Zammara intricata Walker, 1850

Sanborn, Allen F., 2020, The cicadas (Hemiptera: Cicadidae) of Ecuador including the description of five new species, a new subtribe, four new synonymies, and fifteen new records, Zootaxa 4880 (1), pp. 1-80 : 13-14

Zammara intricata Walker, 1850

Zammara intricata Walker 1850: 35 . (Guayaquil, Ecuador)

Zammara erna Schmidt 1919: 390 . (Pucay, Ecuador) n. syn.

Orellana pulla Goding 1925: 25 . (Milagro, Naranjapata, and Macas, Ecuador) n. syn.

Orellana brunneipennis Goding 1925: 27 . (Rio Lliquino and Tenguel, Ecuador) n. syn.

Orellana brunnpennis (sic) Goding 1925: 7 .

REMARKS.— Goemans (2016) demonstrated the Ecuadorian species Orellana brunneipennis and Orellana pulla belong in the genus Zammara rather than Orellana Distant, 1905b and that these species are junior synonyms of Z. intricata Walker, 1850 as is Z. erna . However, Goemans (2016) also disclaimed the nomenclatural acts within the work for nomenclatural purposes as recommended by the Code (ICZN 1999) and has yet to publish his work officially so that the synonymies are listed here as new.

This is one of the large species of Ecuadorian Zammara with body lengths between 25 and 36 mm ( Goemans 2016). Specimens can be distinguished by the more than 50% infuscation of the apical fore wing and the sinusoidal median crossvein of the fore wing. The only other species to have a sinusoidal median crossvein is Z. smaragdina Walker, 1850 but the infuscation pattern is more spotted and less than half the apical area of the fore wing is infuscated in that species ( Goemans 2016).

DISTRIBUTION.—The species has been reported only from Ecuador ( Metcalf 1963a; Sanborn 2013; Goemans 2016). Goemans (2016) mapped the species over much of Ecuador, with a concentration of collection sites in the western portion of the country. Goemans (2016) also demonstrated the reported locality in Puerto Rico is erroneous. Goding (1925) reported the species from Guayaquil, many towns surrounding Guayaquil (as Z. intricata ), Milagro, and Tenegal (as O. brunneipennis ) in Guayas province, Naranjapata, Chimborazo province, Macas, Morona Santiago province (as O. pulla ), and Río Lliquino, Pastaza province.

MATERIAL EXAMINED.— “ Rio Palenque, ECUADOR / Los Rios Prov. - D. Dodson / XI – 15 –1972 ” two males and one female ( AFSC); “Ecuador // W. T. Davis / Collection” one male ( AFSC); “Playas / Ecuador / April1938 / Clark – McIntyre” three males and two females ( AFSC); “ G. Perez Franco, Col. / Guayaquil, Ecuador // ALLAN HANCOCK FOUNDATION / University of Southern / California No.” one male ( LACM), one male ( AFSC) .