Leiopsammodius Rakovic, 1984

Harpootlian, Phillip J., Gordon, Robert D. & Woodruff, Robert E., 2000, Review Of The Genus Leiopsammodius Rakovic (Coleoptera: Scarabaeidae: Aphodiinae: Psammodiini) In America North Of Mexico With Descriptions Of Three New Species, The Coleopterists Bulletin 54 (3), pp. 292-299 : 292-293

Leiopsammodius Rakovic

Psammodius (Leiopsammodius) Rakovic 1981:16 . Type: Psammodius laevicollis Klug 1845:42 , by original designation

Leiopsammodius Rakovic, Pittino 1984:319 , Gordon and Pittino 1992:262, 264

Species of Leiopsammodius, sensu Gordon and Pittino (1992) , can be characterized as follows: length 2.5–4.0 mm; oblong to oblong­oval, usually robust, distinctly dilated posteriorly; usually brown; clypeus emarginate, not dentate; eyes normal; pronotum with at most three transverse, usually vestigial, ridges and furrows with rows of punctures, median groove often present in basal portion; elytra smooth, not granulate; inner wings normal; posterior tibiae ro­

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bust; hind tarsal segments short, at least first segment strongly triangularly expanded, claws corneous.

Key to the Genus LEIOPSAMMODIUS RAKOVIC IN AMERICA NORTH OF MEXICO

1(0) Oval, dilated posteriorly; clypeus with granules small, conical; pronotum with distinct transverse sub­basal band of moderately coarse punctures, base unmargined ( Fig. 3 ); metasternum with median line obsolete; pygidium usually with terminal fringe of 10 setae; hind femora broad, almost twice as wide as middle femora ( Fig. 2 ); frequently attracted to light; widespread Florida endemic ------------------------- L. malkini (Cartwright)

1′ Oblong­oval, subparallel; clypeus with granules large, many elongate; pronotum without distinct transverse sub­basal band of moderately coarse punctures, base margined ( Fig. 6 ); metasternum with median line distinct; pygidium with terminal fringe 0–6 setae; hind femora narrower, not twice as wide as middle femora ( Fig. 11 ); readily attracted to light or not; Florida to North Carolina ----------------------------------------------------------------------- 2

2(1) Head strongly, evenly convex; pronotum with transverse grooves often faint, posterior row of punctures often obsolete ( Fig. 9 ); pygidium with terminal setae often obsolete; dark brown; parameres with ventral notch hooked apically ( Fig. 8 ); central Georgia; readily attracted to blacklight ---------------------------------------------------------------------------------------- L. ocmulgeei , new species

2′ Head convex, less apically; pronotum with transverse grooves and posterior row of moderately coarse punctures distinct ( Fig. 6 ); pygidium normally with 2–6 terminal setae; medium brown; parameres with ventral notch evenly rounded; central Florida or coastal North and South Carolina; not usually attracted to blacklight -------------------------------------------------- 3

3(2) Pronotum with median row of transverse punctures moderate ( Fig. 12 ); males with parameres stout, not distinctly elongate, apically angulate ( Fig. 10 ); coastal North and South Carolina ----------- L. acei , new species

3′ Pronotum with median row of transverse punctures coarse ( Fig. 6 ); males with parameres elongate, apically rounded ( Fig. 5 ); central Florida ------- ------------------------------------------------------------------------------------------- L. deyrupi , new species

Figures

Figs. 1–4. Leiopsammodius malkini. 1) male genitalia; 2) ventral surface; 3) head and pronotum, dorsal aspect; 4) hind tibial spurs.

Figs. 5–7. Leiopsammodius deyrupi. 5) male genitalia; 6) head and pronotum, dorsal aspect; 7) hind tibial spurs.

Figs. 10–12. Leiopsammodius acei. 10) male genitalia; 11) ventral aspect; 12) head and pronotum, dorsal aspect.

Figs. 8–9. Leiopsammodius ocmulgeei. 8) male genitalia; 9) dorsal aspect.