Anthophora (Micranthophora) albata Cresson, 1876

Orr, Michael C., Pitts, James P. & Griswold, Terry, 2018, Revision of the bee group Anthophora (Micranthophora) (Hymenoptera: Apidae), with notes on potential conservation concerns and a molecular phylogeny of the genus, Zootaxa 4511 (1), pp. 1-193 : 23-26

Anthophora (Micranthophora) albata Cresson, 1876

( Figs. 34A , 39B , 44B , 49B , 54B , 59B , 64B , 69B , 74B )

Anthaphora albata Cresson, 1876: 211 (lectotype ( Cresson, 1916): ♀; Denver, Colorado, USA; coll. Putnam, Jun.; ANSP,

2273). Anthophora albata Cresson, 1879: 201–211 (implied correction). Anthophora flexipes Cresson, 1879: 211 (lectotype ( Cresson, 1916): ♀; Nevada, USA; coll. Morrison; ANSP, 2275). NEW

SYNONYM. Anthophora arthuri Cockerell, 1906: 72 (holotype: ♀; Maybell, Colorado, USA; coll. S. Johnson, 1 Aug. 1904, at Asteraceae

(yellow); CASENT, type 15465). NEW SYNONYM.

Diagnosis. This species is unusually distinctive. Males are immediately identifiable by any of the following: forebasitarsus spindly and with black and white tufts, mid leg tarsal segments all elongated, tibial spurs apically pointed and inner hind spur medially bent, and hindbasitarsus irregularly enlarged and vaguely triangular (most visible in Figs. 49B , 54B ). Females are immediately identifiable by the opaque dark brown, apically pointed tibial spurs on the mid and hindlegs. Females may be otherwise determined by the following character combination: galea tessellate, dull; mandible with inferior tooth large, vaguely triangular, clearly angled off main blade ( Fig. 39B ); clypeal maculation an inverted-T, uncommonly interrupted above ( Fig. 39B ); supraclypeus strongly maculated, either trapezoidal or a widened pentagon; scutum shiny, with impunctate areas; and S5 with dark setae apically.

Male description. Head: Facial maculations richly to pale yellow. Labral rim tridentate. Galea reaching rear of foretrochanter or nearly so in repose; medium to dark brown; usually strongly but sometimes moderately tessellate, with weak reflections. Mandible with inferior tooth large and triangular, clearly angled off main blade; maculated from base to near start of inferior tooth ( Fig. 49B ). Labral rim rounded; fully maculated. Clypeal maculation an inverted-T with slight depression to each side of medial upward-projecting line, reaching lateral sides, almost always reaching top medially ( Fig. 49B ); punctation only slightly sparser within maculation, near rim, than without. Paraocular area immaculate. Supraclypeal area strongly maculated; usually entirely filled. Scape maculated; fully along length and width below. Mesosoma: Scutum shiny; with distinct impunctate areas. Tegula transparent to slightly translucent, yellowish to light brown. Foreleg femur and tarsi elongate, basitarsus sinuate and spindly; basitarsus with unusual sculpted inner face and intricate setal tufts of white and black ( Figs. 49B , 54B ). Midleg more elongate: femur roughly equally so, basitarsus and especially other tarsi proportionally longer compared to foreleg ( Fig. 54B ). Hindleg only slightly elongate, mostly in tibia and basitarsus; femur, tibia, and basitarsus swollen, latter two seemingly forming clasping mechanism in conjunction ( Fig. 54B ); with unusually thick and long setae on inner faces. Basitibial plate absent; slight bump may be present anteriorly. Metasoma: T1 usually covered in appressed setae, sometimes with apical appressed setal band; T2–T4 with apical appressed setal bands, variable in thickness and often undulate; T5–T6 with or without apical appressed setal bands. Terga weakly to moderately tessellate between punctures, usually still with moderate reflections; sometimes still apparent through appressed setae. Tergal rims transparent, clear to slightly tan. Male T7 ( Fig. 59B ) without medial longitudinal carina; lateral projections only weakly recurved, blunt flanges; medial projections long and knife-like, the space between them usually filled by setae. Male S5 resembling Fig. 38D , with dense and dark erect setal patches, but denser medially. Male S6 resembling Fig. 38B , clearly medially emarginate with slight transparent integument near base, but with more transparent integument anteriorly and less setae covering rim. Male S7 ( Fig. 64B ) with setae restricted to sides around midpoint of length; widening from midpoint to tip, apical half widest near rounded lateral tips; apical half vaguely heart-shaped, rim rounded outward, with narrow medial emargination. Male S8 ( Fig. 69B ) lateral projections absent; medial projection narrowing to tip, where rounded at narrow medial point. Male genital capsule ( Fig. 74B ) with outer corners, where gonocoxite tips curve inward, nearly absent small bump, best seen in profile; from corner to apex, in profile, gonocoxite with low bump somewhat hidden to interior; tip, in profile, rounded and weakly curved ventrally; gonostylus tip position significantly exceeding that of gonocoxite, measured from above along primary axis of latter. Pubescence: See Fig. 54B . Appears light to medium gray or overall, scutum sometimes tan. Setae white to off-white, though sometimes tan dorsally, except as follows: vertex and scutum only very rarely with any dark intermixing; legs highly variable, but tibia nearly always white; leg inner faces dark brown to black; tergal basal zones dark when present; T5–T6 with or without appressed setae; S5 thick patches always dark brownish.

Female similar to male, except: Head: Facial maculations yellow to off-white, sometimes yellow-green. Labrum rim rounded; fully maculated. Clypeal maculation with upward-directed portion of inverted-T thinner ( Fig. 39B ). Scape immaculate. Mesosoma: Legs unmodified in length. Midtibial spur apically pointed; unusually opaque, brown. Basitibial plate circular distally, teardropped overall. Metasoma: T1 covered in appressed setae; T2–T4 usually with apical appressed setal bands undulate, but highly variable in width, sometimes nearly covering terga but sometimes reduced to half; T5 with or without appressed setae. T4 impunctate rim of short length medially, often only slightly visible. Anal fimbria strongly triangular, covering from nearly half to over half of T5 length medially. Pubescence: See Fig. 44B . Leg inner faces brown; hindbasitarsal brush dark brown to blackish; tergal bands usually thicker; anal fimbria dark brown to blackish or light and whitish east of Colorado’s Front Range; sterna light or dark.

Distribution. See Fig. 7 . Present in USA: Utah, California, Idaho, Colorado, Wyoming, Nevada, Oregon, New Mexico, Arizona, Washington, likely Montana; Mexico: Baja California Norte, possibly Sonora; possibly Canada: possibly British Columbia. Found throughout much of the West, montane in hotter areas, occupying broader range of environments in colder areas. Ecoregional habitance is as follows, based on 1325 specimen records: 16 ecoregions total: Colorado Plateau shrublands (745=56.2%), Great Basin shrub steppe (259=19.5%), Sierra Nevada Forests (143=10.8%), Snake-Columbia shrub steppe (79=6%), Wyoming Basin shrub steppe (38=2.9%), Great Basin montane forests (13=1%), Colorado Rockies forests (12≤≤1%), California montane chaparral and woodlands (9≤≤1%), Wasatch and Uinta montane forests (9≤≤1%), Mojave Desert (6≤≤1%), California coastal sage and chaparral (3≤≤1%), Eastern Cascades forests (3≤1%), Sierra Juarez and San Pedro Martir pine-oak forest (2≤≤1%), South Central Rockies forests (2≤≤1%), Klamath-Siskiyou forests (1≤≤1%), Western short grasslands (1≤≤1%).

Phenology. Recorded from May–October, heavily dependent on elevation and region, generally flies earlier in warmer regions. Univoltine at higher elevations, potentially bivoltine lower.

Nesting biology. Torchio (1968) described the nesting biology of A. albata . This species nests in both horizontal, unconsolidated sand and in vertical sandstone banks. The use of sandstone nests by A. albata has been confirmed by the preservation of specimens within nest cells, but whether or not this species excavated these nests is unclear. Although Torchio (1968) claims that this species was witnessed excavating the sandstone nests, he collected both A. albata and A. peritomae from the site, and the latter is confirmed as a minimally facultative excavator of sandstone nests ( Orr et al. 2016). Further, he marked many of the nest cells collected from that site as “flexipes or peritomae,” despite no mention of A. peritomae anywhere in his paper.

