Ephippiochthonius

Gardini, Giulio, 2013, A revision of the species of the pseudoscorpion subgenus Chthonius (Ephippiochthonius) (Arachnida, Pseudoscorpiones, Chthoniidae) from Italy and neighbouring areas, Zootaxa 3655 (1), pp. 1-151 : 6-9

Key to adults of the Italian, Swiss Tessin and Corsican species of Ephippiochthonius

1 Proximal half of fixed chelal finger with triangular teeth gradually reduced as far as the base of the finger (figs 203–204, 225– 226); proximal half of movable chelal finger with low, rounded, sometime vestigial teeth, consequently without straight, thick marginal lamina (figs 203–204, 225–226) ( C. tetrachelatus group)............................................... 2

- Fixed chelal finger with triangular subequal teeth, followed by few rounded teeth, as far as the proximal third or half of the finger (figs 169–170); proximal third or half of movable chelal finger with straight, thick marginal lamina, sometimes with low and rounded vestigial teeth (more or less fused with high base in C. parmensis , C. cassolai , C. berninii and C. thaleri ) (figs 169–170, 177–178) ( C. fuscimanus group)................................................................. 15

2 Movable cheliceral finger with isolated subapical tooth (di) (figs 4–5); no epistome; 4 eyes with lens.................. 3

- Movable cheliceral finger without isolated subapical tooth (di) (figs 31–32); epistome and eyes present or absent........ 5

3 Carapace with 1 or 2 preocular microsetae on each side; posterior margin of carapace with 2 macrosetae and 1–2 lateral microsetae on each side............................................................................... 4

- Carapace without preocular microsetae; posterior margin of carapace with 2 macrosetae, without lateral microsetae.......................................................................................... [ C. gibbus Beier, 1953 ]

4 Larger: chela length (3) 0.655–0.77 mm; movable chelal finger with 11–14 teeth; distance between successive apices of teeth of fixed chelal finger at level of est-it 0.028–0.034 mm; carapace with anteromedian macrosetae 0.080–0.090 mm long; movable cheliceral finger with spinneret in both sexes (figs 141–142)........................... C. lucanus Callaini, 1984

- Smaller: length of chela (3) 0.60–0.62 mm; movable chelal finger with 8–9 triangular teeth; distance between successive api-

ces of teeth of fixed chelal finger at level of est-it 0.025–0.026 mm; carapace with anteromedian macrosetae 0.066–0.070 mm long; movable cheliceral finger without spinneret in males (fig. 153)............................ C. messapicus n. sp. 5 Movable cheliceral finger with spinneret in both sexes, rarely reduced in males (male of C. altamurae unknown); distance between successive apices of teeth of fixed chelal finger at level of est - it no more than 0.049 mm; epigean and hypogean species from Italy and epigean species from Corsica........................................................... 6

- Movable cheliceral finger without spinneret in males; distance between successive apices of teeth of fixed chelal finger at level of est - it 0.050 mm; hypogean troglomorphic species from Corsica......................... [ C. remyi Heurtault, 1975 ]

6 Epigean species; anterior eyes with convex lens; small or medium size: length of chela 0.48–0.95 (3) 0.53–1.04 (Ƥ) mm, length of movable chelal finger 0.28–0.56 (3) 0.30–0.60 (Ƥ) mm; chela 4.4–6.0 (3) 4.0–5.2 (Ƥ) times as long as deep; ratio of pedipalpal femur/carapace 1.1–1.5; posterior margin of carapace with 2 macrosetae or with 2 macrosetae and a pair of lateral microsetae.......................................................................................... 7

- Endogean or hypogean, micro- or anophthalmic species; medium or large size: length of chela 0.94–1.83 (3) 0.92–1.77 (Ƥ) mm, length of movable chelal finger 0.55–1.14 (3) 0.53–1.07 (Ƥ) mm; chela 4.9–7.9 (3) 4.5–7.4 (Ƥ) times as long as deep; ratio of pedipalpal femur/carapace 1.5–2.0; posterior margin of carapace with 2 macrosetae......................... 9

7 Smaller: length of chela 0.48–0.72 (3) 0.53–0.81 (Ƥ) mm, length of movable chelal finger 0.28–0.41 (3) 0.30–0.45 (Ƥ) mm; ratio of pedipalpal femur/carapace 1.1–1.3; hand of chela depressed at level of trichobothria ib - isb and with hump distad of ib - isb (figs 203–204); fixed chelal finger at level of est-it with 4–6 teeth occupying 0.1 mm (distance between successive apices 0.019–0.022 mm).................................................................................... 8

