Mesembrinella umbrosa Aldrich, 1922

Whitworth, Terry L. & Yusseff-Vanegas, Sohath, 2019, A revision of the genera and species of the Neotropical family Mesembrinellidae (Diptera: Oestroidea), Zootaxa 4659 (1), pp. 1-146 : 136-137

publication ID	https://doi.org/ 10.11646/zootaxa.4659.1.1
publication LSID	lsid:zoobank.org:pub:57309E14-0330-4ED7-BCDA-355EE6618215
persistent identifier	https://treatment.plazi.org/id/03DA87E6-0FBD-FF5D-FF19-B84A35FFFC78
treatment provided by	Plazi
scientific name	Mesembrinella umbrosa Aldrich, 1922
status

Treatment

( Figs 95–96 View FIGURES 93–98 , 185–186 View FIGURES 183–192 , 235 View FIGURES 229–235 , 278 View FIGURES 266–279 , 322 View FIGURES 318–323 , 366 View FIGURES 350–369 , 407 View FIGURES 401–410 , 451 View FIGURES 439–453 , 484 View FIGURES 484–487 , 488 View FIGURE 488 )

Mesembrinella umbrosa Aldrich, 1922: 12 . Holotype male (USNM), examined photographically. Type locality: Tucurrique , Costa Rica.

Mesembrinella umbrosa: Peris & Mariluis (1984: 259) ; Kosmann et al. (2013: 78); Wolff 2013: 121; Wolff & Kosmann (2016: 870) View Cited Treatment ; Marinho et al. (2017: tab. 1); Velásquez et al. (2017: 109): Cerretti et al. (2017: tab. 2).

Diagnosis. A large fly averaging 14.2 mm (13–15/5) in length. Thorax reddish with bright bluish-purple abdomen; wing yellowish, anterior margin from costa to vein R 2+3 darker yellow; section IV of wing 0.69 of section III; scutellum without discal setae; subcostal sclerite setose. Epandrium, cerci and surstyli as in Figs 95–96 View FIGURES 93–98 . Female terminalia as in Fig. 322 View FIGURES 318–323 .

Redescription. Male. Head. Frons 0.025 (0.02–0.03/5) of head width at narrowest. Fronto-orbital and parafacial yellow with whitish tomentum; frontal setae ascending about 60% of distance to vertex; frontal vitta dark red, obliterated midway; gena dark orange with pale tomentum and a horizontal row of stout, black setae on ventral margin, otherwise with scattered tan setae; postgena orange with silvery tomentum and fine golden setae; occiput with dark cuticle covered with heavy silvery tomentum and fine golden setae, tomentum absent along upper margin and midway; median occipital sclerite shiny dark orange; antenna orange, first flagellomere with heavy pale tomentum; palpus typical; eye with median facets 4x size of lateral facets; ocellar triangle medium-sized, anterior ocellus 2x size of posterior ocelli; supravibrissal setae short, brown, forming a dense patch ascending 1/4 of distance to antennal base.

Thorax. Dorsum orange with streaky stripes of irregular, whitish tomentum; pleura orange with pale tomentum; chaetotaxy: ac 2:2 or 2:3, dc 2:3, ia 1, ph 1, ppn 3x 3, kat 2:1, meron with very fine setae in an inverted L, 1 slightly convergent ap, sa and lat absent, 1 stout bas, 1 pb, 0 disc; subscutellum prominently developed; spiracles large, orange; legs entirely orange. Wing entirely yellow, anterior edge from costa to R 2+3 darker, middle of cell r 4+5 hyaline; subcostal sclerite setose; basicosta and tegula orange; section IV 0.69 of section III; discs of upper and lower calypters reddish; rim of upper calypter dark with short reddish setae, rim of lower calypter pale with long reddish setae.

Abdomen. T1+2 yellowish with whitish tomentum, T3–5 more or less metallic, gleaming blue; T4 with row of marginal setae, T5 with row of short marginal setae; disc of T5 with dense, short, fine setae only. Terminalia in lateral view with surstylus curved backward, cercus with apical hook ( Fig. 95 View FIGURES 93–98 ); in posterior view, cerci broad basally and gradually narrowing to tips ( Fig. 96 View FIGURES 93–98 ); phallus in lateral view as in Fig. 185 View FIGURES 183–192 ; in dorsal view, hypophallic lobes rectangular with moderate serrations ( Fig. 186 View FIGURES 183–192 ); T6, STS7+8, pre- and postgonite, ejaculatory sclerite, ST6 and hypandrium as in Fig. 235 View FIGURES 229–235 ; ST1–5 broad, as in Fig. 278 View FIGURES 266–279 .

Female. Similar to male except frons 0.263 (0.24–0.28) of head width at narrowest. T6 of WV shape; T7 continuous, thin, narrowed midway; T8 as separate sclerites ( Fig. 322 View FIGURES 318–323 ); ST6–8 and hypoproct as in Fig. 366 View FIGURES 350–369 ; spermathecae filiform ( Fig. 407 View FIGURES 401–410 ); ST1–5 as in Fig. 451 View FIGURES 439–453 .