Floral specialization. Likely a specialist on Asteraceae , with great breadth, but possibly generalist with strong Asteraceae preference or, less likely, an eclectic specialist on Asteraceae and Cleomaceae ( Cane & Sipes 2006) . Pure Asteraceae loads were quite common among pollen views (11/15), remaining views also with non-negligible proportions of what was likely Cleomaceae pollen, two nearly purely so and two with <50%. Primary recorded visits by 279 females include: Family Asteraceae (61%), Cleomaceae (10%). 423 total floral records from 14 families include: Asteraceae (97 ♂ 169 ♀): Aster chilensis 1 ♂ 1 ♀, Aster falcatus 7 ♂, Aster sp. 2 ♂ 5 ♀, Asteraceae sp. 1 ♂, Asteraceae sp. 2 ♂, Asteraceae sp. 3 ♀, Chaenactis sp. 3 ♂ 12 ♀, Chrysothamnus depressus 1 ♀, Chrysothamnus linifolius 1 ♀, Chrysothamnus sp. 1 ♂ 16 ♀, Chrysothamnus viscidiflorus 1 ♀, Ericameria nauseosa 5 ♂ 6 ♀, Ericameria parryi 1 ♀, Erigeron bellidiastrum 1 ♂, Erigeron sp. 2 ♂ 2 ♀, Erigeron sp. 4 ♂, Grindelia sp. 2 ♀, Grindelia squarrosa 4 ♂ 6 ♀, Haplopappus nuttallii 1 ♀, Helianthus annuus 3 ♂ 7 ♀, Heterotheca villosa 27 ♂ 21 ♀, Hymenopappus filifolius 19 ♀, Machaeranthera canescens 6 ♂ 8 ♀, Machaeranthera tanacetifolia 6 ♂ 4 ♀, Malacothrix sonchoides 1 ♂ 7 ♀, Orochaenactis thysanocarpha 8 ♂ 11 ♀, Senecio douglasii 4 ♂ 5 ♀, Senecio sp. 1 ♂ 1 ♀, Senecio spartioides 1 ♂ 2 ♀, Stephanomeria exigua 5 ♀, Stephanomeria sp. 6 ♂ 12 ♀, Stephanomeria tenuifolia 1 ♂ 3 ♀, Symphyotrichum lanceolatum 5 ♀, Vanclevea stylosa 1 ♀, Boraginaceae (5 ♂ 10 ♀): Cryptantha sp. 8 ♀, Phacelia leucophylla 3 ♂, Phacelia rafaelensis 1 ♀, Phacelia sp. 2 ♂ 1 ♀, Brassicaceae (1 ♀): Thelypodium integrifolium 1 ♀, Cleomaceae (5 ♂ 28 ♀): Cleome lutea 4 ♂ 18 ♀, Cleome serrulata 1 ♂ 5 ♀, Cleome sp. 1 ♀, Cleomella sp. 4 ♀, Crassulaceae (1 ♀): Sedum lanceolatum 1 ♀, Fabaceae (8 ♂ 16 ♀): Lupinus sp. 1 ♀, Medicago sp. 1 ♂, Melilotus alba 1 ♀, Polemoniaceae sp. 6 ♂ 4 ♀, Psoralea lanceolata 1 ♂ 10 ♀, Lamiaceae (14 ♂ 17 ♀): Hedeoma drummondii 2 ♂, Monardella sp. 1 ♀, Poliomintha incana 11 ♂ 11 ♀, Poliomintha sp. 1 ♀, Salvia sp. 1 ♂ 4 ♀, Malvaceae (2 ♀): Sphaeralcea coccinea 2 ♀, Onagraceae (2 ♀): Oenothera sp. 2 ♀, Plantaginaceae (1 ♂ 1 ♀): Penstemon laricifolius 1 ♂, Penstemon strictus 1 ♀, Polemoniaceae (5 ♂ 18 ♀): Gilia congesta 2 ♀, Gilia leptomeria 1 ♂ 5 ♀, Gilia sp. 4 ♂ 11 ♀, Polygonaceae (6 ♂ 11 ♀): Eriogonum corymbosum 2 ♂ 4 ♀, Eriogonum leptocladon 2 ♂, Eriogonum nummulare 2 ♀, Eriogonum racemosum 1 ♂, Eriogonum sp. 5 ♀, Polygonum sp. 1 ♂, Tamaricaceae (1 ♂ 2 ♀): Tamarix chinensis 1 ♂ 2 ♀, Verbenaceae (3 ♂): Verbena bracteata 3 ♂.

Comments. Two names, A. arthuri and A. flexipes , are here synonymized with A. albata . The female type of A. arthuri is only notable for its slightly smaller size, lighter setal color in the anal fimbria, and the color of the tibial spurs ( Cockerell 1906). All three of these characters vary within some species of A. ( Micranthophora ). Cockerell (1906) himself stated “it may be that this is only a variety of albata ,” and the authors here agree and formally synonymize A. arthuri to the senior synonym A. albata . The status of A. flexipes is more problematic. The more western A. flexipes has been distinguished from the more eastern A. albata , in which the terga are covered in appressed setae, including T5, solely on the basis of the reduced amount of appressed pubescence: only apical bands on T1-T4 and none on T5. Numerous exceptions to this have been found in the Sierra Nevada, where some individuals have appressed setae on T5, and the size of the apical bands varies significantly. Further, COI evidence suggests they are the same species ( A. albata from Colorado versus A. flexipes from California have only 1.5% sequence differentiation). Consequently, A. flexipes is also formally synonymized with the senior A. albata .

