Chilelimnophila (Chilelimnophila) wangi Krzemiński, Kania-Kłosok et Arillo, 2024

Krzemiński, Wiesław, Kania-Kłosok, Iwona, Arillo, Antonio, Kopeć, Katarzyna, Santos, Daubian & Soszyńska, Agnieszka, 2024, The evolutionary history and biogeographical distribution of the Mesozoic relic genus Chilelimnophila (Diptera, Limoniidae), Zoological Journal of the Linnean Society 202 (1), pp. 1-16 : 4-7

Chilelimnophila (Chilelimnophila) wangi Krzemiński, Kania-Kłosok et Arillo , sp. nov.

( Fig. 3A–E )

Zoobank registration: urn:lsid:zoobank.org:act : 665347C3- 157F-4E27-88DE-4B2BAB670023

Diagnosis: Vein R 1 elongate, tip of R 1 positioned far beyond fork of R 2 + 3+4; sc-r situated before tip of Sc, approximately nine times the distance from the tip of Sc; outer gonostylus narrow, elongate, slender, dark at the tip; inner gonostylus short and massive, reaching approximately half the length of gonocoxite, slightly curved to the internal part of hypopygium, sharpened at apex.

Material examined: Holotype No. MP/4449 (male), coll. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Kraków, Poland (ISEA PAS). Specimen well preserved, with partially destroyed legs and wing.

Horizon and locality: Lowermost Cenomanian, Hukawng Valley , northern Myanmar. The mining is done at a hill named Noije Bum, near Tanai Village (26 ° 21′ 33.41′′ N, 96 ° 43′ 11.88′′ E).

Description: Body ( Fig. 3D ) 2.69 mm long, brown, wings without colour pattern, pterostigma absent.

Head: Antenna ( Fig. 3A ) 0.85 mm long, longer than head, longer than palpus; scape cylindrical, rather short, only slightly longer than pedicel; approximately 2× as long as wide, pedicel ovoid, constricted at base, scape approximately 1.5× as long as pedicel; first flagellomere fused with second and third; flagellomeres 5‒14 elongate, tiny, approximately 4–5× as long as wide, become more slender to the apex of antenna, first flagellomere longer and wider than second one, second and third flagellomeres very short, fused with the first one; last flagellomere shorter than penultimate one; each flagellomeres with three to five elongate setae, longer than segments bearing them; maxillary palpus foursegmented ( Fig. 3B ), short, 0.2× the length of antenna, 0.21 mm long, palpomeres rather ovoid, short, only slightly longer than wide, third palpomere comparable length to the second, all palpomeres slightly widened in midlength; all palpomeres with few setae, shorter than segments bearing them.

Thorax: Almost as long as high ( Fig. 3D ); wing 2.94 mm long, 0.84mm wide ( Fig. 3E ); h vein situated between the base of wing and fork of Mb and Cu; Sc ending on C at the level of bifurcation of Rs and tip of A 1; sc-r situated at approximately eight of its length from the tip of Sc; Rs slightly curved, originating well proximally to the level of the tip of A 2, only slightly longer than R 2 + 3+4, R 1 reaches almost half the length of petiole; fork of R 2 + 3+4 situated just beyond the level of d-cell, R 3 approximately 0.6× the length of R 4; R 4 longer than R 2 + 3+4, longer than Rs; section of R 5 curved between its origin and point of contact with crossvein r-m; vein M 3 almost straight, approximately 1.5× the length of d-cell; d-cell 3× as long as wide; crossvein m-cu shorter than d-cell; A 1 slightly curved at the tip, tip of A 1 situated beyond fork of Mb; A 2 slightly curved; tip of A 2 just before the level of m-cu, just before the level of fork of Mb, beyond the level of fork of Rb; haltere: stem narrow, elongate, slightly longer than knob; tibial spurs rather thick, not very elongate, shorter than setae which occur on tibia.

Abdomen: Male terminalia ( Fig. 3E )—hypopygium 0.55 mm long, elongate and narrow, gonocoxite approximately 4× as long as wide, tip of gonocoxite and outer gonostylus slightly sclerotized, brown, outer gonostylus narrow, pointed, slightly curved to internal part of hypopygium, inner gonostylus rather slender but lobe-shaped, very pale, rounded at apex.

Etymology: The specific name is dedicated to eminent palaeontologist Professor Bo Wang from the Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, China.

Figures

Figure 3. Chilelimnophila (C.) wangi sp. nov. (Limoniidae), inclusions in Kachin amber, holotype No. MP/4449 (male). A, antenna. B, palpus. C, part of hypopygium: gonocoxite and gonostyle. D, body, lateral view. E, wing venation.Abbreviations:gx, gonocoxite; ing, inner gonostylus; oug, outer gonostylus; scp, scape; ped, pedicel; I–IV, palpomeres 1–4.