- Larger: length of chela 0.65–0.95 (3) 0.55–1.04 (Ƥ) mm, length of movable chelal finger 0.38–0.56 (3) 0.39–0.60 (Ƥ) mm; ratio of pedipalpal femur/carapace 1.3–1.5; hand of chela not depressed (or with a very weak depression) at level of trichobothria ib - isb, without hump distad of ib - isb (figs 225–226, 233–234); fixed chelal finger at level of est-it with 3–4 (rarely 5) teeth occupying 0.1 mm (distance between successive apices 0.024–0.040 mm)............. C. tetrachelatus (Preyssler, 1790)

8 Posterior margin of carapace with 2 macrosetae (exceptionally with 2 macrosetae and pair of lateral microsetae); proximal half of chelal fingers with teeth reduced in size proximally, but clearly distinguishable as far as the base of fingers (figs 203–204)................................................................................... C. siculus Beier, 1961

- Posterior margin of carapace with 2 macrosetae and a pair of lateral microsetae; proximal half of chelal fingers with vestigial teeth, widely rounded proximally, reduced as far as trichobothrium sb, not reaching base of fingers (figs 268–269)................................................................................... C. vachoni Heurtault-Rossi, 1963

9 Fixed chelal finger with 18–25 triangular teeth (proximal rounded teeth excepted); movable chelal finger with 5–12 triangular teeth (proximal rounded or vestigial teeth excepted)........................................................ 10

- Fixed chelal finger with 33–36 triangular teeth (proximal rounded teeth excepted); movable chelal finger with 16–18 triangular teeth (proximal rounded or vestigial teeth excepted) (fig. 194).............................. C. sardous Gardini, 2008

10 Anophthalmous (rarely microphthalmic: C. poseidonis ) hypogean species from Apulia, Sicily and Sardinia; pedipalps more slender: chela 6.4–7.4 (3) 5.6–6.5 (Ƥ) times as long as deep................................................. 11

- Microphthalmic endogean species (locally hypogean in Sardinia) from Liguria, Sicily, Malta and Sardinia; pedipalps less slen- der: chela 4.9–6.1 (3) 4.5–6.0 (Ƥ) times as long as deep..................................... C. concii Beier, 1953

11 Species from Italian mainland (Apulia); carapace with 1 preocular microseta on each side.......................... 12

- Species from Sicily and Sardinia; carapace usually with 2 preocular microsetae on each side (variable in C. poseidonis ).. 13

12 Species of medium size: length of chela 0.92 (Ƥ) mm, length of movable chelal finger 0.53 (Ƥ) mm; pedipalps less slender: chela 5.9 times as long as deep; ratio of pedipalpal femur/carapace 1.6............................ C. altamurae n. sp.

- Species of large size: length of chela 1.78 (3) mm, length of movable chelal finger 1.05 (3) mm; pedipalps more slender: chela 7.4 times as long as deep; ratio of pedipalpal femur/carapace 2.0........................ C. apulicus Beier, 1958

13 Species from Sardinia; larger: length of chela 1.21–1.41 (3) 1.16–1.60 (Ƥ) mm, length of movable chelal finger 0.71–0.85 (3) 0.67–0.94 (Ƥ) mm; fixed chelal finger at level of est-it with 3 teeth occupying 0.1 mm (distance between successive apices 0.035–0.042 mm)................................................................................... 14

- Species from Sicily; smaller: length of chela 1.02 (3) mm, length of movable chelal finger 0.63 (3) mm; fixed chelal finger at level of est-it with 4 teeth occupying 0.1 mm (distance between successive apices 0.027 mm)............. C. elymus n. sp.

