Cybilla Pogue, 2019

Brown, John W., 2019, New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae), Zootaxa 4671 (2), pp. 195-222 : 202

Cybilla Pogue , new genus

Type species: Phalonia hubbardana Busck, 1907 , by monotypy.

Cybilla Pogue and Mickevich, 1990: 322 (nomen nudum).

Cochylini New Genus 3 [ hubbardana group]: Brown 2005: 208; Metzler and Brown 2014: 278.

Cybilla is proposed for the single species Phalonia hubbardana , which traditionally has defied generic assignment. Described in Phalonia by Busck (1907), it was treated as such by McDunnough (1939) and was placed in “incertae sedis” by Powell (1983). Pogue (1986) proposed the name Cybilla, and Brown (2005) and Metzler and Brown (2014) treated it as “ Cochylini New Genus”. Pogue (1986) concluded that Cybilla is most closely related to Decuma (now recognized as Monoceratuncus ), and that the two are the most derived Cochylina genera in North America.

Diagnosis. Cybilla shares with Monoceratuncus an unusually modified uncus, the absence of socii, and the absence of a median process of the transtilla. Cybilla can be distinguished from Monoceratuncus by the presence of small lateral plates fused to the base of the uncus, and by the 4-segmented maxillary palpi.

Description. Head: Vertex and upper frons rough scaled, lower frons with sparse appressed scales; ocellus present; sensory setae of antenna ca. 1.0 times flagellomere diameter in male, shorter, sparser in female; labial palpus upturned, porrect, combined length of all segments ca. 1.2 times diameter of compound eye, segment III exposed; maxillary palpus 4-segmented. Thorax: Posterior crest absent; lateral scale tufts of metanotum flat. Forewing ( Figs. 5, 6 ) length 4.6–5.6 mm, length 3.0–3.1 times width, females usually slightly larger than males; costa straight, without costal fold in male, apex rounded; termen gently curved; Sc slightly less than 0.5 wing length; R 1 originating beyond middle of discal cell; R 2 originating nearer R 3 than R 1; R 5 ending at costa; M 3 and CuA 1 separate; CuA 2 originating ca. 0.75 length of discal cell; CuP absent; A1+2 stalked at ca. 0.5 length. Hindwing moderately broad, costa straight, with costal roll extending about 0.4 length of costa in male; apex rounded; termen straight; Sc+R 1 ca. 0.65 wing length; Rs and M 1 stalked at 0.65–0.75 length of M 1; M 3 and CuA 1 separate; CuA 2 originating ca. 0.65 length of discal cell; frenulum with one acanthus in male, two in female. Abdomen: Unmodified. Male genitalia ( Fig. 22 ) with well-developed uncus bearing small, closely appressed, flat, lateral plates; socius absent; transtilla well developed, concave mesially, lacking median process; valva broad at base, attenuate distally, length ca. 2.1 times width; costa setose, apex produced, sacculus undulate from base to apex of valva, with patch of small spines at outer margin of first undulation from base; vinculum arms free distally. Phallus large, ca. 1.1 times length of valva, straight, parallel-sided in basal 0.33, enlarged in distal 0.5, with ventral spadelike projection; vesica with numerous cornuti. Female genitalia ( Fig. 36 ) with papillae anales curved near middle, bulbous, slightly bilobed, setose; apophyses short, slender, length of apophyses anteriores ca. same as apophyses posteriores, ventral branch of apophyses anteriores joined to sterigma; sterigma a lateral band with prominent short-digitate, caudally-directed, subtriangular median process; ductus bursae slender, membranous throughout; corpus bursae small, rounded, with minute spicules; accessory bursa originating dorsomedially from corpus bursae.

Etymology. The generic name is from the Latin goddess of nature, Cybele, and is feminine in gender.

Figures

FIGURES 1–10. Adults of Cochylini genera. 1. Anacochylidia maderana, Madera Canyon, Santa Rita Mountains, Arizona (EME). 2. Cagiva cephalanthana, Dorman Lake, Oktibbeha Co., Mississippi (USNM). 3. Atroposia oenotherana, Houston, Harris Co., Texas (USNM). 4. Atroposia oenotherana, Radisson, Onondaga Co., New York (USNM). 5. Cybilla hubbardana, Shafter, Presidio Co., Texas (USNM). 6. Cybilla hubbardana, Lordsburg, Hidalgo Co., New Mexico (CNC). 7. Honca grandis, Chimney Gulch, Golden, Colorado (USNM). 8. Pogospinia floridana, Long Key State Park, Monroe Co., Florida (USNM). 9. Nycthia pimama, Portal, Cochise Co., Arizona (USNM). 10. Nycthia yuccatana, Falcon State Park, Starr Co., Texas (USNM).

FIGURES 19–26. Male genitalia of Cochylini genera. 19. Anacochylidia maderana, Palmerlee, Arizona, USNM slide 23,359. 20. Atroposia oenotherana, Padre Island National Seashore, Texas, USNM slide 153,596. 21. Cagiva cephalanthana, Dorman Lake, Mississippi, USNM slide 150,530. 22. Cybilla hubbardana, Tucson, Arizona, USNM slide 29,397. 23. Honca grandis, Ogden Canyon, Utah, USNM slide 153,598. 24. Nycthia pimama, Blair Valley, California, USNM slide 144,042. 25. Nycthia yuccatana, Laguna Atacosa, Texas, USNM slide 120,405. 26. Pogospinia floridana, Key Largo, Florida, USNM slide 23,249.

FIGURES 33–38. Female genitalia of Cochylini genera. 33. Anacochylidia maderana, Palmerlee,Arizona, USNM slide 23,358. 34. Atroposia oenotherana, Padre Island National Seashore, Texas, USNM slide 153,597. 35. Cagiva cephalanthana, Dorman, Mississippi, USNM slide 153,609. 36. Cybilla hubbardana (holotype), Tucson, Arizona, USNM slide 23833. 37. Honca grandis, Wellsville, Utah, USNM slide 153,599. 38. Nycthia pimama, Miller Canyon, Arizona, slide MGP 660.