Cryphalus himalayensis, Buhroo, 2024

Buhroo, Abdul Ahad, 2024, A taxonomic monograph of subfamily Scolytinae (Coleoptera: Curculionidae) in the Western Himalaya, Zootaxa 5533 (1), pp. 1-82 : 3

Abstract................................................................................................... 5 Introduction................................................................................................ 5 Materials and methods....................................................................................... 5 PCR and Sequencing......................................................................................... 7 Data analysis............................................................................................... 7 Results and discussion....................................................................................... 8 Tribe Corthylini LeConte, 1876 ................................................................................ 8 Subtribe Pityophthorina Eichhoff, 1878 .......................................................................... 8 Genus Pityophthorus Eichhoff, 1864 ............................................................................ 8 Key to the Western Himalayan species of Pityophthorus Eichhoff, 1864 ................................................ 8 1. Pityophthorus cedri Wood, 1989 ............................................................................. 8 2. Pityophthorus chilgoza Wood, 1989 .......................................................................... 9 3. Pityophthorus deodara ( Stebbing, 1903) ...................................................................... 10 Tribe Cryphalini Lindemann, 1877 ............................................................................. 11 Genus Cryphalus Erichson, 1836 .............................................................................. 11 Key to the Western Himalayan species of Cryphalus Erichson, 1836 .................................................. 11 4. Cryphalus fulmineus Wood, 1989 ........................................................................... 11 5. Cryphalus himalayensis sp. nov............................................................................. 12 6. Cryphalus major Stebbing, 1903 ............................................................................ 13 7. Cryphalus mangiferae Stebbing, 1914 ....................................................................... 14 8. Cryphalus strohmeyeri Stebbing, 1914 ....................................................................... 15 Tribe Crypturgini LeConte, 1876 .............................................................................. 16 Genus Crypturgus Erichson, 1836 ............................................................................. 16 Key to the Himalayan species of Crypturgus Erichson, 1836 ........................................................ 16 9. Crypturgus beesoni Eggers, 1936 ........................................................................... 16 10. Crypturgus pusillus ( Gyllenhal, 1813) ....................................................................... 17 Tribe Diamerini Hagedorn, 1909 .............................................................................. 18 Genus Sphaerotrypes Blandford, 1894 .......................................................................... 18 Key to the Western Himalayan species of Sphaerotrypes Blandford, 1894 (reproduced after Buhroo & Knížek, 2020)........... 18 11. Sphaerotrypes montanus Buhroo & Knížek, 2020 ............................................................. 18 12. Sphaerotrypes querci Stebbing, 1908 ....................................................................... 19 Tribe Dryocoetini Lindemann, 1877 ............................................................................ 20 Genus Coccotrypes Eichhoff, 1878a ............................................................................ 20 Key to the Western Himalayan species of Coccotrypes Eichhoff, 1878a ................................................ 21 13. Coccotrypes cinnamomi ( Eggers, 1936) ..................................................................... 21 14. Coccotrypes papuanus ( Eggers, 1923) ....................................................................... 22 Genus Dryocoetes Eichhoff, 1864 ............................................................................. 23 Key to the Himalayan species of Dryocoetes Eichhoff, 1864 ........................................................ 23 15. Dryocoetes asperatus sp. nov.............................................................................. 23 16. Dryocoetes brownei Mandelshtam & Petrov, 2010b ............................................................ 25 17. Dryocoetes himalayensis Strohmeyer, 1908a ................................................................. 25 18. Dryocoetes indicus Stebbing, 1914 ......................................................................... 26 19. Dryocoetes quadrisulcatus Strohmeyer, 1908c ................................................................ 27 Genus Taphrorychus Eichoff, 1878b ........................................................................... 28 Key to the Himalayan species of Taphrorychus Eichoff, 1878b ...................................................... 28 20. Taphrorychus betulae Schedl, 1974 ......................................................................... 29 21. Taphrorychus hewetti ( Stebbing, 1908) ...................................................................... 29 Tribe Ernoporini Nüsslin, 1911 ................................................................................ 31 Genus Eidophelus Eichhoff, 1876 ............................................................................. 31 22. Eidophelus indicus ( Wood, 1989) .......................................................................... 31 Genus Ernoporus Thomson, 1859 ............................................................................ 31 23. Ernoporus squamosus sp. nov............................................................................. 32 Tribe Hylesinini Erichson, 1836 ............................................................................... 33 Genus Hylesinus Fabricius, 1801 .............................................................................. 33 24. Hylesinus macmahoni ( Stebbing, 1909) ..................................................................... 33 Tribe Hylurgini Gistel, 1848 .................................................................................. 35 Genus Hylurgus Latreille, 1806 ............................................................................... 35 25. Hylurgus indicus Wood, 1985 ............................................................................. 35 Genus Xylechinus Chapuis, 1869 .............................................................................. 