Anthophora (Micranthophora) maculifrons Cresson, 1879

Orr, Michael C., Pitts, James P. & Griswold, Terry, 2018, Revision of the bee group Anthophora (Micranthophora) (Hymenoptera: Apidae), with notes on potential conservation concerns and a molecular phylogeny of the genus, Zootaxa 4511 (1), pp. 1-193 : 79-81

publication ID	https://doi.org/ 10.11646/zootaxa.4511.1.1
publication LSID	lsid:zoobank.org:pub:754B8A42-E269-42B5-92EB-043F3BEAA055
DOI	https://doi.org/10.5281/zenodo.3798724
persistent identifier	https://treatment.plazi.org/id/03DE87C1-D860-D924-EDE4-4785E5E094A7
treatment provided by	Plazi
scientific name	Anthophora (Micranthophora) maculifrons Cresson, 1879
status

Treatment

Anthophora (Micranthophora) maculifrons Cresson, 1879 View in CoL

( Figs. 38B View FIGURE 38 , 41B View FIGURE 41 , 46B View FIGURE 46 , 51B View FIGURE 51 , 56B View FIGURE 56 , 61B View FIGURE 61 , 66B View FIGURE 66 , 71B View FIGURE 71 , 76B View FIGURE 76 )

Anthophora maculifrons Cresson, 1879: 210 View in CoL (lectotype ( Cresson, 1916): ♀; Nevada, USA; coll. Morrison; ANSP, 2274).

Diagnosis. Males may be determined by the following character combination: galea strongly tessellate, dull; clypeus typically with inverted-T maculation, although upper half frequently weak or absent ( Fig. 51B View FIGURE 51 ); supraclypeus with transverse, often somewhat amorphous, maculation; T2–T6 with well-defined apical appressed setal bands; T7 with slight medial longitudinal carina, lateral projections with tips relatively blunt and only slightly recurved posteriorly, medial projections long with tips rounded ( Fig. 61B View FIGURE 61 ); S5 with dense, light erect setal patches ( Fig. 38B View FIGURE 38 ); and S6 with transparent apical rim medially ( Fig. 38B View FIGURE 38 ). Females may be determined with the following character combination: galea strongly tessellate, dull; clypeus typically with inverted-T maculation, although upper half may be weak or absent ( Fig. 41B View FIGURE 41 ); supraclypeus with strong transverse maculation, filling most of area; terga shiny between punctures; T2–T4 typically with well-defined apical appressed setal bands, though some species with wider bands have almost no basal black zone; and anal fimbria dark brown, often partially obscured by light setae dorsally.

