Oedalea wulingshana, Zeng & Lin & Yang, 2022

Zeng, Wei, Lin, Chen & Yang, Ding, 2022, Four new species of Oedalea (Diptera, Empidoidea, Hybotidae, Oedaleinae) from China with a key to south eastern Asian species, Zootaxa 5141 (6), pp. 553-567 : 560-563

5. Oedalea wulingshana sp. nov.

( Figs 15–21 )

Diagnosis. Palpus dark brown. Antenna 1.2 times as long as head, dark brown except scape black; first flagellomere 7.0 times longer than wide. Eyes with enlarged dorsal ommatidia. Hairs on mesonotum somewhat dense, acr and dc not separated. Wing uniformly brownish. Pterostigma rather narrow, posterior margin far from R 2+3. Halter dark brown with brown base. Basal bridge of epandrium rather narrow; right surstylus rather short, nearly as long as wide, apically widely obtuse; left surstylus about 2.0 times wider than long.

Description. Male ( Fig. 15 ). Body length 3.6 mm, wing length 3.3 mm. Head ( Fig. 16 ) 0.7 mm, black with gray pollinosity. Eyes brownish black, contiguous on frons, except dorsal ommatidia enlarged, dark brownish red; clypeus shiny black. Hairs and bristles on head black, lower occiput with brownish yellow hairs. Ocellar tubercle distinct with 2 oc and 2 posterior hairs; upper occiput with 5–6 slightly long postocular bristles slightly curved forward and inward. Antenna rather long, 1.2 times as long as head height, dark brown except scape black; scape without dorsal bristles; pedicel with circlet of black apical bristles; first flagellomere elongated, 7.0 times longer than wide, short dark brown pubescent; stylus dark brown, nearly thick spine-like, about 0.8 times as long as basal width of first flagellomere. Proboscis 0.6 times as long as head height, brown to dark brown with black hairs; palpus dark brown with black hairs.

Thorax subshiny black with thin gray pollinosity. Hairs on thorax rather short, pale yellow; bristles dark brownish yellow; hairs on mesonotum uniform, somewhat dense; ppn absent; acr and dc not separated; 2 npl (anterior one short, posterior one long), 1 psa, 1 long prsc; 3 pairs of sc. Legs dark yellow to brownish yellow, except coxae nearly yellow, hind tibia blackish except base and tarsi from apex of tarsomere 1 blackish onward. Fore tibia slightly thickened, hind tibia slightly curved. Hairs and bristles on legs brown to dark brown; coxae with brownish yellow hairs and bristles. Fore femur slightly thickened, 1.2 times as wide as mid femur; hind femur distinctly thickened, 2.8 times as wide as mid femur, with row of rather long brownish pd on basal 3/5, with about four rows of black ventral spine-like bristles apically (outer av and pv rather long). Wing ( Fig. 3 ) uniformly brownish; dark brown pterostigma long, oval, posterior margin far from R 2+3; veins dark brown, R 2+3 weakly curved, M 2 complete. Squama dark yellow with dark yellow hairs. Halter dark brown with brown base.

Abdomen nearly straight, somewhat shiny blackish with gray pollen. Hypopygium narrower than pregenital segments. Hairs and bristles on abdomen brownish yellow, except those on apex dark brown.

Terminalia ( Figs 18–21 ): Left and right cerci ( Fig. 18 ) triangular and strongly narrowed. Basal bridge of epandrium ( Fig. 18 ) rather narrow; left and right epandrial lamellae ( Fig. 18 ) thick, not narrowed apically (dorsal view). Right epandrial lamella ( Fig. 20 ) somewhat trapezoidal (lateral view); right surstylus longer than wide, arched ventrally ( Fig. 20 ), basally with outer margin weakly incised, apically widely obtuse. Left epandrial lamella ( Fig. 19 ) somewhat trapezoidal (lateral view); left surstylus somewhat quadrate, short and wide (about 2.0 times wider than long), with straight apical margin (lateral view). Hypandrium ( Fig. 21 ) rather long; apical hook relatively long thick, weakly curved.

Female. Body length 3.7–4.0 mm, wing length 3.3–3.7 mm. Halter paler than male, brown with dark yellow base.

Type material. HOLOTYPE ♂, Hebei, Xinglong, Wulingshan, Mingxi [40°37′33″N, 117°27′42″E, 1019 m], 2018.VI.2, Qilemoge & Ding Yang ( CAU) . PARATYPES: 3 ♀, same data as holotype ( CAU) ; 1 ♀, Hebei, Xinglong, Longtan Scenic Spot [40°25′2″N, 117°30′3″E, 1153 m], 2018. VI .2, Jinlong Ren ( CAU) .

Distribution. China (Hebei).

Remarks. The new species is somewhat similar to O. freyi Chvála from Europe in having the eyes with enlarged dorsal ommatidia, but may be separated from the latter by the antennal stylus thick, spine-like and dark brown pterostigma. In O. freyi , the antennal stylus is somewhat slender, and the pterostigma is indistinct ( Chvála 1983).

Etymology. The specific name refers to the type locality, Wulingshan Mountain.

Figures

FIGURES 15–17. Oedalea wulingshana sp. nov. (male). 15. Adult habitus, lateral view; 16. Head, lateral view; 17. Wing.

FIGURES 18–21. Oedalea wulingshana sp. nov. (male). 18. Genitalia, dorsal view; 19. Left epandrial lamella, lateral view; 20. Right epandrial lamella, lateral view; 21. Hypandrium and phallus, ventral view. Abbreviations: see Material and methods.

FIGURES 1–3. Oedalea linzhiensis sp. nov. (male). 1. Adult habitus, lateral view; 2. Head, lateral view; 3. Wing.