Specimens examined: MEXICO: Baja California Norte: Sierra San Pedro Martir: 2 ♀, 13 Jun 1953, P.H. Arnaud, Jr.. USA: ARIZONA: Apache County: Sanders, 17.07 air km SbE: 2 ♀, 5 Jun 2012, T.L. Griswold; Coconino County: Cameron, 15 mi N: 1 ♀, 3 May 1972, G.E. Bohart, F.D. Parker, P.F. Torchio; Moenkopi, 4.5 mi E: 1 ♀, 10 Jun 1966, J.H. Davidson, M.A. Cazier; 3 ♂, 10 Jun 1966, J.M. Davidson, M.A. Cazier; Page, 10 mi S: 1 ♂, 3 May 1972, P.F. Torchio, G.E. Bohart, F.D. Parker; The Gap: 5 ♀, 25 Sep 1960, G.E. Bohart; The Gap, 15 mi N: 3 ♀, 21 Sep 1966, R.W. Rust, P. Torchio, G. Wood, N. Youssef; 1 ♀, 21 Sep 1966, R.W. Rust, P.F. Torchio, G. Wood, N. Youssef; Navajo County: Hotevilla: 1 ♂ 1 ♀, 3 Jun 1967, J.H. Davidson, J.M. Davidson, M.A. Cazier; 1 ♀, date unknown, collector unknown; Jadito Trading Post: 1 ♀, 28 Jun 1966, J.H. Davidson, J.M. Davidson, M.A. Cazier; CALIFORNIA: Alpine County: Ebbetts Pass: 1 ♀, 29 Aug 1957, E. Mexger; Hope Valley: 1 ♀, 18 Jul 1948, collector unknown; 1 ♀, 12 Jul 1966, R.L. Brumley; Woodfords: 1 ♀, 29 Sep 1963, W.E. Simonds; El Dorado County: Echo Lake: 2 ♂ 1 ♀, 23 Jul 1955, E.L. Schlinger; 1 ♂, 23 Jul 1955, J.C. Downey; Fresno County: Huntington Lake: 1 ♂ 1 ♀, 8 Jul 1919, E.P. Van Duzee; 1 ♂, 15 Jul 1919, F.E. Blaisdell; 1 ♂, 16 Jul 1919, F.C. Clark; 1 ♀, 26 Jul 1919, E.P. Van Duzee; Inyo County: Big Pine Creek: 1 ♀, 20 Jun 1942, R. Bohart; Big Pine, Glacier Lodge: 1 ♂, 23 Jun 1937, E.C. Van Dyke; Bishop: 3 ♂, 2 Aug 1936, G.E. & R.M. Bohart; 1 ♂, 22 Jun 1937, collector unknown; Bishop Creek: 1 ♀, 2 Aug 1936, G.E. Bohart, R.M. Bohart; 1 ♂, 2 Aug 1936, collector unknown; 1 ♀, 22 Jun 1937, C.D. Michener; 1 ♀, 22 Jun 1937, collector unknown; 1 ♀, 28 Jun 1974, T.L. Griswold; Bishop Creek, Bishop: 1 ♂ 1 ♀, 22 Jun 1937, collector unknown; Bishop Vil: 1 ♂ 2 ♀, 22 Jun 1937, collector unknown; Deep Springs, Deep Springs Valley, Inyo Mountains: 1 ♂, 7 Sep 1969, M.S. Wasbauer; Jackass Spring, Panamint Range: 1 ♂ 1 ♀, 6–8 Sep 1994, R.R. Snelling; Owens Valley: 1 ♀, 2 Aug 1936, G.E. Bohart, R.M. Bohart; Westgard Pass, summit: 1 ♀, 15 Jun 1937, C.D. Michener; White Mountains, Goat Spring, 1.06 Air mi NWbW: 1 ♀, 24 Jul 2014, T.L. Griswold; Whitney Portal, Lone Pine Canyon, Inyo National Forest: 3 ♂, 3 Jul 1953, H. Washburn; Wildrose Station, 4 mi E: 1 ♂, 25 Aug 1980, A.&K. Menke, F.D. Parker; Kern County: Walker Pass: 1 ♀, 26 Jun 1949, L.W. Isaak; 1 ♀, 13 Jun 1953, G.D. Butler; 1 ♀, 16 Sep 1960, R.P. Allen; Lassen County: Bridge Creek Camp: 1 ♀, 9 Jul 1949, L. Andres; Hallelujah Jct: 1 ♀, 27 Jun 1949, P.D. Hurd; 1 ♂, 4 Jul 1949, E.I. Schlinger; 1 ♂, 4 Jul 1949, J.E. Gillaspy; 1 ♂, 4 Jul 1949, P.D. Hurd; 1 ♀, 12 Jul 1954, G.A. Schaefers; 1 ♀, 12 Jul 1962, M.E. Irwin; 1 ♂, 2 Jul 1966, collector unknown; 23 ♂ 16 ♀, 27 Jun 1978, R.W. Brooks; 1 ♂, 5 Jul 1978, collector unknown; Hallelujah Junction: 1 ♀, 27 Jun 1949, A.S. Deal; 1 ♀, 27 Jun 1949, W.F. Erhardt; 2 ♂ 1 ♀, 4 Jul 1949, D. Cox; 1 ♀, 4 Jul 1949, E.L. Schlinger; 2 ♂, 4 Jul 1949, W.F. Ehrhardt; 2 ♀, 4 Jul 1949, W.F. Erhardt; 1 ♀, 7 Jul 1949, P.D. Hurd; 1 ♂, 12 Jul 1954, R.M. Bohart; 2 ♂ 2 ♀, 12 Jul 1962, M.E. Irwin; 1 ♂, 2 Jul 1964, S.A. Miller; 1 ♂, 7 Jul 1964, S.A. Miller; 1 ♂, 2 Jul 1966, S.A. Miller; 1 ♂, 4 Jul 1968, D.S. Horning, Jr.; 1 ♀, 28 Jun 1972, M.L. Kok; 1 ♀, 1 Jul 1974, B. Villegas; 2 ♂, 1 Jul 1974, R.M. Bohart; 1 ♂, 29 Jun 1976, R.D. Montgomery; 1 ♀, 27 Jun 1978, L.R. Bronson; 1 ♂ 1 ♀, 27 Jun 1978, M.S Gorman; 1 ♂ 3 ♀, 5 Jul 1978, L.D. French; 2 ♂ 3 ♀, 5 Jul 1978, L.S. Kimsey; 1 ♀, 5 Jul 1978, S.L. Heydon; 1 ♀, 27 Jul 1978, L.D. French; 1 ♀, 30 Jun 1981, L.D. French, P.S. Ward; Hallelujah Junction, Upper Long Valley: 2 ♂ 2 ♀, 4 Jul 1949, D. Cox; 1 ♀, 4 Jul 1949, L. Andres; 1 ♀, 4 Jul 1949, W.F. Ehrhardt; 1 ♀, 28 Jun 1962, R.L. Westcott; Hat Lake: 1 ♂, 25 Jul 1955, J.W. MacSwain; Herlong: 2 ♀, 4 Sep 1969, R.P. Allen; Milford, 7 mi N: 1 ♀, 8 Sep 1957, E.G. Linsley; Mono County: Bridgeport: 1 ♂, 13 Aug 1955, R.C. Bechtel; 1 ♂, 30 Jun 1961, J.G. Rozen; Crooked Creek Research Station, 0.24 mi SWbW: 1 ♀, 29 Jun 2015, T.L. Griswold; Deadman Summit: 1 ♂, 7 Aug 1968, T.R. Haig; 1 ♀, 4 Aug 1969, T.R. Haig; Hot Creek: 1 ♀, 17 Jul 1953, E.L. Schlinger; Leavitt Meadow, Walker River: 3 ♂, 25 Jun 1937, E.C. Van Dyke; 1 ♀, 25 Jun 1937, E.C. VanDyke; Lee Vining, N, Rt 395 N: 1 ♀ 2 ♀, 1 Aug 2012, K. Wright; Mammoth: 1 ♂ 1 ♀, 5 Jul 1933, G.E. Bohart, R.M. Bohart; 1 ♂, 10 Jul 1933, G.E. & R.M. Bohart; 1 ♂, 22 Jul 1936, G.E. Bohart, R.M. Bohart; 1 ♀, 5 Aug 1936, G.E. & R.M. Bohart; Mammoth Lake: 1 ♀, 24 Jul 1936, G.E. Bohart, R.M. Bohart; Mono Basin; Sulfur Spring Rd: 1 ♂, 30 Jul 1980, collector unknown; 2 ♀, 29 Aug 1980, L.S. Kimsey, R.B. Kimsey; 1 ♂ 2 ♀, 29 Aug 1980, collector unknown; 3 ♀, 30 Aug 1980, L.S. Kimsey, R.B. Kimsey; 1 ♀, 19 Aug 1981, collector unknown; Mono Lake, NW corner: 1 ♀, 24 Jun 1937, C.D. Michener; Pickel Meadow: 1 ♂, 11 Aug 1960, A.S. Menke; 1 ♂, 11 Aug 1960, D.Q. Cavagnaro; 1 ♂, 11 Aug 1960, R.R. Montanucci; 1 ♀, 16 Aug 1960, A.S. Menke; 1 ♂, 16 Aug 1960, D.Q. Cavagnaro; Pickle Meadow: 1 ♀, 11 Aug 1960, R.R. Montanucci; Rock Creek ( US 395): 1 ♀, 10 Sep 1956, collector unknown; White Mountains, Blanco's Corral: 1 ♀, 5 Jul 1952, W.D. McLellan; 1 ♂, 6 Jul 1953, J.T. Brooks; 2 ♀, 23 Jul 1953, J.W. MacSwain; 1 ♂, 6 Jul 1988, J.T. Brooks; north side of Mono Lake: 1 ♂ 3 ♀, 11 Jul 1975, S.C. Kuba; Nevada County : Boca: 1 ♀, 3 Jun 