Male description. Head: Facial maculations usually pale yellow, uncommonly richly yellow. Galea roughly reaching foretrochanter in repose, barely exceeding its front margin if at all; medium to dark brown; strongly tessellate, with weak reflections if any. Mandible with inferior tooth unusually expanded and broadly ovular, running about parallel to main blade; maculated from base to start or middle of inferior tooth ( Fig. 51B View FIGURE 51 ). Labral rim tridentate. Clypeal maculation varying from band to inverted-T, very nearly reaching lateral sides, filling about half of clypeal height if not inverted-T ( Fig. 51B View FIGURE 51 ); punctation moderately to weakly sparser within maculation than without. Paraocular area immaculate. Supraclypeal area maculated; usually transverse, sometimes amorphous. Scape maculated; fully along length and most of width below. Mesosoma: Scutum shiny; with impunctate areas, sometimes small or obscure due to invasion by sparse punctures. Tegula translucent to weakly transparent, medium to light brown. Basitibial plate weak but present; weakly outlined, often obscured by setae. Metasoma: T1 covered in appressed setae; T2–T6 with thinner apical appressed setal bands, not or very weakly undulate. Terga moderately to weakly tessellate between punctures, with relatively weak reflections; apparent through appressed setae only where sparse. Tergal rims transparent, slightly tan to clear. Male T7 ( Fig. 61B View FIGURE 61 ) with very weak medial longitudinal carina, sometimes only distinct near base of carina; lateral projections somewhat recurved, wide flanges; medial projections long and apically rounded, sides of each parallel. Male S5 ( Fig. 38B View FIGURE 38 ) with dense and light erect setal patches. Male S6 ( Fig. 38B View FIGURE 38 ) medially emarginate and slightly filled by transparent integument from which long setae arise. Male S7 ( Fig. 66B View FIGURE 66 ) with setae fully banding S7 around midpoint of length; widening from midpoint to tip, apical half widest clearly before rounded lateral tips; apical half vaguely ovular, rim rounded outward, with slight medial bump. Male S8 ( Fig. 71B View FIGURE 71 ) lateral projections distinct; medial projection nearly parallel-sided, apically emarginate. Male genital capsule ( Fig. 76B View FIGURE 76 ) with outer corners, where gonocoxite tips curve inward, marked by obvious flange of about 90 degrees or greater, best seen in profile; from corner to apex, in profile, gonocoxite with low secondary bump but relatively unmodified overall; tip, in profile, pointed and abruptly bent ventrally; gonostylus tip position slightly exceeding that of gonocoxite, measured from above along primary axis of latter. Pubescence: See Fig. 56B View FIGURE 56 . Appears medium to dark gray overall, often somewhat brownish. Setae generally somewhat tan dorsally except on metasoma, whiter there and elsewhere toward venter, except as follows: leg inner faces orange; tergal basal zones dark.

Female similar to male, except: Head: Mandible with inferior tooth ridge-like but expanded, slightly angled off main blade ( Fig. 41B View FIGURE 41 ). Labral rim rounded; fully maculated. Clypeal maculation similarly variable but typically stronger ( Fig. 41B View FIGURE 41 ). Scape immaculate. Mesosoma: Midtibial spur apically curved; tan to orange. Basitibial plate circular distally, teardropped overall. Metasoma: T1 covered in appressed setae; T2–T4 with appressed setal bands of variable thickness, sometimes only absent in very narrow basal zones, usually undulate; T5 with extensive appressed setae. T4 impunctate rim of short to almost moderate length medially, distinct but weak. Anal fimbria triangular or vaguely so, filling around third to under half of T5 length medially. Pubescence: See Fig. 46B View FIGURE 46 . Hindbasitarsal brush dark brown; anal fimbria usually orangish, sometimes invaded posteriorly by brown; S5 with dark apicomedially, hidden by light.

Distribution. See Fig. 19 View FIGURE 19 . Present in: USA: California, Utah, Idaho, Oregon, Nevada, Wyoming, Colorado, Arizona, likely Montana, possibly Washington; Mexico: possibly Baja California Norte. Found throughout much of west, montane in hotter areas, occupying broader range of environments in colder areas. Ecoregional habitance is as follows, based on 352 specimen records: 16 ecoregions total: Mojave Desert (116=33%), Great Basin shrub steppe (65=18.5%), Colorado Plateau shrublands (41=11.6%), Eastern Cascades forests (27=7.7%), Sierra Nevada forests (26=7.4%), Wyoming Basin shrub steppe (23=6.5%), Snake-Columbia shrub steppe (17=4.8%), Wasatch and Uinta montane forests (12=3.4%), Colorado Rockies forests (8=2.3%), California montane chaparral and woodlands (5=1.4%), Blue Mountains forests (5=1.4%), Arizona Mountains forests (2≤1%), Central and Southern Cascades forests (2≤1%), Sonoran Desert (1≤1%), California coastal sage and chaparral (1≤1%), Klamath- Siskiyou forests (1≤1%).

Phenology. Recorded from April–October, almost exclusively late summer to fall, most common in August– September. Likely univoltine, possibly emerging opportunistically on rare occasions.

Nesting biology. Unknown.

