Cephennula medioglabra, Jałoszyński, Paweł, 2011

Jałoszyński, Paweł, 2011, Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia, Zootaxa 3113, pp. 36-52 : 48-49

Cephennula medioglabra sp. n.

( Figs. 9 , 25, 26 )

Holotype: W MALAYSIA: 3, two labels: "W. Malaysia: Pahang \ Cameron Highlands \ 1500m, trails 4, 13: #15 \ Löbl & Calame, 23.3.93 " [white, printed]; " CEPHENNULA \ medioglabra m. \ det. P. Jałoszyński, 2011 \ HOLO- TYPUS " [red, printed] ( MHNG). Paratypes (533, 2ƤƤ): 13, "W. Malaysia:Pahang # 16a \ Cameron Highls 1600m \ Bukit Mentiga, trail 14 \ Löbl&Calame, 23.3.93 " [white, printed]; 333, 2ƤƤ, "W. Malaysia:Pahang # 18b \ Cameron Highls 1550m \ Gunung Jasar, trail 11 \ Löbl&Calame, 24.3.93 "; 13, "W. Malaysia:Pahang \ Cameron Highls 1720m \ Gunung Jasar # 19b \ Löbl&Calame, 25.3.93 " ( MHNG, PCPJ).

Diagnosis. Body length ca. 0.8 mm, not reaching 0.9 mm; antennal club strongly flattened and loosely assembled; head unevenly covered with large and deep punctures, dense on sides of vertex and frons; median part of vertex and frons impunctate; median part of pronotum with large, deep and dense punctures; sublateral pronotal carinae very fine and obliterated in anterior 1/3; subhumeral carinae about as long as 1/ 5x EL, slightly shorter than humeral carinae; males with dense subapical setal field on each elytron; aedeagus in ventral view bottle-shaped with nearly straight apical margin, endophallus very lightly pigmented except of distinct pair of apical lobes slightly curved mesally.

Description. Body of male ( Fig. 9 ) moderately convex, elongate, with barely marked constriction between pronotum and elytra, moderately glossy, light brown to yellowish-brown, covered with yellowish vestiture, legs and antennae slightly lighter. BL 0.79–0.83 mm (mean 0.81 mm).

Head broadest at large, coarsely faceted and strongly convex eyes, HL 0.09–0.10 mm (mean 0.09 mm), HW 0.19–0.20 mm (mean 0.19 mm); vertex and frontoclypeal area convex; supraantennal tubercles barely marked. Punctures on dorsum of head very large, deep and sharply defined, those on sides of vertex and frons dense, narrow median area on vertex and frons impunctate; setae short and sparse, suberect. Antennae with strongly enlarged, strongly flattened, loosely assembled and distinctly 3-segmented club, AnL 0.33 mm; antennomere I about 1.4x as long as broad; II 2.2x as long as broad; III slightly broader than long; IV–VII subequal in length and width, each about 1.4x as long as broad; VIII as broad as VII but distinctly shorter, 2.2x as long as broad; IX much broader and longer than VIII, slightly transverse; X much broader and longer than IX, strongly transverse; XI nearly as broad as X, 1.6x as long as broad.

Pronotum approximately trapezoidal, broadest near anterior 1/3; PL 0.23–0.25 mm (mean 0.24 mm), PW 0.33–0.34 mm (mean 0.33 mm), anterior margin broadly rounded; lateral margins distinctly microserrate, strongly rounded in anterior 1/3, distinctly concave in posterior 1/3 and nearly parallel near nearly straight but acute hind angles; posterior margin very weakly bisinuate; sublateral carinae very fine and obliterated in anterior 1/3. Transverse ante-basal groove shallow but sharply marked and nearly straight; sublateral and lateral ante-basal foveae equally small and relatively deep. Punctures on median part of pronotal disc similar in diameter and depth to those on sides of frons, dense; punctures between sublateral carinae and lateral margins of pronotum distinctly larger and denser than those in middle, nearly adjacent to each other, but not coarse; setae sparse, moderately long, suberect.

Elytra moderately convex, oval, broadest near anterior 1/3; EL 0.46–0.48 mm (mean 0.47 mm), EW 0.38–0.39 mm (mean 0.38 mm), EI 1.23–1.27 (mean 1.24); base of each elytron with very small but distinct basal fovea located nearly in middle between scutellum and base of humeral carina; subhumeral carinae very fine, about as long as 1/ 5x EL; humeral carinae slightly more distinct and slightly longer. Punctures on elytra smaller and shallower than those on head and pronotum, but relatively distinct and dense, especially on circumsutural area in anterior half; setae similar as those on pronotum, except for small subapical field on each elytron composed of dense and nearly recumbent setae directed towards suture.

Legs moderately long and slender; pro- and mesotibiae nearly straight, metatibiae slightly recurved.

Aedeagus ( Figs. 25, 26 ) in ventral view bottle-shaped with broad, subtrapezoidal apex and nearly straight apical margin; AeL 0.18 mm. Endophallus lightly pigmented, with two broad and rounded elongate apical lobes curved mesally; parameres slender and very short, not exceeding apex of median lobe, each bearing single apical seta.

Female. Externally differs from male in lacking subapical setal field on each elytron and slightly more slender body. BL 0.85 mm; HL 0.10 mm, HW 0.18 mm, AnL 0.33 mm; PL 0.25 mm, PW 0.34 mm; EL 0.50 mm, EW 0.39 mm, EI 1.29.

Distribution. West Malaysia (Pahang).

Etymology. The specific epithet “ medioglabra ” refers to the impunctate (Latin glaber, "smooth") median part of the vertex and frons.

Remarks. This is the only known species of Cephennula (and the Cephennomicrus group of genera) with modified elytra in males, bearing a dense subapical setal field. Similar modifications in Cephenniini were known previously only in some species of Cephennodes Reitter, 1884 . Cephennula medioglabra has the aedeagus most similar to that of C. scaphisoma , also occurring in Pahang ( Jałoszyński 2008), but details of the endophallus clearly differ in the latter species.

Figures

FIGURES 8 – 12. Habitus of males. 8 ― Cephennula titiwangsana sp. n.; 9 ― Cephennula medioglabra sp. n.; 10 ― Cephennula sibugana sp. n.; 11 ― Cephennula gombakiana sp. n.; 12 ― Cephennula micropunctata sp. n. Scale bars: 0.2 mm.

FIGURES 25 – 32. Aedeagus in ventral (25, 27, 29, 31) and lateral (26, 28, 30, 32) views. 25, 26 ― Cephennula medioglabra sp. n.; 27, 28 ― Cephennula sibugana sp. n.; 29, 30 ― Cephennula gombakiana sp. n.; 31, 32 ― Cephennula micropunctata sp. n. Scale bars: 0.05 mm.