1954, R.C. Blaylock; 1 ♂ 1 ♀, 3 Jul 1954, R.M. Bohart; 1 ♂, 5 Jul 1954, R.C. Bechtel; Hobart Mills, 7 m N: 1 ♀, 26 Aug 1948, R.F. Smith; Hobart Mills, near: 1 ♂, 18 Jun 1962, M.E. Irwin; Sagehen Creek: 1 ♂, 26 Jun 1968, D.S. Horning, Jr.; 8 ♂ 2 ♀, 15 Jul 1978, L.S. Kimsey; Truckee: 1 ♀, 27 Jul 1939, C.T. Sierra; Placer County: Bear Valley: 1 ♂, 14 Jul 1956, R.R. Snelling; Brockway: 2 ♂ 3 ♀, Jul 1941, G.E. Bohart; Plumas County: Vinton: 1 ♀, 17 Jul 1973, B. Villegas; 1 ♂, 21 Jun 1984, No collector; 2 ♀, 21 Jun 1984, R.M. Bohart; Riverside County: Anza: 1 ♀, 24 Jun 1958, J.I. Stage; Sugarloaf Mountain: 1 ♂ 1 ♀, date unknown, collector unknown; San Bernardino County: Bear Valley: 1 ♂, 1 Jun 1928, E.C. Van Dyke; San Gorgonio Wilderness, Fish Creek Meadows: 1 ♀, 2 Sep 1984, K.W. Cooper; South Fork Santa Ana River, along Seven Oaks Road E of Barton Flats, San Bernardino Mountains, San Bernardino National Forest: 2 ♂, 13 Sep 1962, P.H. Timberlake; 5 ♀, 14 Sep 1962, P.H. Timberlake; Siskiyou County: Macdoel: 1 ♂, 2 Jun–2 Jul 1963, J. Schuh; Trinity County: : 1 ♀, 7 Jun 1934, G.E. & R.M. Bohart; Tulare County: Big Pine Meadow, NW: 1 ♂ 1 ♀, 28 Jun 2011, T.L. Griswold; Chimney Peak, 2.38 Air mi NbE: 1 ♂, 28 Jun 2011, T.L. Griswold; 6 ♂ 4 ♀, 26 Jun 2012, T.L. Griswold; Chimney Peak, 4.38 Air mi N: 9 ♂ 1 ♀, 28 Jun 2011, T.L. Griswold; 1 ♂ 1 ♂, 26 Jun 2012, T.L. Griswold; Crabtree Meadows: 1 ♂, 20 Jul 1935, H. Evans; Kennedy Meadows: 4 ♂, 28 Jun 2011, T.L. Griswold; Kennedy Meadows, Pine Mountain, 2.53 Air mi EbN: 1 ♀, 26 Jun 2012, T.L. Griswold; Rodeo Flat: 1 ♀, 26 Jun 2012, T.L. Griswold; Sirretta Road (Forest Service Road 22S21), 0.85 air mi E of Sherman Pass, 35.59 N, 118.20 W, Sequoia National Forest: 8 ♂ 11 ♀, 9–11 Sep 1995, R.R. Snelling; Troy Meadow: 1 ♂, 28 Jun 2011, T.L. Griswold; Troy Meadow Corral: 2 ♀, 26 Jun 2012, T.L. Griswold; Tussock Bench: 1 ♂ 1 ♀, 28 Jun 2011, T.L. Griswold; nr Troy Meadows, 9 Mile Canyon: 1 ♀, 6 Aug 1961, collector unknown; Tuolumne County: Chipmunk Flat: 3 ♂, 9 Aug 1960, M.E. Irwin; 1 ♂, 9 Aug 1960, R.R. Montanucci; Dardanelle: 1 ♂, 13 Jun 1951, E.L. Schlinger; 2 ♂, 13 Jul 1951, E.I. Schlinger; 2 ♂, 13 Jul 1951, E.L. Schlinger; 1 ♂, 13 Aug 1951, E.L. Schlinger; Dardanelles: 1 ♂, 13 Jul 1951, E. Schlinger; 1 ♀, 2 Jul 1957, J.W. MacSwain; Tioga Pass: 1 ♂, 3 Jul 1933, G.E. Bohart, R.M. Bohart; COLORADO: Alamosa County: Great Sand Dunes National Monument: 2 ♂ 5 ♀, 20–21 Jul 1954, H.E. Evans, M.A. Evans; 1 ♀, 4–7 Aug 1975, H.E. Evans; 1 ♀, 28 Jul 1988, G.C. Eickwort; Jackson County: East Sand Hill, 5 air km WSW; Blevins Ranch entrance: 4 ♀ 2 ♀, 2 Aug 2014, M.C. Orr; North Sand Hill, 5.8 air km E: 1 ♀, 2 Aug 2014, M.C. Orr; Moffat County: Craig, 28 mi W: 1 ♀, 21 Aug 1966, R.B. Roberts, M.G Roberts; Saguache County: Great Sand Dunes National Park (sand sheet): 1 ♀, 15 Aug 2012, J.D. Herndon; Teller County: Florissant: 1 ♂, Jul 1917, T.D.A. Cockerell; IDAHO: Ada County: Star, 1 mi S: 1 ♂, 5 Jul 1968, A.R. Gittins; Bannock County: Lava Hot Springs: 1 ♀, 7 Sep 1967, J.M. Gillespie; Bear Lake County: Bear Lake, N shore: 2 ♀, 10 Jul 1966, G.E. Bohart; Blaine County: Carey, 2 mi W: 1 ♂, 23 Jun 1917, W.F. Barr; Butte County: Craters of the Moon Nat. Mon.: 2 ♂, 20 Jun 1938, E.C. Van Dyke; 3 ♂, 3 Jul 1965, D.S. Horning, Jr.; 1 ♀, 17 Aug 1966, C.J. Horning; 4 ♀, 4 Jul 1967, D.S. Horning, Jr.; Howe, 20 mi NE: 1 ♀, 10 Sep 1968, W.F. Barr; Camas County: Hill City, 10 mi NW: 1 ♂, 15 Jul 1967, A.R. Gittins; Canyon County: Parma, 1mi W: 10 ♀, 28 Jun 1962, A.R. Gittins; Clark County: Lidy Hot Springs: 1 ♀, 6 Jul 1966, S.D. Horning; Custer County: Stanley, 5 mi E: 1 ♂, 2 Aug 1964, R.L. Westcott; Elmore County: Hill City, 9 mi W: 1 ♂, 7 Jul 1956, W.F. Barr; Franklin County: Preston: 1 ♀, 10 Jun 1968, G.E. Bohart; Preston, 3 mi N: 1 ♂ 2 ♀, 15 Sep 1970, P.F. Torchio; Fremont County: Island Park Siding, Island Park: 1 ♂, 2 Aug 1957, B. Knapp; St. Anthony Sand Dunes: 1 ♀, 23 Jun 1961, J.E. Henry; 2 ♀, 23 Jun 1961, W.F. Barr; 1 ♂ 1 ♀, 5 Aug 1967, D.S. Horning, Jr.; St. Anthony sand dunes: 3 ♂ 3 ♀, 1–2 Jul 1980, M.S. Wasbauer; 2 ♂ 3 ♀, 9–10 Jul 1980, M.S. Wasbauer; 2 ♂, 11 Jul 1980, M.S. Wasbauer; St. Anthony, 8 mi NW, sand dunes: 1 ♀, 29 Jun 1977, J.M. Domek; St. Anthony, sand dunes NW: 1 ♂ 4 ♀, 23 Jun 1961, W.F. Barr; 2 ♂, 7 Sep 1967, W.F. Barr; 1 ♂, 28 Jun 1977, J. Domek; 1 ♂, 28 Jun 1977, W.F. Barr; Gooding County: Wood River, 1 mi NE Gooding: 1 ♂, 6–7 Jul 1980, M.S. Wasbauer; Jefferson County: Terreton, 4 mi NW: 1 ♂, 28 Jun 1964, W.F. Barr; Jerome County: Jerome: 1 ♂, 24 Aug 1953, A.R. Gittins; Lincoln County: Shoshone: 1 ♀, 10 Aug 1955, J.E. Gillaspy; Owyhee County: Indian Cove: 1 ♀, 13 Sep 1955, W.F. Barr; Twin Falls County: Bruneau Desert; Twin Buttes, 21.4km SE: 1 ♀, 14 Sep 2010, B. Love; NEVADA:Douglas County : Dagget Pass: 1 ♂, 18 Aug 1957, E.L. Schlinger; Daggett Pass, Toiyabe National Forest: 1 ♂, 18 Aug 1951, E.I. Schlinger; Minden: 1 ♂, 24 Aug 1952, R.M. Bohart; Elko County: Ruby Mountains, Island Lake, below: 1 ♀, 22 Jul 1976, T.L. Griswold; Ruby Mountains, Lamoille Creek: 2 ♂, 22 Jul 1976, T.L. Griswold; 1 ♂, 22 Jul 1977, T.L. Griswold; 6 ♂ 2 ♀, 23 Jul 1977, T.L. Griswold; Humboldt County: Winnemucca: 1 ♀, 30 Aug 1956, T.R. Haig; Washoe County: Carson City: 1 ♂ 1 ♀, 25 Jun 1929, E.P. Van Duzee; Reno, 6 mi W: 2 ♂, 29 Jun 1955, J.C. Downey; Reno, Sky Ranch: 1 ♀, 4 Jul 1952, E.I. Schlinger; Verdi: 1 ♂, 25 