Mecynotarsus magelae, Kejval & Cz, 2013

Kejval, Zbyněk, 2013, Taxonomic revision of the Australian Notoxinae (Coleoptera: Anthicidae), Acta Entomologica Musei Nationalis Pragae (suppl.) 53, pp. 1-98 : 58-60

Mecynotarsus magelae sp. nov.

( Figs 74, 75 , 156–158 , 201 , 202 )

Type locality. Australia, Northern Territory, 9 km SSE of Mudginbarry Homestead, Magela Creek, 12°40′S 132°54′E.

Type material. HOLOTYPE: ♂, ‘ 12.40S 132.54E Magela Creek, N.T. 9 km SSE of Mudginbarry HS. 6.xi.72, at light E. Britton [p] // AUST. NAT. MUS. INS. COLL. [p; green label]’ ( ANIC). PARATYPES: 4 ♂♂ 4 ♀♀, same data as holotype ( ANIC, 1 spec. ZKDC) ; 1 ♂ 1 ♀, ‘ Magela Ck. , NT. 9km. SSE of Mudginbarry HS. 6 Nov. 1972 D. H. Colless [p] // AUST. NAT. MUS. INS. COLL. [p; green label]’ ( ANIC, ZKDC) ; 3 ♀♀, ‘ 12.40S, 132.54E Magela Creek , 9 km SSE. of Mudginbarry HS., N.T. 7.xi.72, M. S. Upton [p] // AUST. NAT. MUS. INS. COLL. [p; green label]’ ( ANIC) ; 1 ♂, ‘ 12.06S 133.04E Cooper Creek, N.T., 19 km E. by S. of Mt. Borradaile , 2.xi.72, at light, E. Britton [p] // AUST. NAT. MUS. INS. COLL. [p; green label]’ ( ANIC) ; 1 ♀, same data, except: 9.xi.72, M. S. Upton ( ANIC) ; 1 ♀, ‘ Cooper Ck., N.T. 19 km.E by S of Mt. Borradaile , 2 Nov. 1972 D. H. Colles [p] // AUST.NAT. MUS. INS. COLL. [p; green label]’ ( ANIC) ; 1 ♀, ‘ 12.17S 133.20E Cooper Creek, NT. 11 km S. by W. of Nimbuwah Rock 1.xi.72, at light E. B. Britton [p] //AUST.NAT. MUS. INS.COLL.[p; green label]’ ( ANIC) ; 5♀♀, ‘ 12.52S, 132.50E Koongarra , 15 km E. of Mt. Cahill, N.T. 15.xi.1972, M. S. Upton [p] // AUST. NAT. MUS. INS. COLL. [p; green label]’ ( ANIC, 2 spec. ZKDC) ; 2 ♂♂ 1 ♀, ‘ 12.47S, 132.51E 19 km NE. By E. of Mt. Cahill, N.T. 16.xi.1972, M. S. Upton [p] // AUST. NAT. MUS. INS. COLL. [p; green label]’ ( ANIC) ; 1 ♂ 3 ♀♀, ‘ Bessie Spring 16.40S 135.51E 8 km ESE of Cape Crawford, NT. 26 Oct. 1975 M. S. Upton [p] // AUST. NAT. MUS. INS. COLL. [p; green label]’ ( ANIC) ; 1 ♂, ‘ 13.45S 131.34E GPS Butterfly Gorge Nature Pk Douglas River NT 19 Jul. 1994, at light T. Weir, A. Roach [p] // AUST. NAT. MUS. INS. COLL. [p; green label]’ ( ANIC) ; 1 ♂, ‘ 8 km ENE of Victoria Riv. Downs, N.T. 12 July, 1973 L. P. Kelsey [p] // At light [p] // ANIC Specimen [p; green label]’ ( ANIC) ; 1 ♂, ‘ 13.15S, 131.06E Adelaide River, N. T. 17.x.72, M.S. Upton [p] // AUST. NAT. INS. COLL. [p; green label]’ ( ANIC) ; 1 ♂, ‘ McArthur River 16.27S 136.05E 48 km SW by S of Borroloola, N.T. 13.iv.1976 at light J.E. Feehan [p] // ANIC Specimen [p; green label]’ ( ANIC) ; 3 ♂♂, ‘ AUSTRALIA, N. Territory, Banka Banka env., road to Tennanth Creek 14°53′S 132°01′E, 316 m, 12.- 13.01.2009, St. Jakl leg. [p]’ ( ZKDC, NMPC) ; 2 ♂♂ 4 ♀♀, ‘ Kimberley Res. Stn. Light Trap W.A. October 1961. K. T. Richards // ANIC Specimen [p; green label]’ ( ANIC) ; 1 ♂ 1 ♀, ‘ Kimberley Research Station , Kununurra, nr. Wyndham, W.A. (15.28S 128.06E) 27.xi.1956 [p] // AUST. NAT. MUS. INS. COLL. [p; green label]’ ( ANIC) ; 1 ♂, ‘ 16.22S 125.12E W.A. Charnley Riv. 2 km SW Rolly Hill CALM Site 25/2 16-20 June 1988 I. D. Naumann [p] // at light, open forest near closed forest margin // Mecynotarsus sp. 4 det. T.A. Weir 1989 [p+h] // AUST. NAT. MUS. INS. COLL. [p; green label]’ ( ANIC) ; 4 ♀♀, same data, but lacking 3rd label ( ANIC, 1 spec. ZKDC) ; 3 ♂♂ 1 ♀, ‘ 14.49S 126.49E Carson escarpment W.A. 9–15 Aug. 1975 I.F.B. Common and M.S. Upton [p] // ANIC Specimen [p; green label]’ ( ANIC) ; 1 ♀, ‘ 15.38S 125.15E W.A. CALM Site 28/3 4 km W of King Cascade 12-16 June 1988 T.A.Weir [p] at light, open forest [p] // Mecynotarsus sp. 5 det. T.A.Weir 1989 [p+h] // AUST. NAT. INS. COLL. [p; green label]’ ( ANIC) ; 2 ♀♀, ‘ 12.27S 142.38E QLD Moreton , 10Dec. 1992 W. Dressler, P. Zborowski, at light [p] // AUST. NAT. INS. COLL. [p; green label]’ ( ANIC) ; 1 ♂, ‘ Qld. Greenvale 70 Km SW at light 1-13 Jan. 96 A.J.Watts [p] // SAMA Database No. 25-029048 [p]’ ( SAMA) ; 2 ♀♀, same data, except: ‘ 14-24 Mar 1995 ’ and database number 25-028959 ( SAMA) ; 1 ♀, ‘ Karumba , Qld. 27–28.v.1972 G.B. & S.R. Monteith [p]’ ( QMBA) .