Type material examined. HOLOTYPE ♂ ( Costa Rica, Cartago; USNM; examined photographically: Fig. 484 View FIGURES 484–487 ), labeled: Tucurrique / Costa Rica; CollSchild / & Burgdorf; Mesembrinella / umbrosa / Ald.; Type No. / 25246 / U.S. N.M. [orange label]; USNMENT / 01295435.

Additional material examined. Costa Rica, Alajuela. 1 ♀ *, 1 ♀, San Ramón, Villa Blauca , 1115 m, 18.xi.2008, Tp. Frutas, L _N_242482 483371 #95287, R. Rojas Vasquez ( INBIO) . Cartago. 1 ♂, Grano de Oro, 1120 m, Chir- ripo, Turrialba , xi.1992, L_N_ 200250 595900, P. Campos ( INBIO) ; 1 ♂, La Suiza de Turrialba , 9.x.1921, P. Schild ( USNM) ; 1 ♀, P.N. Tapanti, 1150 m, vii.1993, L_N_194000 559800 #2801, G. Mora ( INBIO) ; 1 ♀ ♦ ( TLW338 ) , P.N. Barbilla, Turrialba, Tayutic, Campamento 2, 1200 m, 6–9.v.2005, Libre, L_N_213371 600782 #80273, J.D. Gutierrez ( INBIO) . Guanacaste. 1 ♂, Macizo Miravalles, Estación Cabro Muco., 1100 m, 27.vi–2.vii.2003, L_N_ 299769 411243 #74526, Tp de Luz , B. Hernández ( INBIO) ; 1 ♂, same data except 24.ix–5.x.2003, Libre, #75498 ( INBIO) . Puntarenas. 2 ♂♂*, 2 ♂♂, Monteverde , 25–30.viii.1991, 1500 m, D.M. Wood ( CNC) ; 1 ♂, same data ex- cept 18–24.viii.1987 ( INBIO) ; 1 ♂, same data except San Luis , 1040 m, 24.viii–15.ix.1992, L_N_ 250850 449250, F.A. Quesada ( INBIO) ; 1 ♀, same data except 14–20.v.1990, D.M. Wood ( CNC) ; 1 ♀ ♦ ( TLW370 ) , same data except 20–22.viii.1993 ( INBIO) ; 1 ♂*, Est. Pittier , PILA-ACLA, 1670 m, 5–18.1995, L_N_330900 577400 #4437, R. Vil- lalobos ( INBIO) ; 1 ♂, same data except 4–22.i.1996, L_S_ 220900 577400 #6813, E. Navarro ( INBIO) ; 1 ♂, Send. a Cerro Pittier , 600 m, NO. de la Estación, 1750 m, 18–24.xii.1995, L_S_331250 577150 #7401, on excrement, M. Moraga ( INBIO) ; 1 ♂, Estación Agujas. Rio Agujas. Send. Zamia. , 300 m, 14–24.viii.1996, L_S_276750 526550 #8477, A. Azofeifa ( INBIO) ; 1 ♀, Est. La Casona, Res. Biológica, Monteverde , 1520 m, x.1990, L_N_253250 449250, N. Obando ( INBIO) . San José. 1 ♀ *, Zurquí de Moravia , 1600 m, 21.x.2012, #105240, Bait trap with hu- man dung, ZADBI ( LACM) ; 1 ♀ ♦ ( TLW342 ) , same data except 24.ix.2012, #105110 ( LACM) ; 1 ♀ ♦ ( TLW341 ) , same data except 6–12.ix.2012, #105003, Malaise trap #1 ( LACM) ; 1 ♀ ♦ ( TLW340 ) , same data except 22.x– 1.xi.2012, #105314, Malaise trap #1 pan ( LACM) ; 1 ♀, 5400 ft., HWY 2, N of San Isidro , 6.vi.1988, [no collector] ( LACM) . Ecuador, Pastaza. 1 ♀, Napa Sarayacu , 29.x.2015, 1312 m, M. Domínguez ( FSCA) .

Distribution. Costa Rica, Ecuador. Marinho et al. (2017) also listed it from Colombia and Panama.

Remarks. Five specimens ( TLW 338, TLW 340–342 and TLW 370) were barcoded, and grouped together close to M. apollinaris ( Fig. 488 View FIGURE 488 ).

USNM	Smithsonian Institution, National Museum of Natural History
CNC	Canadian National Collection of Insects, Arachnids, and Nematodes
LACM	Natural History Museum of Los Angeles County
FSCA	Florida State Collection of Arthropods, The Museum of Entomology

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Taxonomy

Kingdom	Animalia
Phylum	Arthropoda
Class	Insecta
Order	Diptera
Family	Calliphoridae
SubFamily	Mesembrinellinae
Genus	Mesembrinella