Jun 1954, R.M. Bohart; 1 ♂, 25 Jun 1961, F.D. Parker; 1 ♂, 18 Jun 1964, R.M. Bohart; 1 ♂, 30 Jun 1964, F.D. Parker; 1 ♂, date unknown, collector unknown; Washoe City: 1 ♂, 8 Sep 1960, A.S. Menke; NEW MEXICO: McKinley County: 19 mi N of Gallup: 1 ♂, 14 Aug 1972, J.G. Rozen, R.J. McGinley; San Juan County: Bloomfield, 30 m S: 1 ♂, 15 Sep 1966, R.W. Rust, P.F. Torchio, G. Wood, N. Youssef; Chaco River, N of Tsaya: 3 ♂, 8 May 1989, T.L. Griswold; Shiprock: 1 ♂, 24 Sep 1970, G.E. Bohart, R.M. Bohart; OREGON: Crook County: Bend, 53 mi E: 1 ♀, 17 Jul 1927, H.A. Scullen; Harney County: 20 mi S of Burns: 1 ♂, 13 Jul 1927, H.A. Scullen; Lake County: Silver Lake, 52 km SW; hwy 86 @ mile marker 18: 2 ♀, 21 Jul 2010, M.E. Irwin, F.D. Parker; Silver Lake, 26 km NW; on hwy 31: 2 ♀, 21 Jun–29 Jul 2010, M.E. Irwin, F.D. Parker; Malheur County: Adrian: 1 ♂, 7 Sep 1959, P.F. Torchio; 1 ♀, 8 Sep 1959, P.F. Torchio; Morrow County: Boardman: 1 ♀, 23 Jun 1961, C.E. Osgood; 1 ♀, 24 Jun 1961, C.E. Osgood; 2 ♀, 14 Sep 1961, C.E. Osgood; 1 ♀, 13 May 1962, collector unknown; 2 ♀, 5 Jul 1962, collector unknown; UTAH: Cache County: Cornish: 10 ♂ 3 ♀, 10 Jul 1959, No collector; 1 ♂, 10 Jul 1959, collector unknown; 2 ♀, 29 Jul 1959, No collector; 1 ♀, 4 Aug 1959, No collector; 1 ♀, 6 Aug 1959, D. Davis; 1 ♂, 6 Aug 1959, D.W. Davis; 2 ♀, 6 Aug 1959, G.E. Bohart; 9 ♂, 15 Aug 1962, G.E. Bohart, P.F. Torchio; 3 ♂, 15 Aug 1962, collector unknown; 1 ♀, 23 Aug 1962, R.M. Bohart; 1 ♀, 26 Jun 1963, G.E. Bohart; 1 ♀, 10 Jul 1963, G.E. Bohart, P.F. Torchio; 2 ♂, 13 Aug 1963, G.E. Bohart; 1 ♀, 7 Sep 1968, C.D. Michener; 1 ♂ 1 ♀, 2 Jul 1970, P.F. Torchio; 1 ♀, 12 Jul 1971, E.E. Grissell; 1 ♀, 8 Sep 1972, F.D. Parker; Cornish, 4 mi NE: 2 ♂ 2 ♀, 28 Jul 1970, P.F. Torchio; Cornish, near: 2 ♂, 8 Aug 1963, No collector; Logan: 1 ♂, 4 Aug 1965, R.M. Bohart; White Pine Lake Trail, Logan Canyon: 1 ♂, 3 Aug 2013, D. Denlinger; Duchesne County: Duchesne: 1 ♂, 11 Aug 1962, R.M. Bohart; Emery County: 2 mi ESE Temple Mountain,: 2 ♂, 2 Jun 1994, R.R. Snelling; Buckskin Spring, N Goblin Valley: 1 ♂ 1 ♀, 29 May 1981, F.D. Parker; 1 ♂, 2 Jun 1982, D.R. Frohlich, D.F. Veirs; 1 ♂, 3 Jun 1982, T.L. Griswold; Buckskin Springs, Goblin Valley: 1 ♀, 21–26 May 2002, F.D. Parker, M.E. Irwin; Calf Canyon, San Rafael Swell: 1 ♀, 7 Jun 1982, T.L. Griswold, R.T. Griswold; 2 ♂, 7 Jun 1982, collector unknown; Cottonwood Spring, 2.5 mi SE: 4 ♀, 14 Sep 1983, F.D. Parker, T.L. Griswold; Dugout Spring, 2.5 air mi NE Big Flat Top: 1 ♂, 28 Jul 1983, F.& A. Parker, T.L. Griswold; 7 ♀, 14 Sep 1983, F.D. Parker, T.L. Griswold; Gilson Butte Well: 1 ♀, 18 Sep 1980, W.P. Nye; 1 ♂ 3 ♀, 23 Aug 1981, W.P. Nye; 1 ♂ 3 ♀, 12 Sep 1981, W.P. Nye; 7 ♀, 19 Jun 1982, W.P. Nye; 9 ♀ 5 ♀, 4 Sep 1982, W.P. Nye; 1 ♀, 27 Aug 2001, M.E. Irwin, F.D. Parker, M. Metz, M. Hauser, C. Lambkin; Gilson Butte Well, 25 km NNE Hanksville: 4 ♀, 22–26 Aug 2001, F.D. Parker, C. Lambkin, M. Metz; 5 ♀, 22–26 Aug 2001, F.D. Parker, C. Lambkin, M. Metz, M. Hauser, M.E. Irwin; Gilson Butte, 4 air mi N: 6 ♀, 16 Sep 1980, F.D. Parker; 1 ♂ 3 ♀, 16 Sep 1980, T.L. Griswold; 1 ♂ 3 ♀, 13 Sep 1983, F.D. Parker, T.L. Griswold; Goblin Valley: 1 ♂, 14 Sep 1979, W. Nye; Goblin Valley Road, Junction Highway 24: 2 ♀, 8 Jun 1988, W.P. Nye; Goblin Valley State Reserve, 2 mi N: 1 ♂ 1 ♀, 25 Aug 1980, A.&K. Menke, F.D. Parker; Goblin Valley turnoff, 25 mi N Hanksville: 6 ♂ 8 ♀, 16 Sep 1979, F.D. Parker; 3 ♀, 16 Sep 1979, F.D. Parker, D.F. Veirs; 1 ♀, 5 Jun 1981, D.F. Veirs; Goblin Valley, sand dunes: 5 ♀, 16 Sep 1979, C.L. Hatley, G. Briggs; 1 ♂ 8 ♀, 16 Sep 1979, F.D. Parker, D.F. Veirs; 1 ♂ 2 ♀, 20 Jun 1980, F.D. Parker; Goblin Valley, wash: 1 ♂ 7 ♀, 16 Sep 1979, F.D. Parker, D.F. Veirs; Hanksville, 12 mi N, Emery County line: 1 ♂ 4 ♀, 22 Aug 1979, F.D. Parker; Hatt Ranch, 17 mi W Green River: 1 ♂, 23–24 Aug 2001, M.E. Irwin, F.D. Parker; 4 ♂, 24–25 Aug 2001, M.E. Irwin, F.D. Parker; Hatt's Ranch, 2 km S: 3 ♀, 21–26 May 2002, M.E. Irwin, F.D. Parker; Iron Wash, 2.5 mi SW: 1 ♂ 1 ♀, 2 Jun 1982, T.L. Griswold; Little Flat Top: 1 ♀, 26 Aug 2001, F.D. Parker, C. Lambkin, M. Metz, M. Hauser, M.E. Irwin; Little Flat Top, San Rafael Desert: 1 ♂, 22 Aug 1989, J. Burner; Little Gilson Butte, 0.5 air mi NE: 1 ♀, 12 Sep 1983, F.D. Parker, T.L. Griswold; Little Gilson Butte, 0.5 mi E: 1 ♀, 27 Aug 1985, T.L. Griswold; Little Gilson Butte, 2 air mi E: 1 ♂, 29 May 1981, F.D. Parker; 2 ♀, 23 Jul 1981, D.F. Veirs, T.L. Griswold, F.D. Parker; 11 ♀, 25 Aug 1981, D.F. Veirs, T.L. Griswold, F.D. Parker; 1 ♂, 3 Jun 1982, D.R. Frohlich, D.F. Veirs; 3 ♀, 3 Jun 1982, F.D. Parker, T.L. Griswold; 1 ♀, 14 Sep 1983, F.D. Parker, T.L. Griswold; 1 ♂, 16 Sep 1989, T.L. Griswold; Little Gilson Butte, 2 air mi W: 2 ♀, 16 Sep 1989, D.F. Veirs; 2 ♀, 16 Sep 1989, F.D. Parker; 3 ♂ 4 ♀, 16 Sep 1989, T.L. Griswold; Little Gilson Butte, 3.2 air mi NE: 1 ♀, 3 Jun 1981, F.D. Parker; 1 ♂ 6 ♀, 3 Jun 1982, D.R. Frohlich, D.F. Veirs; 1 ♂ 4 ♀, 4 Jun 1982, F.D. Parker, T.L. Griswold; 2 ♀, 14 Jun 1983, F.D. & J.H. Parker; 1 ♂ 3 ♀, 13 Sep 1983, F.D. Parker, T.L. Griswold; 1 ♂, 28 May 1985, F.D. Parker; 1 ♂, 28 May 1985, R.T. Griswold; 1 ♂, 28 May 1985, T.L. Griswold; Little Gilson Butte, E side: 4 ♀, 4 Jun 1982, F.D. Parker; 3 ♀, 4 Jun 1982, F.D. Parker, T.L. Griswold; Little Gilson Butte, near: 11 ♂ 6 ♀, 25 Aug 1980, A.