Floral specialization. Reported as a specialist on Chrysothamnus (Asteraceae) ( Moldenke & Neff 1974). Appears mostly true; known associations suggest specialization on Chrysothamnus and Ericameria as well as other fall flowering shrubs in Asteraceae . Primary recorded visits by 131 females include: Family Asteraceae (98%); Genus Ericameria (38%), Chrysothamnus (35%), Gutierrezia (23%); Species Ericameria nauseosa (34%). 190 total floral records from three families include: Asteraceae (58 ♂ 128 ♀): Aster sp. 1 ♀, Centaurea sp. 1 ♂, Chrysothamnus nauseosus 1 ♀, Chrysothamnus sp. 14 ♂ 41 ♀, Chrysothamnus viscidiflorus 5 ♂ 3 ♀, Ericameria nauseosa 25 ♂ 48 ♀, Ericameria sp. 6 ♂ 2 ♀, Grindelia squarrosa 1 ♂, Gutierrezia microcephala 12 ♀, Gutierrezia sarothrae 1 ♂ 11 ♀, Gutierrezia sp. 1 ♂ 8 ♀, Helianthus sp. 1 ♂, Machaeranthera canescens 1 ♀, Machaeranthera pinnatifida 1 ♂, Vanclevea stylosa 2 ♂, Orobanchaceae (1 ♀): Cordylanthus sp. 1 ♀, Polygonaceae (1 ♂ 2 ♀): Eriogonum effusum 1 ♀, Eriogonum nummulare 1 ♀, Eriogonum trichopes 1 ♂.