14 Anophthalmic species with marked troglomorphic facies; chela 6.85 (3) 6.1–6.5 (Ƥ) times as long as deep; ratio of pedipalpal femur/carapace 1.7–1.9; fixed chelal finger with 18–22 triangular teeth (proximal rounded teeth excepted)................................................................................................ C. grafittii Gardini, 1981

- Microphthalmic or oculated species with weak troglomorphic facies; chela 6.4–6.7 (3) 5.6–6.1 (Ƥ) times as long as deep; ratio of pedipalpal femur/carapace 1.5–1.8; fixed chelal finger with 21–25 triangular teeth (proximal rounded teeth excepted).................................................................................. C. poseidonis Gardini, 1993

15 Movable cheliceral finger with isolated subapical tooth (di).................................................. 16

- Movable cheliceral finger without isolated subapical tooth (di)............................................... 34

16 Chelal hand (in lateral view) with a rounded hump distad of trichobothria ib-isb and a more or less marked hollow just before base of movable finger, making hand abruptly constricted; proximal half of movable chelal finger with thick marginal lamina, with teeth rounded or vestigial and more or less fused with high base (figs 177–178, 240–241)...................... 17

- Chelal hand (in lateral view) without a rounded hump distad of trichobothria ib-isb, sometimes weakly depressed at level of ib/ isb, without hollow just before base of movable finger, making hand gradually constricted; proximal half of movable chelal finger with straight, thick marginal lamina, sometimes undulated (figs 74–75, 119–120)........................... 20

17 Posterior margin of carapace with 4 macrosetae (lateral macrosetae slightly shorter, exceptionally 2 macro- and 2 microsetae in C. parmensis ); ocular series with 6 macrosetae (fig. 173).................................................... 18

- Posterior margin of carapace with 2 macrosetae; ocular series with 4 macrosetae and 2 microsetae (fig. 237)................................................................................................ C. thaleri Gardini, 2009

18 Epigean species with eyes; carapace with 2 preocular microsetae on each side; fixed chelal finger with 12–16 triangular teeth (proximal rounded teeth excepted); smaller: length of chela 0.45–0.57 (3) 0.50–0.66 (Ƥ) mm, length of movable chelal finger 0.26–0.35 (3) 0.28–0.40 (Ƥ) mm; pedipalps less slender: chela 4.0–5.3 (3) 3.6–4.6 (Ƥ) times as long as deep; ratio of pedipalpal femur/carapace 1.0–1.2........................................................................... 19

- Endogean or hypogean, anophthalmic or microphthalmic species; carapace with 1 preocular microseta on each side; fixed chelal finger with 17–21 triangular teeth (proximal rounded teeth excepted); larger: length of chela 0.89–1.12 (3) 0.84–1.37 (Ƥ) mm, length of movable chelal finger 0.54–0.68 (3) 0.50–0.78 (Ƥ) mm; pedipalps more slender: chela 5.7–5.9 (3) 5.0–5.1 (Ƥ) times as long as deep; ratio of pedipalpal femur/carapace 1.4–1.6.......................... C. cassolai Beier, 1973

19 Proximal half of fixed chelal finger with 2–5 rounded teeth merging in an evident high bulge of marginal lamina (figs 177– 178); species from Italian mainland................................................... C. parmensis Beier, 1963

- Proximal half of fixed chelal finger with 2–5 rounded and spaced teeth, marginal lamina weakly raised (figs 26–27); species from Sicily and Sardinia............................................................ C. berninii Callaini, 1983

20 Epigean species with eyes (or microphthalmic, with posterior eyes reduced to ocular spot: C. intemelius ); movable chelal finger with 5–8 triangular teeth.......................................................................... 21

- Hypogean or endogean anophthalmic species; movable chelal finger with (6)7–14 triangular teeth................... 24

21 Smaller: length of chela 0.435–0.65 (3) 0.47–0.73 (Ƥ) mm, length of movable chelal finger 0.25–0.37 (3) 0.265–0.40 (Ƥ) mm; pedipalps less slender: ratio of pedipalpal femur/carapace 1.1–1.3; fixed chelal finger at level of est-it with 5 teeth occupying 0.1 mm (distance between successive apices 0.020–0.031 mm).......................................... 22

- Larger: length of chela 1.0–1.12 (3) 1.13–1.25 (Ƥ) mm, length of movable chelal finger 0.52–0.58 (3) 0.58–0.63 (Ƥ) mm; pedipalps more slender: ratio of pedipalpal femur/carapace 1.3–1.7; fixed chelal finger at level of est-it with 3–4 teeth occupying 0.1 mm (distance between successive apices 0.043–0.050 mm)........................ C. fuscimanus Simon, 1900

22 Anterior eyes with distinctly convex lens; epistome mostly absent (rarely present in C. tyrrhenicus ); movable cheliceral finger with spinneret in both sexes............................................................................23

- Anterior eyes with weakly convex lens, posterior ones reduced to ocular spot, without tapetum; epistome mostly present; movable cheliceral finger with spinneret only in females (sometimes vestigial in 3: figs 116–117)......... C. intemelius n. sp.