&K. Menke, F.D. Parker; North Temple Wash: 1 ♂, 14 Jun 1983, F.D. & J.H. Parker; 1 ♂ 1 ♀, 15 Jun 1983, T.L. Griswold; San Rafael Desert, near Goblin Valley: 1 ♀, Sep 1980, G.E. Bohart; Sinbad Country, E of Block Mountain: 1 ♀, 24 Jul 1981, Parker, D. Veirs, T.L. Griswold; South Temple Wash: 3 ♂ 7 ♀, 15 Sep 1980, T.L. Griswold; 1 ♂ 1 ♀, 21 Aug 2001, F.D. Parker, M.E. Irwin, M. Metz, M. Hauser, C. Lambkin; 1 ♀, 22–26 Aug 2001, F.D. Parker, C. Lambkin, M. Metz, M. Hauser, M.E. Irwin; Temple Mountain, 3 mi SSE: 2 ♀, 3 Jun 1982, F.D. Parker, T.L. Griswold; Wild Horse Creek, N Goblin Valley: 3 ♀, 16 Sep 1980, F.D. & J.H. Parker; 5 ♀, 16 Sep 1980, F.D. Parker; 2 ♀, 16 Sep 1980, T.L. Griswold; 8 ♀, 22 Jul 1981, D.F. Veirs, T.L. Griswold, F.D. Parker; 1 ♀, 3 Jun 1982, D.R. Frohlich, D.F. Veirs; 4 ♂ 2 ♀, 3 Jun 1982, F.D. Parker, T.L. Griswold; 1 ♂ 1 ♀, 14 Jun 1983, T.L. Griswold; 1 ♀, 13 Sep 1983, F.D. Parker, T.L. Griswold; 2 ♀, 16 Aug 1992, T.L. Griswold; 1 ♂, 5 Aug 1997, F.D. Parker; Woman Wash: 2 ♂, 7 Jun 1982, F.D. Parker; Garfield County: Big Flat: 1 ♂, 15 Aug 2001, C.M. Davidson; 1 ♂ 1 ♀, 15 Aug 2001, O.J. Messinger; 1 ♂, 15 Aug 2001, R. Andrus; 2 ♂, 15 Aug 2001, S. Messinger; Boulder, 6.2 km SE: 1 ♀, 27 Aug 2015, M.C. Orr; Calf Creek: 1 ♂, 29 Aug 1985, T.L. Griswold; 2 ♂, 10 Sep 2001, B. Morgan; 1 ♂, 30 Aug 2002, O.J. Messinger; 1 ♂, 30 Sep 2002, B. Bradley; 1 ♀, 23 Sep 2003, S.M. Higbee; Deer Creek: 3 ♀, 2 Jun 2000, T.L. Griswold; 1 ♀, 25 May 2001, K. Moredock; Deer Creek Campground, 7 km SE Boulder: 2 ♀, 23–27 May 2002, F.D. Parker, M.E. Irwin; 1 ♀, 21 Jun 2005, F.D. Parker; Duffy Mesa, 1.8 mi NNW: 1 ♂, 31 Aug 2000, M. Sunseri; 6 ♀, 7 Jun 2001, O.J. Messinger; 2 ♀ 11 ♀, 7 Jun 2001, R. Andrus; 7 ♂, 14 Aug 2001, O.J. Messinger; 6 ♂ 2 ♀, 28 Aug 2001, R. Andrus; 1 ♂ 3 ♀, 31 Aug 2001, O.J. Messinger; 3 ♂, 10 Sep 2001, K. Moredock; 2 ♂, 10 Sep 2001, R. Andrus; 1 ♂ 2 ♀, 27 Aug 2002, C.M. Davidson; 1 ♀, 12 Sep 2002, L. Wilson; 2 ♀ 1 ♀, 27 Sep 2002, J. Hawk; 3 ♀, 9 Oct 2002, H. Ikerd; 1 ♀, 3 Jul 2003, H. Ikerd; 1 ♀, 1 Sep 2003, S.M. Higbee; Duffy Mesa, 2 mi NNW: 1 ♂ 1 ♀, 28 Aug 2001, R. Andrus; 3 ♂, 10 Sep 2001, R. Andrus; 1 ♀, 27 Sep 2002, J. Hawk, L. Wilson; 1 ♀, 2 Jun 2003, H. Ikerd; 2 ♂, 29 Jul 2003, C. Boyers; 1 ♂ 1 ♀, 12 Aug 2003, B. Bradley; 8 ♂ 3 ♀, 12 Aug 2003, H. Ikerd; 2 ♂ 1 ♀, 1 Sep 2003, A. Johansen; 6 ♂ 2 ♀, 1 Sep 2003, S.M. Higbee; 1 ♂ 1 ♀, 15 Sep 2003, A. Johansen; 5 ♂ 1 ♂, 15 Sep 2003, J. Tolliver; 3 ♂ 1 ♂, 29 Sep 2003, A. Johansen; 2 ♂ 1 ♀, 29 Sep 2003, S.M. Higbee; Escalante Rim, 1.13 mi NW: 1 ♀, 22 Aug 2003, H. Ikerd; Escalante Riv. Jct Death Hollow, 1.1 mi W: 2 ♂, 30 Aug 2002, J. Hawk; Escalante Riv., jct Death Hollow, 1.3 mi W: 1 ♀, 28 May 2001, C.M. Davidson; 1 ♀, 30 Aug 2002, C.M. Davidson; Escalante River: 3 ♂ 1 ♀, 28 May 2001, S. Messinger; Escalante River, jct Hwy 12: 2 ♂ 1 ♂, 22 Sep 2003, J. Tolliver; Escalante, 20 km N: 1 ♂, 25–28 May 2002, M.E. Irwin, F.D. Parker; Hanksville, 34 mi S: 2 ♂ 16 ♀, 18 Sep 1980, W.J. Hanson, G.E. Knowlton; Hot Canyon, 0.9 mi S: 1 ♂ 4 ♀, 7 Jun 2003, H. Ikerd; Hot Canyon, 1.1 mi E: 1 ♂ 1 ♀, 7 Jun 2003, H. Ikerd; Hot Canyon, 1.2 mi SSE: 1 ♀ 5 ♀, 7 Jun 2003, H. Ikerd; Long Canyon: 1 ♂, 31 Aug 2000, S. Jenkins; 1 ♂ 1 ♀, 1 Sep 2003, T.L. Griswold; 1 ♂ 3 ♀, 15 Sep 2003, S.M. Higbee; 1 ♀, 29 Sep 2003, J. Tolliver; Long Cyn, 0.8 mi NNW mouth: 1 ♀ 1 ♀, 31 Aug 2000, T.L. Griswold; 1 ♂, 25 Sep 2000, A. Worley; 2 ♀, 25 May 2001, E. Mullaney; Milk Creek, SE of Mount Hillers: 1 ♀, 16 Jun 1983, T.L. Griswold; Mount Hillers, E slope: 1 ♀, 16 Jun 1983, F.D. Parker, J. Parker; 1 ♀, 16 month unknown 1983, No collector; North Wash, SSE Hanksville: 1 ♀, 28 Aug 1985, T.L. Griswold; Pine Creek, N of jct Deep Creek: 1 ♂, 21 Jun 2000, T.L. Griswold; Rattlesnake Bench, 1.3 mi SSW: 2 ♀, 25 May 2001, C.M. Davidson; Red Breaks, 1.6 mi W: 5 ♀ 1 ♀, 25 Jun 2003, H. Ikerd; 1 ♀, 1 Oct 2003, S.M. Higbee; Red Breaks, 2.3 mi NW: 3 ♂ 1 ♀, 25 Jun 2003, H. Ikerd; 2 ♂, 8 Aug 2003, H. Ikerd; 4 ♀ 1 ♀, 1 Oct 2003, J. Tolliver; 1 ♀, 1 Oct 2003, S.M. Higbee; 2 ♀, 4 Oct 2003, A. Johansen; Steep Creek, 0.5 mi E jct Long Canyon: 2 ♂ 1 ♀, 1 Sep 2003, T.L. Griswold; Tenmile Spr., 1.86 mi WNW: 1 ♀, 1 Oct 2003, S.M. Higbee; The Gulch: 1 ♂ 1 ♀, 25 May 2001, C.M. Davidson; 1 ♀ 4 ♀, 25 May 2001, S. Messinger; White Point, 2.3 mi NNW: 1 ♀, 4 Jun 2001, K. Moredock; 3 ♀, 4 Jun 2001, R. Andrus; 1 ♂ 1 ♀, 23 Aug 2001, R. Andrus; White Point, 3.4 mi NNW: 1 ♀, 4 Jun 2001, R. Andrus; Grand County: : 1 ♀, 11 Jun 1963, L.B. Coons; Arches National Park, Salt Valley, NW: 1 ♂, 4 Jun 1982, R. & T. Donotuse Griswold; Moab: 1 ♀, 30 Sep 1960, G.E. Bohart; 4 ♀ 1 ♀, 9 Jun 1963, G.E. Bohart, R.L. Brumley; 1 ♀, 6 Sep 1967, P.F. Torchio; Moab, 14 mi S: 5 ♀, 3 Oct 1965, G.E. Bohart; 2 ♂ 3 ♀, 15 Oct 1965, P.F. Torchio; 10 ♀, 15 Oct 1965, collector unknown; Moab, 16 mi S: 23 ♂ 14 ♀, Sep 1963, P.F. Torchio; Moab, 25 mi S: 1 ♂, 10 Jun 1963, G. Bohart, R.L. Brumley; Seven Mile Canyon, 13 mi NW Moab: 1 ♀, 6 Jun 1982, R. & T. Donotuse Griswold; 1 ♀, 6 Jun 1982, T.L. Griswold, R.T. Griswold; Juab County: Little Sahara Sand Dunes: 1 ♂ 1 ♀, 14 Aug 1992, T.L. Griswold; 2 ♂ 1 ♀, 14 Sep 1993, T.L. Griswold; Kane County: Billy Pasture, 0.4 mi N: 4 ♀, 26 Jun 2003, H. Ikerd; Burnt Ridge, 1 mi N: 1 ♂, 2 Jul 2003, H. Ikerd; Coral Pink Sand Dunes: 1 ♂, 5 Jul 1968, L.S. Hawkins, Jr.; Coral Pink Sand Dunes State Park: 1 ♂, 13 Jul 1998, R.W. Baumann, et al; Cougar Knoll, 0.61mi SSW: 2 ♂, 31 May 2000, C.M. Davidson; Cougar Knoll, 1.32 mi SSW: 