Specimens examined: USA: ARIZONA: Maricopa County: Usery Mountains, near Mesa: 1 ♂, 24 Apr 2008, J. Alcock; Mohave County: Hualapai Mountains: 1 ♀, 12 Sep 1994, T.L. Griswold; Yavapai County: 2 mi SW of Prescott: 1 ♂, 7 Sep 1964, C.W. O'Brien, L.B. O'Brien; Kingman, 55 mi E: 1 ♂, 2 Sep 2012, K. Wright; Prescott: 1 ♀, 19 Sep 1953, P.H. Timberlake; CALIFORNIA: Alpine County: Woodfords: 1 ♀, 25 Oct 1959, W.E. Simonds; El Dorado County: Lake Tahoe: 1 ♂, 11 Aug 1940, E.E. Kenaga; Stateline: 2 ♂, 20 Sep 1938, F.L. Blanc; Inyo County: 12 mi SW of Little Lake: 1 ♂ 3 ♀, 16 Sep 1969, M.S. Wasbauer; Bishop: 1 ♂, 2 Aug 1936, G.E. & R.M. Bohart; Cerro Gordo Mine: 1 ♀, 29 Aug 1958, A.S. Menke, L.A. Stange; Coso Bridge, 1 mi SW Coso Peak: 1 ♀, 14 Sep 1996, G. Pratt, C. Pierce; Independence: 1 ♀, 26 Oct 1963, J.D. Birchim; Independence, 7 km E: 2 ♀, 17 Sep 1993, T.L. Griswold; Jackass Spring, Panamint Range: 4 ♂, 31 Aug–1 Sep 1991, collector unknown; 18 ♂ 15 ♀, 6–8 Sep 1994, R.R. Snelling; Little Lake: 1 ♂, 3 Jun 1937, W.C. Bush; Manzanar, 3 km E: 1 ♀, 17 Sep 1993, T.L. Griswold; Owens River, 7 km SE Lone Pine: 1 ♂, 19 Sep 1993, T.L. Griswold; Owens Valley: 1 ♂, 2 Aug 1932, G.E. Bohart, R.M. Bohart; Pinon Bridge, Coso Mountains: 1 ♂, 13 Aug 1995, G.R. Ballmer; Tinemaha Reservoir: 1 ♀, 13 Oct 1974, T.L. Griswold; White Mountains, Wyman Canyon: 1 ♀, 13 Aug 1999, A.S. Menke, N.D. Menke; Kern County: Tehachapi, 10 mi W: 1 ♀, 17 Oct 1958, G.M. Ferguson; Valley Vista Camp [may be Chula Vista Camp]: 1 ♂, 9 Sep 1971, collector unknown; Walker Pass: 1 ♂, 22 Sep 1957, H.R. Moffit; 1 ♂, 22 Sep 1957, J.C. Hall; 1 ♀, 26 Sep 1957, E.I. Schlinger; 1 ♂ 1 ♀, 26 Sep 1957, J. Hall; 1 ♂, 26 Sep 1957, J.C. Hall; 1 ♀, 27 Sep 1957, H.R. Moffit; 1 ♂ 9 ♀, 8 Oct 1957, P.H. Timberlake; 1 ♂, 23 Sep 1981, E.M. Fisher; Lassen County: 9 mi W of Termo: 1 ♀, 12 Aug 1985, T.R. Haig; Bridge Creek Camp: 1 ♂ 1 ♀, 14 Aug 1952, R.C. Bechtel; Litchfield: 1 ♀, 8 Sep 1957, E.G. Linsley; Standish: 1 ♀, 27 Aug 1960, collector unknown; Los Angeles County: Claremont: 1 ♀, date unknown, Baker; Modoc County: Cedar Pass: 1 ♀, 11 Oct 1952, E.I. Schlinger; 1 ♀, 3 Sep 1961, W.E. Simonds; Fandango Pass: 1 ♀, 16 Jul 1991, T.R. Haig; Lake City: 1 ♂, 2 Aug 1922, C.L. Fox; Mono County: Banner Springs: 1 ♀, 8 Aug 2014, T.L. Griswold; Grants Lake: 1 ♂, 5 Aug 1948, P.D. Hurd, J.W. MacSwain; Mammoth: 1 ♂, 5 Aug 1936, G.E. & R.M. Bohart; 1 ♀, 7 Aug 1936, G.E. Bohart, R.M. Bohart; Sherwin Summit, Hwy 395: 1 ♀, 8 Sep 1975, R.R. Snelling; Nevada County : Boca Dam, 11 E Truckee: 1 ♀, 1 