23 Pedipalpal hand (in lateral view) with gentle slope between trichobothria ib-isb and eb-esb-ist (hand with a more rectangular profile: figs 169–170); generally larger: length of chela 0.52–0.65 (3) 0.56–0.73 (Ƥ) mm, length of movable chelal finger 0.29–0.37 (3) 0.31–0.40 (Ƥ) mm; distance between successive apices of teeth of fixed chelal finger at level of est-it 0.026– 0.031 mm .......................................................................... C. nanus Beier, 1953

- Pedipalpal hand (in lateral view) with abrupt slope between trichobothria ib-isb and eb-esb-ist (hand with a more square profile: figs 259-260); generally smaller: length of chela 0.435–0.54 (3) 0.47–0.60 (Ƥ) mm, length of movable chelal finger 0.25– 0.31 (3) 0.265–0.34 (Ƥ) mm; distance between successive apices of teeth of fixed chelal finger at level of est-it 0.020 mm ....................................................................................... C. tyrrhenicus n. sp.

24 Hypogean species of medium or large size: length of chela 0.75–1.70 (3) 0.75–1.86 (Ƥ) mm, length of movable chelal finger 0.44–0.96 (3) 0.44–1.07 (Ƥ) mm; pedipalps more slender: chela 6.4–8.4 (3) 5.3–8.45 (Ƥ) times as long as deep; ratio of pedipalpal femur/carapace 1.3–2.05 (Ƥ); fixed chelal finger with 12–21 triangular teeth, at level of est-it with 3–4 teeth occupying 0.1 mm (distance between successive apices 0.025–0.048 mm)............................................ 25

- Endogean species of small size: length of chela 0.49–0.51 (3) 0.55–0.565 (Ƥ) mm, length of movable chelal finger 0.29–0.30 (3) 0.32–0.325 (Ƥ) mm; pedipalps less slender: chela 5.6–5.7 (3) 5.0–5.1 (Ƥ) times as long as deep; ratio of pedipalpal femur/ carapace 1.2 (Ƥ); fixed chelal finger with 10–11 triangular teeth, at level of est-it with 5 teeth occupying 0.1 mm (distance between successive apices 0.020–0.021 mm)................................................... C. aeneae n. sp.

25 Species from Northern Italy........................................................................... 26

- Species from Central Italy............................................................................ 31

26 Species from southwestern Alps and Ligurian Apennines.................................................... 27

- Species from southeastern Alps (from Bergamasque pre-Alps to Carnic Alps)................................... 30

27 Movable cheliceral finger, besides the isolated subapical tooth (di), with a median prominent group of 5 teeth (figs 272–273); species from Ligurian Apennines....................................................................... 28

- Movable cheliceral finger, besides the isolated subapical tooth (di), with 7–12 teeth (figs 245-247); species from Maritime and Ligurian Alps and Ligurian pre-Alps.................................................................... 29

28 Movable cheliceral finger with isolated subapical tooth (di) proximal with respect to the spinneret (fig. 87); smaller: length of chela 0.75 (Ƥ) mm, length of movable chelal finger 0.44 (Ƥ) mm; species from the neighbourhood of Genoa............................................................................................ C. genuensis Gardini, 1990

- Movable cheliceral finger with isolated subapical tooth (di) at same level as spinneret (figs 272-273); larger: length of chela 0.75–1.06 (3) 0.92–1.07 (Ƥ) mm, length of movable chelal finger 0.45–0.63 (3) 0.54–0.62 (Ƥ) mm; species from Eastern Liguria............................................................................... C. zoiai Gardini, 1990

29 Movable cheliceral finger of 3 without spinneret (fig. 245); smaller: length of chela 1.0–1.29 (3) 0.77–1.39 (Ƥ) mm, length of movable chelal finger 0.60–0.745 (3) 0.455–0.80 (Ƥ) mm; pedipalps less slender: chela 6.7–7.6 (3) 5.45–6.8 (Ƥ); ratio of pedipalpal femur/carapace 1.6–1.85 (3) 1.3–1.7 (Ƥ); fixed and movable chelal fingers respectively with 13–17 and 8–11 triangular teeth, at level of est-it with 3–4 teeth occupying 0.1 mm (distance between successive apices 0.027–0.031 mm)...................................................................................... C. troglophilus Beier, 1930

