Chilelimnophila (Ribeironia) amorimi Krzemiński, Kania-Kłosok et Arillo,, 2024

Krzemiński, Wiesław, Kania-Kłosok, Iwona, Arillo, Antonio, Kopeć, Katarzyna, Santos, Daubian & Soszyńska, Agnieszka, 2024, The evolutionary history and biogeographical distribution of the Mesozoic relic genus Chilelimnophila (Diptera, Limoniidae), Zoological Journal of the Linnean Society 202 (1), pp. 1-16 : 3-4

Chilelimnophila (Ribeironia) amorimi Krzemiński, Kania-Kłosok et Arillo, sp. nov.

( Fig. 2A–F )

Zoobank registration: urn:lsid:zoobank.org:act:7C7A1D3BFE45-4FF5-BC5E-ADF0518F3A05

Diagnosis: Vein R 1 elongate, tip of R 1 positioned far beyond fork of R 2 + 3+4; sc-r situated before tip of Sc, approximately five times the distance from the tip of Sc; outer gonostylus narrow, elongate, strongly sclerotized, pointed, slightly curved to the internal part of hypopygium; inner gonostylus longer than half the length of the outer one.

Material examined: Holotype No. MCNA 9845 (male) housed in in the Museo de Ciencias Naturales de Álava, (Vitoria-Gasteiz, Álava, Spain). Specimen well preserved, with partially destroyed legs and left antenna.

Horizon and locality: Lower Cretaceous (Upper Albian); Peñacerrada I amber site ( Peñacerrada I , Moraza), eastern area of the Basque-Cantabrian Basin, Burgos, northern Spain.

Description: Body ( Fig. 2A, B ) 1.80 mm long, dark brown, wings without colour pattern, pterostigma absent.

Head: Antenna ( Fig. 2C ) 0.50 mm long, longer than head, longer than palpus; scape cylindrical, very elongate, approximately 4× as long as wide, pedicel ovoid, constricted at base, scape approximately twice as long as pedicel; first flagellomere ovoid, constricted at base; first flagellomere ovoid; flagellomeres 2‒11 elongate, approximately twice as long as wide, becoming more slender to the apex of antenna, first flagellomere only slightly longer and wider than second one, first flagellomere partially fused with the second one, last flagellomere equal in length to penultimate one; last flagellomere with two, rather short setae at apex, each other flagellomeres with four not very elongate setae, shorter than segments bearing them; maxillary palpus foursegmented, rather short, 0.3× the length of antenna, 0.20 mm long, palpomeres more or less cylindrical, not very elongate, longer than wide, rather narrow, third palpomere longer than second and fourth, slightly widened in midlength, second and fourth palpomere 2× as long as wide, third palpomere approximately 2.5× as long as wide; all palpomeres ( Fig. 2D ) with few not very elongate setae, shorter than segments bearing them.

Thorax: Almost as long as high; wing 2.20 mm long, 0.80 mm wide ( Fig. 2F ); h vein situated between the base of wing and fork of Mb and Cu; Sc ending on C before the level of bifurcation of Rs and tip of A 1; sc-r situated at approximately six of its length from the tip of Sc; Rs slightly curved, originating well proximally to the level of the tip of A 2, only slightly longer than R 2 + 3+4, R 1 reaches 0.3 the level of petiole, fork of R 2 + 3+4 situated just beyond the level of d-cell, R 3 approximately 0.6× the length of R 4; R 4 longer than R 2 + 3+4, longer than Rs; section of R 5 between its origin and point of contact with crossvein r-m straight; vein M 3 almost straight, approximately 1.5× the length of d-cell; d-cell 2× as long as wide; crossvein m-cu shorter than d-cell; A 1 almost straight, tip of A 1 situated beyond fork of Mb; A 2 slightly curved; tip of A 2 just before the level of m-cu; tip of A 2 situated just before the level of fork of Mb, beyond the level of fork of Rb; haltere: stem narrow, elongate, slightly longer than knob; tibial spurs rather thick, not very elongate, shorter than setae which occur on tibia.

Abdomen: Male terminalia ( Fig. 2E )—hypopygium 0.20 mm long, elongate and narrow, gonocoxite approximately 3× as long as wide, tip of gonocoxite and outer gonostylus strongly sclerotized, almost black, outer gonostylus narrow, pointed, slightly curved to internal part of hypopygium, outer gonostylus rather slender but lobe-shaped, very pale, rounded at apex.

Etymology: The specific name is dedicated to eminent entomologist and paleontologist, Professor Dalton de Souza Amorim from the Departamento de Biologia (FFCLRP), Universidade de Sao Paulo, Brazil.

Remarks: The antenna of C. (R.) amorimi subgen. et sp. nov. is reduced and 11-segmented, while other species of Chilelimnophila are characterized by 16-segmented antenna.

Figures

Figure 2. Chilelimnophila (R.) amorimi subgen. et sp. nov. (Limoniidae), inclusions in Spanish amber, holotype No. MCNA 9845 (male). A, body, latero-frontal view, drawing A. Arillo; B, body, latero-frontal view. C, antenna.D, palpus. E, part of hypopygium: gonocoxite and gonostyle; F.wing venation. Abbreviations: gx, gonocoxite; ing, inner gonostylus; oug, outer gonostylus; scp, scape; ped, pedicel; I–IV, palpomeres 1–4.