Sep 1957, J.M. Linsley; Hobart Mills, 1 m S: 1 ♀, 1 Sep 1957, J.M. Linsley; Hobart Mills, 7 m N: 1 ♂, 26 Aug 1948, E.G. Linsley; 1 ♂, 26 Aug 1948, M.A. Cazier; 2 ♀, 26 Aug 1948, R.E. Smith; San Bernardino County: Barton Flats: 1 ♂, 10 Sep 1946, N. Crickmer; 1 ♂, 3 Oct 1946, N. Crickmer; Cedar Canyon: 1 ♀, 3 Oct 1980, T.L. Griswold; Cedar Wash: 1 ♂, 1 Sep 1980, T.L. Griswold; Fish Creek Meadow, San Gorgonio Wilderness: 1 ♂, 2 Sep 1980, K.W. Cooper; Hole-in-the-Wall: 2 ♀, 23 Sep 1993, T.L. Griswold; Kessler Spring: 2 ♀, 9 Sep 1994, T.L. Griswold; Lanfair Valley: 3 ♀, 23 Sep 1993, T.L. Griswold; 1 ♀ 1 ♀, 19 Sep 1994, T.L. Griswold; Macedonia Canyon: 1 ♀, 23 Sep 1993, T.L. Griswold; Mid Hills, SSE of Cima: 1 ♂, 14 Sep 1975, T.L. Griswold; New York Mountains: 7 ♀ 1 ♀, 23 Sep 1993, T.L. Griswold; Round Valley: 1 ♂ 3 ♀, 3 Oct 1980, T.L. Griswold; 5 ♀, 23 Sep 1993, T.L. Griswold; South Fork Campground, E of Barton Flats, San Bernardino Mountains, San Bernardino National Forest: 1 ♀, 10 Sep 1944, P.H. Timberlake; South Fork Santa Ana River at Hwy 38, San Bernardino Mountains, San Bernardino National Forest: 1 ♂, 22 Aug 1952, P.H. Timberlake; Teutonia Peak: 9 ♀, 9 Sep 1994, T.L. Griswold; Watson Wash: 1 ♂ 1 ♀, 19 Sep 1994, T.L. Griswold; 2 ♀, 20 Sep 1994, T.L. Griswold; Wild Horse Canyon: 8 ♀ 1 ♀, 23 Sep 1993, T.L. Griswold; Shasta County: Old Station: 1 ♂ 1 ♀, 16 Aug 1952, R.C. Bechtel; Sierra County: Sierraville: 1 ♂ 1 ♂, 20 Aug 1953, E.I. Schlinger; Yuba Pass: 1 ♂, 20 Aug year unknown, A.A. Grigarick; Siskiyou County: 4 mi NE of Bray: 1 ♀, 24 Aug 1979, T.R. Haig; Gazelle: 1 ♀, 11 Oct 1950, M.F. McClay; Lava Beds National Monument: 1 ♀, 22 Aug 1954, G. Ferguson; Macdoel: 4 ♂ 3 ♀, 21 Aug 1954, G. Ferguson; 1 ♂, 19 Aug 1963, J. Schuh; Tuolumne County: Sonora Pass: 1 ♂, 16 Aug 1960, R.R. Snelling; COLORADO: Jackson County: Walden, 7.3 km N; Jc Rd 10: 2 ♂, 2 Aug 2014, M.C. Orr; Moffat County: Craig, 28 mi W: 1 ♀, 21 Aug 1966, R.&M. Roberts; Maybell, 8 mi E: 3 ♀, 17 Aug 1959, E. Ordway; IDAHO: Adams County: Oxbow Dam: 1 ♂, 8 Aug 1976, A.R. Moldenke; Bear Lake County: Paris, 3 mi W: 1 ♀, 18 Jul 1966, A.R. Gittins; Bingham County: Sterling: 1 ♂, 24 Sep 1954, A.R. Gittins; Cassia County : Elba-Basin Pass: 1 ♀, 23 Aug 1958, W.F. Barr; Fremont County: Saint Anthony Sand Dunes: 1 ♂ 1 ♀, 9 Aug 2013, A. Payne; Jefferson County: Terreton: 1 ♂, 29 Jul 1958, A.R. Gittins; Minidoka County: Acequia: 1 ♂, 9 Jul 1969, W.F. Barr; None County: Rock Creek Ranger Station, Sawtooth Nat Forest: 1 ♂, 6 Aug 1958, A.R. Gittins; Owyhee County: