Ledocas, OLIVIER, 1885

Bocakova, Milada, Campello-Gonçalves, Lucas & Silveira, Luiz Felipe Lima Da, 2022, Phylogeny of the new subfamily Cladodinae: neotenic fireflies from the Neotropics (Coleoptera: Lampyridae), Zoological Journal of the Linnean Society 195, pp. 1181-1199 : 1189

LEDOCAS OLIVIER, 1885

( FIGS 2D, 2J , 3B, C , 4B, C , 5B, C , 6A , 7A, B , 8A–D , 9A, B, 10B1–C2)

Ledocas Olivier, 1885: 136 .

Type species: Ledocas parallelus Olivier, 1885 (by monotypy).

Diagnosis: Pronotum and elytra subparallel-sided. Eyes small to medium-sized, mandibles semicircular, never right-angled, antennae uniflabellate, fan-like, branches of median antennomeres about 4× longer than the antennomere body, flabella of antennomere III markedly shorter than the following branches, translucent spots on pronotum absent. Abdomen strongly laterally lobed, sternum VIII bisinuate, mucronate, pygidium usually as long as wide, widest medially, with median extension about half as long as the lateral corners, lateral margins rounded. Male genitalia small, parameres as long as phallus, parameral apices simply straight.

Species included: Ledocas detectus Olivier, 1908 , L. emissus (Olivier, 1908) comb. nov., L. pikillactanus sp. nov., L.. parallelus Olivier, 1885 , L. remixtus (Olivier, 1911) comb. nov., L. xanthomus Olivier, 1894 .

Distribution: Argentina, Brazil, Peru, Venezuela, Paraguay.

Figures

Figure 2. Representatives of the Cladodinae; A, Cladodes flabellatus (photo: A. Reyes); B, Cladodes cincticollis (photo: V. Valdés); C, Cladodes ater (photo: M. Cid); D, Ledocas remixtus (photo: S. Quipildor-Sánchez); E, Nyctocrepis demoulini (photo: A. Madruga); F, Nyctocrepis stellata (photo:A.Katz); G, Dodacles elegans (photo: P. Smith); H, Brasilocladodes illigeri (photo: L. Silveira); I, Brasilocladodes delalandei (photo: A. Bouzan); J, Ledocas proteus (photo: M. Rojas-Quintero).

Figure 3. Habitus, dorsal view; A, Cladodes flabellatus; B, Ledocas parallelus; C, Ledocas pikillactanus sp. nov.; D, Nyctocrepis malleri; E, Andecladodes ovalis; F, Andecladodes cosangensis, lamella of antennomere III damaged; G, Brasilocladodes carinatus; H, Dryptelytra cayennensis; I, Dodacles elegans; J, Dodacles niger; K, Dodacles lateralis; L, Dryptelytra pampahermosae sp. nov.. Scales: 5 mm.

Figure 5. Antenna: A, Cladodes melanurus; B, Ledocas parallelus; C, Ledocas pikillactanus sp. nov.; D, Nyctocrepis malleri; E, Andecladodes ovalis; F, Andecladodes cosangensis; G, Dodacles elegans; H, Dryptelytra cayennensis; I, Dryptelytra pampahermosae sp. nov.; J, Vesta chevrolatii. Scales: 1 mm.

Figure 6. Head. A, ventral view; Ledocas pikillactanus sp. nov.; B, C, dorsal view: B, Andecladodes cosangensis; C, Andecladodes ovalis. Scales: 0.5 mm.

Figure 7. A, C–H, Abdomen, ventral view: A, Ledocas pikillactanus sp. nov.; C, Andecladodes cosangensis sp.nov.; D, Andecladodes ovalis; E, Nyctocrepis malleri; F, Dodacles niger; G, Dodacles lateralis; H, Dryptelytra pampahermosae sp.nov.; B, distal portion of hind tibia and tarsus, Ledocas pikillactanus sp. nov.

Figure 8. A, C, E, G, I, K, M, O, Q, S, U, W, Sternum VIII; B, D, F, H, J, L, N, P, R, T, V, X, Pygidium; A, B, Ledocas parallelus; C, D, Ledocas pikillactanus sp. nov.; E, F, Andecladodes cosangensis; G, H, Andecladodes ovalis; I, J, Dryptelytra cayennensis; K, L, Dryptelytra pampahermosae sp. nov.; M, N, Dodacles elegans; O, P, Dodacles niger; Q, R, Dodacles lateralis; S, T, Nyctocrepis malleri; U, V, Brasilocladodes illigeri; W, X, Brasilocladodes carinatus. Scales: 1 mm.

Figure4. Pronotum:A,Cladodescincticollis;B,Ledocasparallelus;C,Ledocaspikillactanus;D,Nyctocrepismalleri;E,Andecladodes ovalis;F,Andecladodes cosangensis, lamellaof antennomere IIIdamaged;G, Brasilocladodescarinatus; H,Dryptelytra cayennensis; I, Dodacles elegans; J, Dodacles niger; K, Dodacles lateralis; L, Dryptelytra pampahermosae sp. nov.. Scales: 1 mm.