- Movable cheliceral finger of 3 with spinneret (fig. 90); larger: length of chela 1.50–1.70 (3) 1.65–1.86 (Ƥ) mm, length of

pedipalpal movable finger 0.86–0.96 (3) 0.955–1.07 (Ƥ) mm; pedipalps more slender: chela 7.5–8.1 (3) 7.25–8.45 (Ƥ) times as long as deep; ratio of pedipalpal femur/carapace 1.8–2.05 (Ƥ); pedipalpal fixed and movable fingers respectively with 18– 21 and 11–14 triangular teeth, at level of est-it with 2–3 teeth occupying 0.1 mm (distance between successive apices 0.040 mm)............................................................................. C. gestroi Simon, 1896 30 Tergite I with 4 setae; movable cheliceral finger of 3 without spinneret; isolated subapical tooth (di) of Ƥ proximal with respect to the spinneret (figs 182–183); smaller: length of chela 0.75–1.04 (3) 0.93–1.14 (Ƥ) mm, length of movable chelal finger 0.44–0.59 (3) 0.57–0.66 (Ƥ) mm; pedipalps less slender: chela 6.5–7.4 (3) 6.3–6.9 (Ƥ) times as long as deep; ratio of pedipalpal femur/carapace 1.3–1.7 (Ƥ); fixed and movable chelal fingers respectively with 12–14 and 7–9 triangular teeth, fixed finger at level of est-it with 3–4 teeth occupying 0.1 mm (distance between successive apices 0.030–0.036 mm)......................................................................................... C. pieltaini Beier, 1930

- Tergite I with 2 setae; movable cheliceral finger with spinneret in both sexes; isolated subapical tooth (di) level with the spinneret (figs 158–159); larger: length of chela 1.15–1.27 (3) 1.24–1.35 (Ƥ) mm, length of movable chelal finger 0.68–0.77 (3) 0.73–0.81 (Ƥ) mm; pedipalps more slender: chela 7.05–7.9 (3) 6.7–7.5 (Ƥ) times as long as deep; ratio of pedipalpal femur/ carapace 1.7–2.0 (Ƥ); fixed and movable chelal fingers respectively with 14–17 and 8–11 triangular teeth, fixed finger at level of est-it with 3–4 teeth occupying 0.1 mm (distance between successive apices 0.040–0.048 mm).. C. monguzzii n. sp.

31 Carapace with 2 preocular microsetae on each side; epistome absent; fixed chelal finger at level of est-it with 3 teeth occupying 0.1 mm ........................................................................................... 32

- Carapace without preocular microsetae; epistome present, even if only weakly developed (fig. 146); fixed chelal finger at level of est-it with 4 teeth occupying 0.1 mm ...................................................... C. magrinii n. sp.

32 Larger: length of chela 1.07–1.19 (3) 1.11–1.27 (Ƥ) mm, length of movable chelal finger 0.64–0.70 (3) 0.65–0.74 (Ƥ) mm; pedipalps more slender: chela 7.2–8.4 (3) 6.6–7.9 (Ƥ); ratio of pedipalpal femur/carapace 1.7–1.9 (Ƥ).............. 33

- Smaller: length of chela 0.81 (3) 0.83–0.90 (Ƥ) mm, length of movable chelal finger 0.48 (3) 0.48–0.53 (Ƥ) mm; pedipalps less slender: chela 6.75 (3) 5.3–6.4 (Ƥ) times as long as deep; ratio of pedipalpal femur/carapace 1.4–1.5 (Ƥ).................................................................................................. C. etruscus n. sp.

33 Smaller: length of chela 1.07–1.095 (3) 1.11–1.25 (Ƥ) mm, length of movable chelal finger 0.64–0.66 (3) 0.65–0.73 (Ƥ) mm; pedipalps less slender: chela 7.2–7.4 (3) 6.6–6.9 (Ƥ) times as long as deep; ratio of pedipalpal femur/carapace 1.7–1.8 (Ƥ); movable cheliceral finger with isolated subapical tooth (di) mostly proximal with respect to the spinneret C. sulphureus n. sp.

