Anacochylidia maderana Brown, 2019

Brown, John W., 2019, New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae), Zootaxa 4671 (2), pp. 195-222 : 198

Anacochylidia maderana Brown , new species

Figs. 1 , 19 , 33

Diagnosis. Anacochylidia maderana is a small species with a narrow triangular forewing and somewhat distinctive maculation: the ground color is white, with an ill-defined, pale rust patch in the subterminal region and an ill-defined, concolorous, oblique dash from the middle of the hind margin extending to about the middle of the discal cell ( Fig. 1 ). The species is superficially similar to Cochylis temerana ( Busck, 1907) and Mimcochylis fulvotincatana ( Walsingham, 1884) , but the genitalia have little in common with those species. Anacochylidia maderana is easily distinguished by the male genitalia ( Fig. 19 ), which have a broad, concavely rounded top of the tegumen; a very short, wide valva similar to species of Cochylidia ; and a long, curved, slender, apically-attenuate rod from the phallus, extending ca. 0.5 the length of the phallus. The male genitalia lack the conspicuous elongate process from the base of the costa of the valva that is present in all species Cochylidia .

Description. Head: Frons and vertex buff; labial palpus light buff with scattered fuscous scales, cream on inner surface; antenna mixed fuscous and cream, sensory setae about 0.9 times flagellomere diameter in male, shorter in female. Thorax: Mesonotum and tegula buff in anterior 0.3, remainder pale fuscous; forewing length 4.5–4.8 mm (n = 4); forewing ground white, with ill-defined, pale rust, submedian dash extending from near middle of hind margin to lower edge of discal cell; small concolorous spot at costa ca. 0.6 distance from base to apex; subterminal area with broad concolorous patch extending from costa almost to tornus. Fringe white. Hindwing cream-white. Fringe concolorous with wing. Abdomen: Dorsal scaling brown, venter shiny white. Male genitalia ( Fig. 19 ) as described for genus. Female genitalia ( Fig. 33 ) as described for genus.

Types. Holotype ³, Madera Canyon , [Santa Cruz Co.], Arizona, 3‒5 Sep 1969, J. Powell ( EME) . Paratypes (6³, 1♀). USA: Arizona: Cochise Co.: Palmerlee, no date (5³, 1♀) ( USNM). Santa Cruz Co. : Madera Canyon, 3‒5 Sep 1969 (1³), J. Powell ( EME) .

Etymology. The specific epithet refers to the type locality of Madera Canyon in Arizona.

Distribution and biology. This species is known from Palmerlee (Cochise Co.) (n = 7) (USNM) and Madera Canyon (Santa Cruz Co.) (n = 2) (EME) of southern Arizona. The only two specimens with capture data are from September. Nothing is known of the early stages.

Figures

FIGURES 1–10. Adults of Cochylini genera. 1. Anacochylidia maderana, Madera Canyon, Santa Rita Mountains, Arizona (EME). 2. Cagiva cephalanthana, Dorman Lake, Oktibbeha Co., Mississippi (USNM). 3. Atroposia oenotherana, Houston, Harris Co., Texas (USNM). 4. Atroposia oenotherana, Radisson, Onondaga Co., New York (USNM). 5. Cybilla hubbardana, Shafter, Presidio Co., Texas (USNM). 6. Cybilla hubbardana, Lordsburg, Hidalgo Co., New Mexico (CNC). 7. Honca grandis, Chimney Gulch, Golden, Colorado (USNM). 8. Pogospinia floridana, Long Key State Park, Monroe Co., Florida (USNM). 9. Nycthia pimama, Portal, Cochise Co., Arizona (USNM). 10. Nycthia yuccatana, Falcon State Park, Starr Co., Texas (USNM).

FIGURES 19–26. Male genitalia of Cochylini genera. 19. Anacochylidia maderana, Palmerlee, Arizona, USNM slide 23,359. 20. Atroposia oenotherana, Padre Island National Seashore, Texas, USNM slide 153,596. 21. Cagiva cephalanthana, Dorman Lake, Mississippi, USNM slide 150,530. 22. Cybilla hubbardana, Tucson, Arizona, USNM slide 29,397. 23. Honca grandis, Ogden Canyon, Utah, USNM slide 153,598. 24. Nycthia pimama, Blair Valley, California, USNM slide 144,042. 25. Nycthia yuccatana, Laguna Atacosa, Texas, USNM slide 120,405. 26. Pogospinia floridana, Key Largo, Florida, USNM slide 23,249.

FIGURES 33–38. Female genitalia of Cochylini genera. 33. Anacochylidia maderana, Palmerlee,Arizona, USNM slide 23,358. 34. Atroposia oenotherana, Padre Island National Seashore, Texas, USNM slide 153,597. 35. Cagiva cephalanthana, Dorman, Mississippi, USNM slide 153,609. 36. Cybilla hubbardana (holotype), Tucson, Arizona, USNM slide 23833. 37. Honca grandis, Wellsville, Utah, USNM slide 153,599. 38. Nycthia pimama, Miller Canyon, Arizona, slide MGP 660.