36 26. Xylechinus padus Wood, 1988a ............................................................................ 36 Tribe Ipini Bedel, 1888 ...................................................................................... 37 Genus Ips DeGeer, 1775 ..................................................................................... 37 Key to the Western Himalayan species of Ips DeGeer, 1775 (reproduced after Buhroo & Lakatos, 2011)...................... 37 27. Ips longifolia ( Stebbing, 1909) ............................................................................. 37 28. Ips stebbingi Strohmeyer, 1908b ........................................................................... 38 Genus Pityogenes Bedel, 1888 ................................................................................ 39 Key to the Himalayan species of Pityogenes Bedel, 1888 ........................................................... 39 29. Pityogenes scitus Blandford, 1893a ......................................................................... 39 30. Pityogenes spessivtsevi Lebedev, 1926 ....................................................................... 40 Tribe Micracidini LeConte, 1876 .............................................................................. 41 Genus Pseudothysanoes Blackman, 1920 ....................................................................... 41 31. Pseudothysanoes kashmirica Buhroo and Knížek, 2020 ......................................................... 42 Tribe Polygraphini Chapuis, 1869 ............................................................................. 43 Genus Carphoborus Eichhoff, 1864 ............................................................................ 43 Key to the Himalayan species of Carphoborus Eichhoff, 1864 ....................................................... 43 32. Carphoborus costatus Wichmann, 1915 ..................................................................... 44 33. Carphoborus zhobi ( Stebbing, 1909) ........................................................................ 44 Genus Polygraphus Erichson, 1836 ............................................................................ 45 Key to the species of Himalayan Polygraphus Erichson, 1836 ...................................................... 46 34. Polygraphus aterrimus Strohmeyer, 1908b ................................................................... 46 35. Polygraphus difficilis Wood, 1988b ......................................................................... 47 36. Polygraphus longifolia Stebbing, 1903 ...................................................................... 48 37. Polygraphus major Stebbing, 1903 ......................................................................... 49 38. Polygraphus pini Stebbing, 1914 ........................................................................... 49 39. Polygraphus setosus Schedl, 1979 .......................................................................... 50 40. Polygraphus trenchi Stebbing, 1905 ........................................................................ 51 Tribe Scolytini Latreille, 1804 ................................................................................ 53 Genus Scolytus Geoffroy, 1762 ............................................................................... 53 41. Scolytus kashmirensis Schedl, 1958 ........................................................................ 53 42. Scolytus major Stebbing, 1903 ............................................................................. 54 43. Scolytus nitidus Schedl, 1936a ............................................................................. 55 44. Scolytus stepheni Mandelstam and Petrov, 2010a .............................................................. 56 Tribe Scolytoplatypodini Blandford, 1893 ....................................................................... 57 Genus Scolytoplatypus Schaufuss, 1891 ........................................................................ 57 45. Scolytoplatypus daimio Blandford, 1893b .................................................................... 58 46. Scolytoplatypus darjeelingi Stebbing, 1914 ................................................................... 59 47. Scolytoplatypus denticauda sp. nov......................................................................... 59 48. Scolytoplatypus kunala Strohmeyer, 1908a ................................................................... 61 49. Scolytoplatypus minimus Hagedorn, 1904 .................................................................... 62 50. Scolytoplatypus raja Blandford, 1893b ( Figure 63 )............................................................. 63 Tribe Trypophloeini Nüsslin, 1911 ............................................................................. 64 Genus Hypothenemus Westwood, 1834 ......................................................................... 64 Key to the Western Himalayan species of Hypothenemus Westwood, 1834 ............................................. 64 51. Hypothenemus birmanus ( Eichhoff, 1878a) .................................................................. 64 52. Hypothenemus ficivorus sp. nov........................................................................... 65 Tribe Xyleborini LeConte, 1876 .............................................................................. 66 Genus Cyclorhipidion Hagedorn, 1912 ......................................................................... 67 53. Cyclorhipidion inarmatum ( Eggers, 1923) ................................................................... 67 Genus Diuncus Hulcr & Cognato, 2009 ......................................................................... 68 54. Diuncus corpulentus ( Eggers, 1930) ........................................................................ 68 Genus Xyleborinus Reitter, 1913 .............................................................................. 69 55. Xyleborinus saxesenii ( Ratzeburg, 1837) .................................................................... 69 Genus Xyleborus Eichhoff, 1864 .............................................................................. 70 56. Xyleborus perforans ( Wollaston, 1857) ...................................................................... 71 Genus Xylosandrus Reitter, 1913 .............................................................................. 72 57. Xylosandrus crassiusculus ( Motschulsky, 1866) ............................................................... 72 The endemism and distribution of scolytine fauna in the Western Himalaya............................................. 75 Acknowledgement......................................................................................... 76 References................................................................................................ 76

Figures

FIGURE 63. Scolytoplatypus raja. Male: dorsal view A, lateral view B, frons C; female: dorsal view D.