- Larger: length of chela 1.14–1.19 (3) 1.27 (Ƥ) mm, length of movable chelal finger 0.66–0.70 (3) 0.74 (Ƥ) mm; pedipalps more slender: chela 8.2–8.4 (3) 7.9 (Ƥ) times as long as deep; ratio of pedipalpal femur/carapace 1.8–1.9 (Ƥ); movable cheliceral finger with isolated subapical tooth (di) at level with spinneret................................ C. latellai n. sp.

34 From Corsica...................................................................................... 35

- From Northern Italy................................................................................. 37

35 Epigean species, with eyes, of small or medium size: length of chela 0.47–0.67 (3) 0.53–0.61 (Ƥ) mm, length of movable chelal finger 0.28–0.39 (3) 0.30–0.35 (Ƥ) mm; pedipalps less slender: ratio of pedipalpal femur/carapace 1.1–1.4 (Ƥ); fixed chelal finger with 10–12 triangular teeth, at level of est-it with 4–6 teeth occupying 0.1 mm (distance between successive apices 0.019–0.026 mm)................................................................................ 36

- Endogean microphthalmic species of large size: length of chela 1.03–1.21 (3) 1.14–1.28 (Ƥ) mm, length of movable chelal finger 0.61–0.70 (3) 0.65–0.735 (Ƥ) mm; pedipalps more slender: ratio of pedipalpal femur/carapace 1.7–1.75 (Ƥ); fixed chelal finger with 13–15 triangular teeth, at level of est-it with 2–3 teeth occupying 0.1 mm (distance between successive apices 0.038–0.043 mm).......................................................... [ C. siscoensis Heurtault, 1975 ]

36 Larger: length of chela 0.655–0.67 (3) mm, length of movable chelal finger 0.38–0.39 (3) mm; pedipalps more slender: chela 5.5–5.8 (3) times as long as deep; ratio of pedipalpal femur/carapace 1.35–1.4 (3); fixed finger, at level of est-it, with 4 teeth occupying 0.1 mm (distance between successive apices 0.026 mm)........................ [ C. corsicus Callaini, 1981 ]

- Smaller: length of chela 0.47–0.54 (3) 0.53–0.61 (Ƥ) mm, length of movable chelal finger 0.28–0.31 (3) 0.30–0.35 (Ƥ) mm; pedipalps less slender: chela 4.8–5.15 (3) 4.1–4.3 (Ƥ) times as long as deep; ratio of pedipalpal femur/carapace 1.1–1.2 (Ƥ); fixed finger, at level of est-it, with 5–6 teeth occupying 0.1 mm (distance between successive apices 0.019–0.021 mm)..................................................................................... [ C. giustii Callaini, 1981 ]

37 Smaller: length of chela 0.51–0.63 (3) 0.56–0.69 (Ƥ) mm, length of movable chelal finger 0.30–0.38 (3) 0.32–0.41 (Ƥ) mm; fixed finger, at level of est-it, with 5 teeth occupying 0.1 mm (distance between successive apices 0.020–0.025 mm); species from Ligurian pre-Alps.............................................................................. 38

- Larger: length of chela 0.63–1.03 (3) 0.69–1.22 (Ƥ) mm, length of movable chelal finger 0.35–0.56 (3) 0.37–0.62 (Ƥ) mm; fixed finger, at level of est-it, with 3–4 teeth occupying 0.1 mm (distance between successive apices 0.032–0.035 mm); species from Eastern Alps................................................................... C. boldorii Beier, 1934

38 Endogean anophthalmic species; movable cheliceral finger with spinneret in both sexes (figs 80-81); pedipalps more slender: ratio of movable chelal finger/hand 1.45–1.55 (Ƥ), chela 5.6–5.5.7 (3) 5.5–5.6 (Ƥ) times as long as deep, ratio of pedipalpal femur/carapace 1.3 (Ƥ).................................................................... C. gallii n. sp.

- Epigean microphthalmic species; movable cheliceral finger mostly without spinneret in males (figs 133-134); pedipalps less slender: ratio of movable chelal finger/hand 1.3–1.4 (Ƥ), chela 5.1–5.3 (3) 4.6–4.75 (Ƥ) times as long as deep, ratio of pedipalpal femur/carapace 1.1–1.25 (Ƥ).......................................................... C. ligur n. sp.