Lycoriella eurystylata, Menzel & Vilkamaa, 2021

Menzel, Frank & Vilkamaa, Pekka, 2021, New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region, Zootaxa 5072 (6), pp. 501-530 : 507-509

Lycoriella eurystylata sp. n.

Figs 4B , 6C , 13A

Material studied. Holotype male. CANADA, Quebec, Old Chelsey , 3.VIII.1958, J. R. Vockeroth (in CNC).

Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale brown. Eye bridge 3 facets wide. Face with 14 setae. Clypeus with 4 setae. Maxillary palpus with 3 segments, 1 st segment as long as 3 rd segment, 2 nd segment shortest; 1 st segment with 5 setae, with small dorsal pit with sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.2x as long as wide, neck slightly shorter than wide, longest setae longer than width of flagellomere. Thorax. Brown; setae pale. Anterior pronotum with 5 setae. Proepisternum with 8 setae. Scutellum with 2 long and some short and fine setae. Wing. Fumose. Length 1.7 mm. Width/length 0.45. Anal lobe small. Veins distinct, except for stM. c/w 0.75. R1/R not detectable in the specimen studied. stM shorter than fork of M. bM and r-m subequal in length, stCuA shortest. bM and r-m non-setose. Halter yellow. Legs. Brown; setae pale. Fore tibial organ forming large patch of many setae in demarcated depression. Fore tibial spur slightly shorter than apical width of tibia. Abdomen. Pale brown; setae pale and moderately long. Hypopygium ( Fig. 4B ). Brown, like abdomen. Intergonocoxal area rather long, with short setose lobe. Gonocoxa longer than gonostylus, slightly roundish laterally, with rather dense and fine setosity. Gonostylus ( Fig. 6C ) widest subbasally, narrowed towards apex, strongly impressed medially, densely setose apically, with strong and curved apical tooth; with 5 medial megasetae in apical half, megasetae slender, long and straight, oblique, the apicalmost (with sockets) shorter than the others, as long as apical tooth; with well-differentiated long whip-lash seta in basal third of gonostylus. Tegmen ( Fig. 13A ) much wider than long, smoothly curved and membraneous apically, straight laterally, sclerotized basolaterally, with small area bearing a few aedeagal teeth. Aedeagal apodeme moderately long.

BIN. Unknown.

Discussion. Lycoriella eurystylata sp. n. resembles L. taimyrensis sp. n. and L. tundrae sp. n. in having the tegmen smoothly curved apicolaterally, whereas L. acutostylia Mohrig & Menzel, 1990 [described in Menzel et al. (1990)], L. jakovlevi sp. n. and L. kinbasketi sp. n. have the tegmen distinctly angled apicolaterally ( Fig. 13 ). L. eurystylata differs from L. taimyrensis and L. tundrae in having the tegmen very short, strongly sclerotized basolaterally, semicircular apically and without tooth-like structures, whereas the other two have their tegmen only slightly sclerotized laterally, less curved apically and with tooth-like structures. Furthermore, L. eurystylata has five, L. tundrae four and L. taimyrensis eight to nine gonostylar megasetae and the fore tibial organ of L. eurystylata is more distinctly demarcated than that of L. taimyrensis and L. tundrae .

L. eurystylata is similar to L. acutostylia , L. jakovlevi and L. kinbasketi in having rather long antennal flagellomeres (4 th flagellomere 2.1–2.3 times as long as wide), whereas L. taimyrensis and L. tundrae have shorter flagellomeres (4 th flagellomere 1.4–1.6 times as long as wide). To distinguish L. taimyrensis and L. tundrae , see under the former, and to distinguish L. acutostylia , L. jakovlevi and L. kinbasketi , see under L. jakovlevi sp. n.

Etymology. The name is derived from the Greek words eurys, broad, and stylos, style, referring to the thick gonostylus of the male hypopygium.

Figures

FIGURE 4. Hypopygium, ventral. A. Lycoriella acutostylia Mohrig & Menzel, 1990 (from Finland, Ilmajoki). B. L. eurystylata sp. n. (holotype). Scale 0.1 mm.

FIGURE 6. Gonostylus, ventral. A. Lycoriella jakovlevi sp. n. (holotype). B. L. kinbasketi sp. n. (holotype) C. L. eurystylata sp. n. (holotype). Scale 0.1 mm.

FIGURE 13. Tegmen, ventral. A. Lycoriella eurystylata sp. n. (holotype). B. L. acutostylia Mohrig & Menzel, 1990 (from Finland, Sadinvaara). C. Lycoriella jakovlevi sp. n. (paratype). D. L. kinbasketi sp. n. (holotype). E. L. taimyrensis sp. n. (holotype). F. L. tundrae sp. n. (holotype). Scale 0.1 mm.