Antocha, Osten Sacken, 1860

Markevičiūtė, Radvilė, Podenas, Sigitas & Saldaitis, Aidas, 2021, New Antocha Osten Sacken (Diptera: Limoniidae) from Sichuan, China, Zootaxa 4969 (2), pp. 280-292 : 282

Key to species of the genus Antocha from Sichuan, China (males)

1 Crossvein m-cu of wing as long as or longer than distance between it and base of cell dm ( Fig. 1 )..................... 3

– Crossvein m-cu of wing shorter than distance between it and base of cell dm ( Fig. 2 )............................... 2

2 Caudal margin of tergite 9 nearly straight between two small bumps ( Fig. 3 )............................. A. pallidella

– Caudal margin of tergite 9 gently emarginate and weakly crenulate, without two small bumps ( Fig. 4 )........ A. nigribasis

3 Caudal margin of tergite 9 with pair of median lobes ( Fig. 5 ), bumps ( Figs 6–8 ), small tubercles ( Fig. 9 ) or deep U-shaped median notch ( Fig. 10 )................................................................................ 4

– Caudal margin of tergite 9 smooth ( Figs 11–14 , 16, 18–20 ).................................................... 9

4 Caudal margin of tergite 9 with rounded, median bumps ( Figs 6–8 ), small tubercles ( Fig. 9 ) or deep U-shaped median notch ( Fig. 10 )............................................................................................ 5

– Caudal margin of tergite 9 with two obliquely truncated median lobes ( Fig. 5 ).......................... A. emarginata

5 Apex of outer gonostylus narrowed to acute point ( Figs 6–8 ).................................................. 6

– Apex of outer gonostylus bifid ( Figs 9, 10 )................................................................ 8

6 Apex of gonocoxite broadly rounded ( Figs 7, 8 )............................................................ 7

– Apex of gonocoxite narrowly elongated ( Fig. 6 )................................................... A. fortidens

7 Outer gonostylus strongly bent ( Fig. 8 )............................................ A. nebulipennis nebulipennis

– Outer gonostylus arched, not bent ( Fig. 7 )........................................... A. nebulipennis immaculata

8 Interbase well developed, appearing loop-shaped; paramere approximately as long as gonocoxite, appearing as slender, simple rod ( Fig. 9 )................................................................................... A. bidens

– Interbase small, narrow to acute point; paramere shorter than gonocoxite ( Fig. 10 ).......................... A. bifida

9 Aedeagus connected with lateral structures ( Figs 11–14 )..................................................... 10

– Aedeagus covered by wide parameral sheath ( Figs 15 , 16 ) or without parameral sheath and any lateral structures ( Figs 18–20 ) .................................................................................................. 13

10 Lateral structures branched on tip of aedeagus ( Figs 12–14 ).................................................. 11

– Lateral structures very slender, twisted into complete spiral turn at distal end ( Fig. 11 )...................... A. spiralis

11 Lateral structure consists of two small branches ( Figs 13, 14 )................................................. 12

– Lateral structure consists of three small branches ( Fig. 12 ).......................................... A. constricta

12 Outer branch of lateral structure approximately three times in length smaller than inner branch ( Fig. 14 ).... A. quadrifurca

– Outer branch of lateral structure very short, narrowed to acute point ( Fig. 13 )......................... A. multidentata

13 Aedeagus covered by wide parameral sheath ( Figs 15 , 16 )................................................... 14

– Aedeagus not covered by parameral sheath ( Figs 18–20 )..................................................... 15

14 Parameral sheath forms four horns ( Fig. 15 )................................................. A. pulchra sp. nov.

– Parameral sheath simple, without any structures ( Fig. 16 )......................................... A. minuticornis

15 Apex of outer gonostylus obtuse or rounded ( Figs 18–20 ).................................................... 16

– Apex of outer gonostylus is gradually narrowed to acute tip ( Fig. 17 ).................................. A. lacteibasis

16 Apex of aedeagus bifid ( Fig. 18 )................................................................. A. setigera

– Apex of aedeagus curved, not bifid ( Figs 19, 20 )........................................................... 17

17 Paramere twisted, spiral-shaped ( Fig. 20 )............................................................. A. bella

– Paramere straight, tapered ( Fig. 19 )................................................................ A. indica

Figures

FIGURES 1–10. Distinctive features of genus Antocha in Sichuan, China. 1. wing of A. pulchra sp. nov., paratype; 2. wing of A. nigribasis, paratype from Markevičiūtė et al. (2019, fig. 11); 3. male tergite 9 of A. pallidella, from Alexander (1933, fig. 37); 4. male tergite 9 of A. nigribasis, from Alexander (1932, fig. 42); 5. male tergite 9 of A. emarginata, holotype from Markevičiūtė et al. (2019, fig. 9); 6. male gonocoxite, gonostyli and tergite 9 of A. fortidens; 7. male gonocoxite, gonostyli and tergite 9 of A. nebulipennis immaculata; 8. male gonocoxite, gonostyli and tergite 9 of A. nebulipennis nebulipennis from Markevičiūtė et al. (2019, figs 46, 49); 9. male tergite 9 from Alexander (1932, fig. 43) and genital structures of A. bidens, holotype; 10. male tergite 9 and genital structures of A. bifida.

FIGURES 11–15. Distinctive features of genus Antocha in Sichuan, China. 11. male genitalia and genital structures, dorsal view of A. spiralis; 12. male genital structures and tergite 9 of A. constricta; 13. male genital structures and tergite 9 of A. multidentata; 14. male genital structures and tergite 9 of A. quadrifurca; 15. male genital structures and tergite 9 of A. pulchra sp. nov., paratype.

FIGURES 16–20. Distinctive features of genus Antocha in Sichuan, China. 16. male genitalia, dorsal view of A. minuticornis, holotype from Markevičiūtė et al. (2019, fig. 21); 17. male gonocoxite and gonostyli of A. lacteibasis, paratype; 18. male genital structures and tergite 9 of A. setigera, paratype; 19. male genital structures and tergite 9 of A. indica; 20. male genital structures and tergite 9 of A. bella, paratype from Markevičiūtė et al. (2019, figs 12, 26, 28).