Riddle, 5 mi N: 1 ♀, 18 Aug 1971, D.E. Foster; NEVADA: Douglas County : Daggett Pass, Toiyabe National Forest: 1 ♂, 18 Aug 1951, R.C. Bechtel; Genoa, 3 mi S: 1 ♂, 23 Aug 1955, G.E. Bohart; Minden: 1 ♂, 24 Aug 1952, R.C. Bechtel; 1 ♂, 24 Aug 1952, R.M. Bohart; Spooner Summit: 1 ♀, 12 Sep 1956, T.R. Haig; Elko County: Elko, 27 mi E: 1 ♂, 20 Aug 1955, W.P. Stephen; Texas Spring: 1 ♀, 14 Sep 1991, W.P. Nye; Eureka County: Eureka: 1 ♀, 27 Sep 1938, I. McCracken; Humboldt County: Summer Place, 7 mi SE Summit Lk Ind Res: 1 ♀, 6 Sep 1997, F.D. Parker; Lander County: Campbell Creek Ranch: 1 ♂, May 1981, J.B. Knight; Washoe County: Little Valley: 1 ♂, 3 Sep 1981, R.W. Rust; Sparks, 17 m N: 3 ♀, 2 Sep 1957, E.G. Linsley; Wingfield Springs, N Sparks: 1 ♀, 3 Sep 1997, F.D. Parker; OREGON: Crook County: Bend, 82 mi E: 1 ♀, 20 Aug 1948, A.B. Black; Deschutes County: Bend, 5 mi S: 1 ♀, 9 Aug 1939, Schuh, Gray; Sisters: 1 ♀, 10 Aug 1926, H.A. Scullen; Three Sisters: 1 ♀, 10 Aug 1926, H.A. Scullen; Tumalo: 1 ♂, 7 Aug 1939, Gray, Schuh; 2 ♀, 7 Aug 1939, Schuh, Gray; 1 ♀, 7 Aug 1939, collector unknown; Grant County: Antelope Mountain: 1 ♀, 13 Aug 1931, D.K. Frewing; 1 ♂, 17 Aug 1931, D.K. Frewing; Harney County: Antelope Mountain: 1 ♂, 13 Aug 1931, D.K. Frewing; 1 ♂, 14 Aug 1931, D.K. Frewing; Klamath County: Bly Mountain: 1 ♂ 1 ♀, 31 Aug 1958, J. Schuh; Bly, 6 mi W: 1 ♀, 18 Aug 1955, W.P. Stephen; Grizzly Butte: 1 ♂ 1 ♀, 18 Aug 1939, Gray, Schuh; 3 ♀, 18 Aug 1939, K. Gray, Schuh; 2 ♀, 18 Aug 1939, Schuh, Gray; Klamath Falls Experiment Station: 1 ♂, 3 Aug 1960, P.F. Torchio; Lake of the Woods: 1 ♂, 6 Aug 1959, P.F. Torchio; Lake County: La Pine, 24 mi SE, T24S R13E S24: 2 ♀, 9 Sep 1957, G.F. Kraft; UTAH:Box Elder County: Snowville: 1 ♂, 1 Sep 1970, G.E. Knowlton, J.H. Judd; Yost: 1 ♀, 27 Aug 1982, W.J. Hanson, C.R. Nelson; Cache County: Logan Canyon: 1 ♂, 9 Aug 1940, W.P. Nye; Emery County: Goblin Valley: 1 ♀, 14 Sep 1979, W.P. Nye; Goblin Valley State Reserve, 2 mi N: 1 ♂, 25 Aug 1980, A.&K. Menke, F.D. Parker; Goblin Valley, wash: 1 ♂, 16 Sep 1979, F.D. Parker, D.F. Veirs; Hatt Ranch, 17 mi W Green River: 2 ♂, 23–24 Aug 2001, M.E. Irwin, F.D. Parker; 4 ♂, 24–25 Aug 2001, M.E. Irwin, F.D. Parker; Little Flat Top: 1 ♂, 22–26 Aug 2001, F.D. Parker, C. Lambkin, M. Metz, M. Hauser, M.E. Irwin; Little Flat Top, 8 km E: 2 ♂, 22–26 Aug 2001, M.E. Irwin, F.D. Parker, M. Metz, M. Hauser, C. Lambkin; Little Gilson Butte, 0.5 mi E: 1 ♂, 27 Aug 1985, T.L. Griswold; Little Gilson Butte, 2 air mi E: 1 ♂, 25 Aug 1981, D.F. Veirs, T.L. Griswold, F.D. Parker; Little