucht, 1989: 129 (valid name for genus; notes; supplement for key to genera of Lohse, 1964).

— Campbell and Davies, 1991: 94 (checklist of species; Alaska and Canada).

— Silfverberg, 1992: 23 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Boháč, 1993: 42 (list of species of Czech Republic and Slovakia).

— Terlutter, 1995: 7 (list of some species of Germany).

— Ciceroni and Zanetti, 1995: 10 (list of species of Italy).

— M. Hansen, 1996: 99 (list of Danish species).

— Downie and Arnett, 1996: 441 (characters; key to species of Northeastern North America).

— Schülke, 1998i: 126 (characters).

— Assing, Frisch, Kahlen, et al., 1998: 127 (characters).

— Newton, Thayer, Ashe, and Chandler, 2000: 380 (3 Nearctic species; notes; characters in key).

— Naomi, Kuranishi, Saito, and Maruyama, 2000: 107 (collected from North Kuril Islands).

Ancyrophorus Kraatz, 1857d: 886 (species included: rosenhaueri , flexuosus , longipennis , omalinus , venustulus , angustatus ). TYPE SPECIES: Trogophloeus omalinus Erichson , fixed by subsequent designation by Thomson, 1959: 44.

— Redtenbacher, 1857: 990 (characters).

— G. Waterhouse, 1858: 30 (catalog of British species).

— Thomson, 1859: 44 (characters; type species: omalinus ).

— Schaum, 1859: 31 (catalog; European species).

— LeConte, 1861a: 69 (characters).

— Thomson, 1861: 139 (characters).

— Gredler, 1863: 119 (list of species from Tirol).

— LeConte, 1863: 26 (list of North American species).

— Wencker and Silbermann, 1866: 35 (list of species; collecting notes; France).

— Jacquelin du Val, 1868: 78 (catalog; European species).

— Fauvel, 1869: 493 (synonym of Ochthephilus ).

— Fauvel, 1871a: 165 [= 1872: 139] (characters; key to species).

— Sharp, 1871d: 14 (list of British species).

— Redtenbacher, 1874: 259 (characters; key to Austrian species).

— Fauvel, 1875a: IX [= 1875b: 211] (catalog).

— LeConte, 1877: 241 ( Ancyrophorus ; characters; notes).

— Fauvel, 1878d: 264 [= 1878e: 100] (notes; key to North American species).

— Heyden, 1880: 81 (list of species of Siberian region).

— Duvivier, 1883: 106 (catalog).

— Fowler, 1888: 384 (notes; key to British species).

— Ganglbauer, 1895: 666 (adult and larval characters; key to middle European species).

— Heyden, 1896: 36 (list of species of Siberian region).

— Fauvel, 1897d: 250 (catalog of species of Barbary).

— Heyden, Reitter, and Weise, 1906: 138 (list of species of Europe and Caucasus).

— Sainte-Claire Deville, 1907: 59 (notes; key to and annotated list of species of Seine Basin, France).

— Reitter, 1909: 174 (characters; key to German species).

— Bernhauer and Schubert, 1911: 92 (world catalog; 13 species).

— Petri, 1912: 56 (list of species of Siebenbürgen).

— Leng, 1920: 95 (catalog of North American species).

— Winkler, 1925: 339 (catalog for Palaearctic region).

— Cameron, 1925: 105 (catalog of Indian species).

— Porta, 1926: 36 (characters; key to species of Italy).

— Portevin, 1929: 423 (characters; key to French species).

— Cameron, 1930: 176 (characters; key to species of British India).

— Roubal, 1930: 323 (catalog; Slovakia).

— Joy, 1932: 144 (key to British species).

— Scheerpeltz, 1933: 1079 (world catalog supplement).

— Normand, 1935: 358 (list of species of Tunisia).

— Tottenham, 1939d: 228 (type species designation: omalinus ).

— Kloet and Hincks, 1945: 162 (list of British species; type species: omalinus ).

— Jeannel and Jarrige, 1949: 325 (cave species; type species: rosenhaueri ).

— Tottenham, 1949: 360, 409 (type species: omalinus Erichson ; list of British species).

— Jarrige, 1949: 59 (key to French species).

— Scheerpeltz, 1950: 54 (key to Palaearctic species).

— Blackwelder, 1952: 52 (type species).

— Tottenham, 1954: 40 (characters; key to British species).

— Hatch, 1957: 85, 90 (characters; key to species of Pacific Northwest).

— Kocher, 1958: 96 (checklist of species; Morocco).

— Smetana, 1959a: 197 (list of species collected in Albania).

— Watanabe and Shibata, 1961: 6 (key to species of Japan).

— Horion, 1963: 167 (list of central European species).

— Lohse, 1964: 74 (characters; key to central European species).

— Smetana, 1967f: 384 (key to Czechoslovakian species).

— Kasule, 1968: 120 (larval characters; collecting notes).

— Scheerpeltz, 1968a: 21 (catalog of Austrian species).

— Herman, 1970: 383 (synonym of Ochthephilus ).

— Tikhomirova, 1973a: 143 (checklist of species of USSR).

— Scheerpeltz, 1976: 19 (key to species of Himalayan region).

— Pope, 1977: 24 (synonym of Ochthephilus ).

— M. Dvořák, 1979: 110 (some species collected in Slovakia).

— Dettner, 1987: 31 (defensive glands).

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Trogophloeus

Loc

Trogophloeus segnis

Herman, Lee H. 2001
2001
Loc

kerstensi

Scheerpeltz 1954: 7
1954
Loc

subtilicornis

Roubal 1946
1946
Loc

Carpelimus

Leach 1819
1819
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