1 ♀, 3 Sep 2003, T.L. Griswold; Dry Fork: 1 ♂, 26 Aug 2003, S.M. Higbee; Dry Fork, N: 2 ♂ 4 ♀, 26 Aug 2003, S.M. Higbee; 2 ♂, 18 Sep 2003, S.M. Higbee; GSENM; Mollies Nipple, 7.1 air km W; roadside: 1 ♂ 1 ♂, 25 Sep 2014, M.C. Orr; 1 ♂, 13 Sep 2015, M.C. Orr, D. Denlinger; Georgie Hollow, 0.6 mi NE: 1 ♀, 31 Jul 2003, C. Boyers; Jody's Knoll, 1.0 mi SSW: 1 ♀ 1 ♀, 26 Jun 2003, H. Ikerd; 1 ♂ 1 ♀, 20 Aug 2003, S.M. Higbee; 1 ♂ 1 ♀, 5 Sep 2003, T.L. Griswold; Kitchen Corral Rd.: 2 ♀, 4 Sep 2003, T.L. Griswold; Left Hand Collet Cyn, 2.7 mi SE: 1 ♂, 1 Aug 2000, O.J. Messinger; Lick Wash: 1 ♀, 26 Aug 2002, J. Hawk; Mount Carmel Junction, 3.5 mi SE: 1 ♀, 29 Aug 1990, T.L. Griswold; Pool Hollow, nr.: 2 ♂, 1 Aug 2002, J.S. Wilson; 4 ♂ 4 ♂, 15 Aug 2002, S. Messinger; Red Wash, 2.6 mi SSE Carly Knoll: 1 ♂, 6 Jun 2001, R. Andrus; Sunset Natural Arch, 0.72 mi NE: 1 ♂, 8 May 2001, T.L. Griswold; Millard County: Delta: 1 ♀, 15 Aug 1952, G.E. Bohart, E.A. Cross; McCormick: 1 ♂, 19 Jun 1950, G.F. Knowlton, G.E. Bohart; Meadow, 19 mi N: 3 ♀, 8 Jun 1969, S.M. Hogue; Pahvant: 1 ♀, 20 Jun 1952, E.A. Cross; 1 ♀, 20 Jun 1952, G.E. Bohart, E.A. Cross; 1 ♀, 20 Jun 1952, collector unknown; Pahvant, sand dunes: 1 ♀, 20 Jun 1952, G.E. Bohart, E.A. Cross; Rich County: Laketown: 1 ♀, 21 Jul 1966, W.J. Hanson; San Juan County: La Sal Junction, 6 m N: 1 ♂ 5 ♀, 3 Oct 1965, R.W. Thorp; 1 ♂, 3 Oct 1965, collector unknown; Moab, 12 mi S: 4 ♂ 1 ♀, 22–23 Jul 1979, F.D. Parker; Moab, 16 mi S: 1 ♀, Sep 1963, P.F. Torchio; 1 ♂ 1 ♀, 2 Oct 1965, P. Torchio; Sevier County: Fremont Junction, 15 mi W: 1 ♀, 18 Aug 1966, Bohart, Cross; Tooele County: 0.7 mi NW Little Granite Mt.: 1 ♀, 10 Jun 2003, O.J. Messinger; 1 ♀, 14 Jun 2005, E. Jarrell; 1 ♂ 1 ♀, 14 Jun 2005, J.S. Wilson; 1 ♀ 2 ♀, 22 Jun 2005, E. Jarrell; 3 ♂ 1 ♀, 22 Jun 2005, J.S. Wilson; 1 ♂, 29 Sep 2005, T.L. Griswold; 1.3 mi W Little Granite Mt.: 1 ♂ 1 ♂, 14 Jun 2005, J.S. Wilson; 1 ♂, 29 Sep 2005, K.T. Huntzinger; 1 ♂, 29 Sep 2005, T.L. Griswold; 1.9 mi N Little Granite Mtn.: 1 ♂, 10 Jun 2003, R. Andrus; 2 mi N Little Granite Mt.: 1 ♀, 10 Jun 2003, O.J. Messinger, R. Andrus; 1 ♀, 11 Sep 2005, K.T. Huntzinger; 1 ♂, 14 Sep 2005, K.T. Huntzinger; 1 ♀, 14 Sep 2005, O.J. Messinger; 1 ♀ 1 ♀, 29 Sep 2005, K.T. Huntzinger; 2.8 mi W Simpson Buttes: 3 ♀, 28 Sep 2005, K.T. Huntzinger; 1 ♀ 3 ♀, 28 Sep 2005, T.L. Griswold; 1 ♀, 28 Sep 2005, T.L. Griswold, K.T. Huntzinger; 3.2 mi SSE Wig Mt.: 1 ♀, 12 Jun 2003, O.J. Messinger; 1 ♂, 12 Jun 2003, R. Andrus; 4.6 mi WSW Little Granite Mtn.: 1 ♂, 28 Sep 2005, K.T. Huntzinger; 1 ♂, 28 Sep 2005, T.L. Griswold; Camels Back Ridge, 3 mi NNE: 1 ♀, 19 Jun 2003, O.J. Messinger; Uintah County: Bonanza: 2 ♀, 30 Aug 1975, G.E. Bohart; 1 ♀, 18 Sep 1975, G.E. Bohart; Bonanza, SW1: 2 ♂, 11 Aug 1981, M. Schwartz, R. Miller; 1 ♂, 21 Aug 1981, M. Schwartz, R. Miller; 3 ♀, 28 Aug 1981, M. Schwartz, R.E. Miller; 1 ♂, 5 Sep 1981, M. Schwartz, R. Miller; Little Brush Creek: 1 ♂, 12 Aug 1981, W.J. Hanson, S. Keller; Utah County: Provo: 1 ♂ 1 ♀, 24 Jun 1912, T. Spalding; Washington County: Stave Spr., 0.80 mi WSW: 1 ♀, 19 Jun 2007, H. Ikerd; Zion Canyon, Zion National Park: 1 ♀, 6 May 1948, G.E. Bohart; Zion Cyn: 1 ♀, 6 May 1948, G.E. Bohart; Wayne County: Big Flat Top, 7 mi E: 3 ♂ 2 ♀, 29 May 1985, F.D. Parker; 3 ♂ 2 ♀, 29 May 1985, T.L. Griswold; 1 ♂, 30 May 1985, R.T. Griswold; Capitol Reef National Park: 1 ♀, 16 month unknown 1979, No collector; 1 ♀, 15 month unknown 1981, No collector; 1 ♀, 4 Sep 1982, W.P. Nye; 1 ♀, 4 month unknown 1982, W.P. Nye; E edge: 1 ♀, 15 Sep 1979, C.L. Hatley, G. Briggs; 1 ♀, 16 Sep 1979, F.D. Parker, D.F. Veirs; 3 ♀, 15 Sep 1981, C.L. Hatley, G. Briggs; Fremont River, 27 mi W Hanksville: 1 ♀, 27 Aug 1980, A.S. Menke, F.D. Parker, K.A. Menke; 1 ♀, 27 month unknown 1980, No collector; Hanksville: 4 ♀, 31 Aug 1982, R.M. Bohart; 2 ♀, 5 Sep 1982, W.P. Nye; Hanksville, 5.5 mi N: 2 ♀, 12 Sep 1983, T.L. Griswold; Jct Bull Mountain Rd & Hwy 95, 12 km SW: 1 ♀, 27 May 2002, M.E. Irwin, F.D. Parker; WASHINGTON: Benton County: Hanford Site, 1200 Ft Rd, dirt road: 1 ♂ 1 ♀, 5 Jun 1995, R.S. Zack; Chelan County: Wenatchee: 1 ♂, 20 Aug 1907, A.L. Melander; WYOMING: Albany County: Dirt Farm, 8 mi SE Laramie: 1 ♂, 17 Jul 1974, V.J. Tepedino; 1 ♂, 19 Jul 1974, V.J. Tepedino; 1 ♂, 26 Jul 1974, V.J. Tepedino; 1 ♂, 14 Jul 1975, V.J. Tepedino; 1 ♂, 2 Aug 1975, V.J. Tepedino; 2 ♂, 12 Jul 1976, V.J. Tepedino; 1 ♂ 1 ♂, 14 Jul 1976, V.J. Tepedino; 1 ♂ 1 ♀, 23 Jul 1976, V.J. Tepedino; Carbon County: Bear Mountain, SE Slope: 1 ♀, 22 Jun 2005, B. Bradley; Ferris Mt E, 7.5km SE; Ferris Dunes: 2 ♂ 3 ♀, 26 Jul 2012, M.C. Orr; Ferris Mt E, 8km S; Ferris Rd: 1 ♀, 26 Jul 2012, M.C. Orr; Fremont County: Boysen Dam: 1 ♀, 20 Aug 1969, C.D. Michener; Shoshoni: 1 ♂, 21 Jul 1965, G. Bohart, P. Torchio; Lincoln County: Kemmerer: 1 ♂, 13 Jul 1955, No collector; Sweetwater County: Boar's Tusk; vegetated sand dunes: 1 ♂, 9 Jun 2004, T.L. Griswold, S. Laske; 1 ♀, 15 Jun 2004, S. Laske; 4 ♀, 9 Sep 2004, S. Laske; Farson, 20 mi E: 1 ♀, 4 Aug 1955, G.E. Bohart; Jim Bridger Mine: 1 ♂, 24 Jun 1979, No collector; Jim Bridger Mine, Point of Rocks: 1 ♂, 24 Jun 1979, D.G. Krementz; Little America: 1 ♂, 25 Jun 1960, G.I. Stage; Little America, 22 mi W Green River: 1 ♂ 1 ♀, 25 Jun 1960, G.I. Stage; Little America, 22 mi W of the Green River: 1 ♀, 25 Jun 1960, G.I. Stage; Red Desert: 3 ♂, 6 Sep 1979, F.D. Parker.