Gilson Butte, 2 air mi W: 1 ♂, 16 Sep 1980, D.F. Veirs; Little Gilson Butte, 3.2 air mi NE: 1 ♂, 13 Sep 1983, F.D. Parker, T.L. Griswold; Little Gilson Butte, 3.2 mi NE: 1 ♂, 13 Sep 1983, F.D. Parker, T.L. Griswold; North Temple Wash: 1 ♀, 17 Oct 1988, T.L. Griswold; Garfield County: Blubber Creek, jct Right Fork: 1 ♀, 20 Jul 2000, B.S. Heath; Boulder: 1 ♀, 11 Sep 1949, C.J. Stewart; Panguitch Lake: 1 ♂, 17 Aug 1995, V.J. Tepedino, F.D. Parker; Ruby's Inn, NE: 1 ♂ 3 ♂, 17 Aug 1995, V.J. Tepedino, F.D. Parker; Juab County: Little Sahara Sand Dunes: 4 ♂ 3 ♀, 14 Sep 1993, T.L. Griswold; Tooele County: 0.5 mi E Wig Mountain: 1 ♀, 16 Sep 2003, O.J. Messinger; 0.8 mi NNE Little Granite Mt.: 1 ♂ 2 ♀, 23 Sep 2003, H. Ikerd; 1.3 mi W Little Granite Mt.: 3 ♀, 23 Sep 2003, T.L. Griswold; 1.9 mi N Little Granite Mtn.: 3 ♀, 23 Sep 2003, T.L. Griswold; 4 ♀, 2 Oct 2003, T.L. Griswold; 2 mi N Little Granite Mt.: 1 ♀, 29 Sep 2005, T.L. Griswold; 2.6 mi WNW Simpson Butte: 1 ♀, 16 Sep 2003, H. Ikerd; 3 ♀ 1 ♀, 16 Sep 2003, O.J. Messinger; 2.8 mi W Simpson Buttes: 1 ♂ 1 ♂, 28 Sep 2005, K.T. Huntzinger; 3.2 mi SSE Wig Mt.: 1 ♀, 16 Sep 2003, O.J. Messinger; 1 ♀, 2 Oct 2003, O.J. Messinger; 1 ♀, 2 Oct 2003, T.L. Griswold; 4.17 mi SE Wig Mt.: 1 ♂ 1 ♀, 13 Sep 2005, K.T. Huntzinger; 1 ♀, 13 Sep 2005, O.J. Messinger, K.T. Huntzinger; 3 ♀, 26 Sep 2005, K.T. Huntzinger; 1 ♀ 1 ♀, 26 Sep 2005, T.L. Griswold; Washington County: Mountain Meadow Monument: 1 ♂ 4 ♀, 3 Sep 1962, C.W. O'Brien; Wayne County: Carcass Creek, SSE of Torrey: 1 ♂ 3 ♀, 5 Sep 2008, T.L. Griswold; Hanksville, 9 mi NE: 1 ♂, 28 Aug 1985, T.L. Griswold; Teasdale, 4 mi SE: 2 ♂ 4 ♂, 15 Aug 1995, V.J. Tepedino, F.D. Parker; WYOMING: Albany County: Dirt Farm, 8 mi SE Laramie: 1 ♀, 23 Aug 1974, V.J. Tepedino; 1 ♂, 19 Aug 1975, V.J. Tepedino; 1 ♀, 10 Sep 1975, V.J. Tepedino; 1 ♂, 9 Aug 1976, V.J. Tepedino; 1 ♂ 1 ♀, 16 Aug 1976, V.J. Tepedino; 1 ♀ 1 ♀, 24 Aug 1976, V.J. Tepedino; Fremont County: Shoshoni: 1 ♀, 17 Aug 1965, G.E. Bohart; Sublette County: Pinedale: 1 ♂, 11 Aug 1962, J.G. Rozen, B.L. Rozen; Sweetwater County: Eden, 24km ESE; Killpecker Dunes: 1 ♂, 24 Jul 2012, M.C. Orr; Eden, 25km ESE; nr Killpecker Dunes: 2 ♂, 24 Jul 2012, M.C. Orr; Eden, 25km SE; Chilton Rd: 2 ♂ 1 ♀, 23 Jul 2012, M.C. Orr; Eden, 6km SSE; US Hwy 191: 1 ♂, 24 Jul 2012, M.C. Orr; Farson, 20 mi E: 1 ♂ 1 ♀, 4 Aug 1955, G.E. Bohart; Granger: 1 ♀, 25 Aug 1954, G.E. Bohart; Red Desert: 3 ♀, 6 Sep 1979, F.D. Parker; Uinta County: Lyman, 11km SSE; WY HWY 414: 1 ♂, 22 Jul 2012, M.C. Orr.