Anacochylidia Brown, 2019

Brown, John W., 2019, New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae), Zootaxa 4671 (2), pp. 195-222 : 197

Anacochylidia Brown , new genus

Type species: Anacochylidia maderana Brown , new species, by monotypy.

Diagnosis. The short broad valvae and the long, slender secondary process of the phallus of Anacochylidia ( Fig. 19 ) are most similar to those of Cochylidia Obraztsov, 1956 , but other features of the male genitalia deviate from those of all known Cochylidia . In Anacochylidia the top of the tegumen is broad with a shallow, rounded concavity; the costa of the valva is short, unmodified, and upcurved, without a basal process; and the phallus has an elongate secondary process ca. 0.5 times as long as the phallus itself. In contrast, the male genitalia of Cochylidia are characterized by a dorsally attenuate tegumen, often with a narrowly bifurcate process at the top; the base of the costa of the valva has a long, erect, somewhat digitate, setose process; and the phallus has a short, curved digitate process apically. The latter is probably homologous with the extraordinarily long process of the phallus of Anacochylidia.

Description. Head: Vertex and upper frons rough scaled, lower frons with sparse appressed scales; ocellus present; sensory setae of antenna about 0.9 times flagellomere diameter in male, shorter, sparser in female; labial palpus porrect, combined length of all segments ca. 1.2 times diameter of compound eye, segment III exposed; maxillary palpus inconspicuous. Thorax: Posterior crest absent; lateral scale tufts of metanotum flat. Forewing ( Fig. 1 ) length 4.5–4.8 mm, length ca. 2.8 times width; costa slightly and evenly arched throughout, without costal fold in male, apex rounded; termen obliquely angled toward hind margin; Sc ca. 0.45 times wing length; R 1 originating from about middle of discal cell; R 2 originating nearer R 3 than R 1; R 5 ending at costa well before apex; M 3 and CuA 1 separate; CuA 2 originating about 0.75 times length of discal cell; CuP absent; A1+2 stalked in distal 0.65. Hindwing length ca. 2.5 times width; costa weakly undulate, male with narrow costal roll in basal 0.5 of wing; apex rounded; termen slightly concave near middle; Sc+R 1 ca. 0.25 times wing length, Rs and M 1 stalked ca. 0.3 length of M 1; M 3 and CuA 1 separate; CuA 2 originating ca. 0.6 length of discal cell; frenulum with one acanthus in male, two acanthi in female. Abdomen: Unmodified. Male genitalia ( Fig. 19 ) with tegumen short, broad, with shallow, rounded excavation at top bordered laterally by short lobelike processes; subscaphium subrectangular, semi-sclerotized; socius inconspicuous; transtilla well developed, broad, weakly bilobed with small rounded median lobe; valva short, broad, subquadrate, pointed apically, outer margin undulate, length ca. 0.4 times width; costa sclerotized, curved dorsally; sacculus represented by narrow, curved, sclerotized edge, lacking free apical process; vinculum arms weakly joined by membrane distally. Phallus elongate, ca. 4.0 times length of valva, strongly curved in basal 0.4, with slender curved spine originating dorsally just beyond curve, extending ca. 0.5 length of phallus, vesica with dense bundle of 7–8 small, non-deciduous cornuti. Female genitalia ( Fig. 33 ) with ovipositor long, slender, telescopic; papillae anales slender, slightly pointed posteriorly; apophyses posteriores ca. 2.0 times as long as apophyses anteriores; sterigma mostly membranous, lamella postvaginalis a subquadrate plate, attenuate and distinctly emarginate posteriorly, with a pair of narrow, submedial sclerites extending anteriorly from plate; colliculum sclerotized with longitudinal wrinkles, occupying entire ductus bursae; corpus bursae rounded, differentiated from ductus bursae by absence of sclerotization, signum lacking; origin of accessory bursa uncertain.

Etymology. The name comes from the similar genus Cochylidia and the Greek “ana” (= against) and is feminine in gender.

Figures

FIGURES 19–26. Male genitalia of Cochylini genera. 19. Anacochylidia maderana, Palmerlee, Arizona, USNM slide 23,359. 20. Atroposia oenotherana, Padre Island National Seashore, Texas, USNM slide 153,596. 21. Cagiva cephalanthana, Dorman Lake, Mississippi, USNM slide 150,530. 22. Cybilla hubbardana, Tucson, Arizona, USNM slide 29,397. 23. Honca grandis, Ogden Canyon, Utah, USNM slide 153,598. 24. Nycthia pimama, Blair Valley, California, USNM slide 144,042. 25. Nycthia yuccatana, Laguna Atacosa, Texas, USNM slide 120,405. 26. Pogospinia floridana, Key Largo, Florida, USNM slide 23,249.

FIGURES 1–10. Adults of Cochylini genera. 1. Anacochylidia maderana, Madera Canyon, Santa Rita Mountains, Arizona (EME). 2. Cagiva cephalanthana, Dorman Lake, Oktibbeha Co., Mississippi (USNM). 3. Atroposia oenotherana, Houston, Harris Co., Texas (USNM). 4. Atroposia oenotherana, Radisson, Onondaga Co., New York (USNM). 5. Cybilla hubbardana, Shafter, Presidio Co., Texas (USNM). 6. Cybilla hubbardana, Lordsburg, Hidalgo Co., New Mexico (CNC). 7. Honca grandis, Chimney Gulch, Golden, Colorado (USNM). 8. Pogospinia floridana, Long Key State Park, Monroe Co., Florida (USNM). 9. Nycthia pimama, Portal, Cochise Co., Arizona (USNM). 10. Nycthia yuccatana, Falcon State Park, Starr Co., Texas (USNM).

FIGURES 33–38. Female genitalia of Cochylini genera. 33. Anacochylidia maderana, Palmerlee,Arizona, USNM slide 23,358. 34. Atroposia oenotherana, Padre Island National Seashore, Texas, USNM slide 153,597. 35. Cagiva cephalanthana, Dorman, Mississippi, USNM slide 153,609. 36. Cybilla hubbardana (holotype), Tucson, Arizona, USNM slide 23833. 37. Honca grandis, Wellsville, Utah, USNM slide 153,599. 38. Nycthia pimama, Miller Canyon, Arizona, slide MGP 660.