Figures

FIGURE 7. Distribution of Anthophora albata. Generated with 1325 specimen records from 313 locations.

FIGURE 34. Mandible, inferior tooth. Scale=0.5 mm. A) Triangular, tip clearly angled (A. albata female, BBSL510289). B) Ovular, tip and general form rounded (A. estebana male, BBSL276281). C) Ridge-like, concave above, often projecting off main blade at tip (A. abroniae female, BBSL502235). D) Absent (A. abroniae male, BBSL502217).

FIGURE 38. Male sterna, S5 setae density and S6 rim form. Scale=0.5 mm. A) S5 with sparse setae; S6 medially emarginate, at least vaguely triangularly, evenly so (A. estebana, 72476). B) S5 setae density high, appearing tufted; S6 medially emarginate, at least vaguely triangularly, but at least slightly filled medially, sometimes tufted (A. maculifrons, 82068). C) S5 with sparse setae; S6 entire or nearly so, with transparent square of integument (A. petrophila, BBSL655748). D) S5 setae density high; S6 entire, medially knobbed (A. flavocincta, BBSL504065).

FIGURE 39. Female face of A) Anthophora abroniae (BBSL920115), B) A. albata (10234), C) A. caudata (BBSLID99019), D) A. chihuahua (BBSL502616), E) A. cockerelli (BBSL502283), F) A. columbariae (BBSL853114). Scale=0.5 mm.

FIGURE 44. Female habitus of A) Anthophora abroniae (BBSL920116), B) A. albata (BBSL513478), C) A. caudata (BBSL506712), D) A. chihuahua (BBSL502616), E) A. cockerelli (BBSL502289), F) A. columbariae (KWC0014759). Scale=1 mm.

FIGURE 49. Male face of A) Anthophora abroniae (BBSL502212), B) A. albata (BBSL504186), C) A. caudata (BBSLID98988), D) A. chihuahua (BBSL502617), E) A. cockerelli (BBSL502260), F) A. columbariae (BBSL853110). Scale=0.5 mm.

FIGURE 54. Male habitus of A) Anthophora abroniae (BBSL502223), B) A. albata (10046), C) A. caudata (BBSLID92552), D) A. chihuahua (BBSL502617), E) A. cockerelli (72472), F) A. columbariae (BBSL853157). Scale=1 mm.

FIGURE 59. Male T7 of A) Anthophora abroniae (BBSL348460), B) A. albata (9988), C) A. caudata (BBSL506708), D) A. chihuahua (BBSL502618), E) A. cockerelli (BBSL502252), F) A. columbariae (FDP80368). Scale=0.25 mm.

FIGURE 64. Male S7 of A) Anthophora abroniae (BBSL502216), B) A. albata (BBSL843758), C) A. caudata (BBSL506707), D) A. chihuahua (BBSL502617), E) A. cockerelli (BBSL502279), F) A. columbariae (BBSL502330). Scale=0.25 mm.

FIGURE 69. Male S8 of A) Anthophora abroniae (BBSL502211), B) A. albata (BBSL843758), C) A. caudata (BBSL506707), D) A. chihuahua (BBSL502617), E) A. cockerelli (BBSL502279), F) A. columbariae (BBSL502330). Scale=0.25 mm.

FIGURE 74. Male genital capsule of A) Anthophora abroniae (BBSL502216), B) A. albata (BBSL843758), C) A. caudata (BBSL506707), D) A. chihuahua (BBSL502617), E) A. cockerelli (BBSL502263), F) A. columbariae (BBSL502332). Scale=0.25 mm.