Description (holotype, male). Body length 1.8 mm. Head largely brown black, somewhat paler ventrally, pronotum and elytra reddish brown, elytra with vague darker marking; legs and antennae reddish.

Antero-lateral margins of frons simple (not raised near antennal insertion). Gular rugules anteriorly coarser, contiguous to fused as in Figs 138, 139 . Clypeal granules minute. Setation of head short and appressed, rather fine medially on vertex (erect setae absent), distinctly coarser to scaly around eyes and ventro-laterally. Antennae moderately long; antennomeres III–V about twice, X as long as wide; setation mostly fine, coarser on basal four antennomeres, especially antennomere I with some whitish scales.

Pronotum moderately transverse, 1.4 times as long as wide, its lateral margins rather strongly, somewhat unevenly convex in dorsal view; posterior collar narrow but distinct. Pronotal horn rather robust and wide, triangular, its posterior angles projecting in dorsal view ( Fig. 156 ); horn margins armed with 4 lobules on each side, apical lobule simple, widely rounded; horn crest very conspicuous, strongly raised, short and situated rather posteriorly, forming nearly complete semicircular rim with single small, separate rugule posteriorly on each side; submarginal rugules numerous, distinctly spaced; single large, longitudinal median rugule and several minute granules posteriorly. Setation whitish, scaly, much finer and rather inconspicuous on pronotal horn dorsally; scales on pronotal disc quite appressed, somewhat variable in length (not clearly of two sizes), longitudinally oval, rounded to subtruncate apically, very densely spaced and somewhat glossy; antebasal paired setae present both medially and laterally (median pair shorter and coarser); additional tactile setae absent.

Elytra 1.6 times as long as wide; omoplates and postbasal impression absent. Setation greyish and brownish, forming vague brownish markings ( Fig. 201 ), scaly, appressed and evenly ordered; scales similar to those on pronotal disc, but smaller and rather opaque, very densely spaced (surface not visible); erect tactile setae absent.

Male characters. Sternum VII slightly produced medially. Tergum VIII and aedeagus as in Figs 74, 75 ; parameres evenly curved in apical half.

Variation. Body length (♂ ♀) 1.7–2.4 mm. Pronotal horn with 3–4 (mostly 4) lobules on each side, varying in shape; apical lobule simply and widely rounded to nearly bilobed; rugules of crest margins mostly fused and forming evenly shaped rim ( Figs 156, 157 ), rarely separate ( Fig. 158 ). Dark markings of elytra reduced ( Fig. 201 ) to extensive ( Fig. 202 ) and more or less sharply outlined.

Differential diagnosis. Mecynotarsus magelae sp. nov. may resemble M. pilbarensis sp. nov., especially for specimens with more extensive dark markings of the elytra (cf. Fig. 202 versus 206) or lacking the evenly shaped crest rim of the pronotal horn ( Fig. 158 ). It differs from this species by the shorter and more compact, posteriorly situated crest of the pronotal horn (rugules mostly fused and forming a semicircular rim as in Figs 156, 157 ), the single and rather simply shaped median rugule, the dense fringe of whitish scales on the posterior margin of the pronotum (sparser, pale and rather linear scales in the other species), and by the male characters (cf. Figs 74, 75 versus 102, 103).

Etymology. The species name is a noun in genitive case; named after the type locality.

Distribution. Australia: Northern Territory, Queensland, Western Australia.

Figures

Figs 74–81. 74–75 – Mecynotarsus magelae sp. nov.: 74 – tergum VIII, 75 – aedeagus in dorsal view. 76–79 – M. mastersii MacLeay, 1872: 76 – sternum VII, 77 – tergum VIII, 78 – tegmen in dorsal view, 79 – median lobe in dorsal view. 80–81 – M. mollis sp. nov.: 80 – tergum VIII, 81 – aedeagus in dorsal view. Scale (0.2 mm): A – Fig. 75; (0.5 mm): B – Fig. 76; C – Fig. 77; D – Figs 80, 81; E – Figs 74, 78, 79.

Figs 150–158. Pronotal horn in dorsal view: 150 – Mecynotarsus festivus sp. nov., 151 – M. grandior sp. nov., 152 – M. granulatus sp. nov., 153 – M. hortensis Lea, 1922 (syntype, SAMA), 154 – M. kingii MacLeay, 1872, 155 – M. leai Pic, 1942 (Richmond), 156 – M. magelae sp. nov., 157 – the same (paratype, Carson Escarpment), 158 – the same (paratype, Charnley River).

Figs 194–201. Habitus: 194 – Mecynotarsus festivus sp. nov., 195 – M. grandior sp. nov., 196 – M. granulatus sp. nov., 197 – M. hortensis Lea, 1922 (Fowlers Gap), 198 – M. kingii MacLeay, 1872, 199 – M. kreusleri King, 1869 (Mount Tozer), 200 – M. leai Pic, 1942 (Richmond), 201 – M. magelae sp. nov.

Figs 202–209. Habitus: 202 – Mecynotarsus magelae sp. nov. (Charnley River), 203 – M. mastersii MacLeay, 1872 (Cedar Creek National Park), 204 – M. obesus sp. nov., 205 – M. parvulus sp. nov., 206 – M. pilbarensis sp. nov. (paratype, Millstream Homestead), 207 – M. regalis sp. nov., 208 – M. weiri sp. nov., 209 – M. ziczac King, 1869 (Newcastle Grain Terminal).

Figs 133–140. 133 – Mecynotarsus setulosus sp. nov., head in lateral view (paratype, Fowlers Gap), 134 – M. excavatus sp. nov., apex of elytra in female (paratype, Aoue-Enga), 135 – M. grandior sp. nov., apex of elytron in male (paratype, Kimberley Research Station), 136 – M. obesus sp. nov., metatibia with tarsus (paratype, Hull River), 137 – M. excavatus sp. nov., male protibia (paratype, Aoue-Enga). 138–140 – head in ventral view: 138 – M. auripilosus sp. nov. (paratype, Kambah Pool), 139 – M. bullatus sp. nov. (paratype, Regans Ford), 140 – M. ziczac King, 1869 (Swain Reefs).