Muisca, Spinola, 1844

Opitz, Pitz Pinola Weston, 2018, Classification, Natural History, and Evolution of the Subfamily Peloniinae O (Coleoptera: Cleroidea: Cleridae). Part IX. Taxonomic revision of the New World genus Muisca S, Linzer biologische Beiträge 50 (1), pp. 587-653 : 596-598

Key to species of Muisca SPINOLA

My efforts in the production of this key strive to use convenient characteristics of external morphology. But, the distinction between closely related species is sometimes small and when possible the reader should in most cases rely in the shape of the phallus for definitive species identifications.

1 Pronotum with secondary tubercle (Fig. 47) (M. insigna- group) ..................................2

1' Pronotum without secondary tubercle ...........................................................................3

2(1) Elytral asetiferous punctures terminate at elytral middle ( Fig. 125) ( Bolivia, Brazil, Paraguay) ........................................................................ Muisca dilatata (CHEVROLAT)

2' Distribution of elytral asetiferous punctures extend beyond elytral middle ( Brazil) ( Fig. 126) ......................................................................... Muisca insigna (CHEVROLAT)

3(1') Body form ovate ( Fig. 127)...........................................................................................4

3' Body form more rectangulate ( Fig. 139).......................................................................9

4(3) Antennal capitulum extraordinarily long ( Fig. 15) (M. apicalis- group)........................5

4' Antennal capitulum not extraordinarily long ................................................................6

5(4) White marking at elytral apex sharply defined ( Fig. 127) ( Bolivia, Brazil).................... ............................................................................................. Muisca apicalis (SPINOLA)

5' White marking at elytral apex not sharply defined ( Fig. 128) ( Bolivia, Brazil).............. ........................................................................................ Muisca dozieri OPITZ, nov.sp.

6(4') Elytral disc vested with extraordinarily stout setae; hind body subovoid ( Fig. 129) ( Brazil) (M. irrorata- group) ( Brazil, Bolivia)..................... Muisca irrorata (GORHAM)

6' Elytral disc not vested with extraordinarily stout setae; hind body not subovoid ( M. hirtula group)................................................................................................................7

7(6') Elytral disc without spots, disc partially castaneous (M. xanthura- group) ( Bolivia, Brazil, French Guiana) ( Fig. 105) ................................ Muisca xanthura OPITZ, nov.sp.

7'. Elytral disc with spots, disc not partially castaneous ....................................................8

8(7') Elytral dark spot in form of "checkmark" ( Brazil) ( Fig. 130) .... Muisca hirtula (KLUG)

8' Elytral dark spot not in form of "checkmark" ( Brazil) ( Fig. 131) ................................... .......................................................................................... Muisca togata (CHEVROLAT)

9(3') Eyes very small; frons twice width of eyes (M. angulicollis- group) ...........................10

9' Eyes not very small; frons not twice width of eyes.....................................................11

10(9) Pronotal disc impunctate at midline ( Colombia) ( Fig. 136) ............................................ ......................................................................................... Muisca menda OPITZ, nov.sp.

10' Pronotal disc punctate at midline ( Colombia) ( Fig. 135) ................................................ .................................................................................. Muisca angulicollis (CHEVROLAT)

11(9') Elytral disc with very few asetiferous punctures, broad region near sutural margin impunctate (M. fera- group) ........................................................................................12

11' Elytral disc asetiferous punctures not reduced in numbers as described above...........13

12(11) Pronotum with dark brown lines at sides ( Peru) ( Fig. 134)............................................. ........................................................................................... Muisca signa OPITZ, nov.sp.

12' Pronotum without dark brown lines at sides ( Costa Rica) ( Fig. 133).............................. ................................................................................................. Muisca fera (WOLCOTT)

13(11') Pronotal disc cribrate (M. maculosa- group)................................................................14

13' Pronotal disc not cribrate ............................................................................................15

14(13) Pronotal disc with 2 black markings ( Brazil) ( Fig. 138) ........ Muisca maculosa (GORHAM)

14' Pronotal disc uniformly castaneous ( Brazil, Peru) ( Fig. 137) ......................................... ............................................................................................ Muisca hexa OPITZ, nov.sp.

15(13') Posterior margin of male pygidium broadly concave (M. mestolinea- group) .............16

15' Posterior margin of male pygidium not broadly concave............................................18

16(15) Pronotum without dark brown markings; each elytral disc with 3 diagonally positioned faint markings ( Colombia) ( Fig. 139) ........... Muisca anachyma OPITZ, nov.sp.

16' Pronotum with dark brown markings ..........................................................................17

17(16') Pronotum with 2 dark lines at sides ( Ecuador) ( Fig. 140) ............................................... .................................................................................. Muisca mestolinea OPITZ, nov.sp.

17' Pronotum with 2 dark spots ( Colombia, Ecuador) ( Fig. 141) ......................................... .......................................................................................... Muisca omma OPITZ, nov.sp.

18(15') Antennal funicle longer than length of capitulum (M. octonotata- group)...................19

18' Antennal funicle not longer than length of capitulum .................................................23

19(18) Elytral disc with dark fascia ........................................................................................20

19' Elytral disc with spots .................................................................................................21

20(19) Midelytral fascia boldly expressed ( Costa Rica, Panamá) ( Fig. 142).............................. ..................................................................................... Muisca heppneri OPITZ, nov.sp.

20' Midelytral fascia faintly expressed ( Colombia) ( Fig. 143).............................................. ............................................................................. Muisca lateripunctata (SCHENKLING)

21(19') Each elytral disc with 6 spots ( Colombia) ( Fig. 144) ...................................................... .................................................................................. Muisca magdalena OPITZ, nov.sp.

21' Each elytral disc with 4 spots......................................................................................22

22(21') Elytral asetiferous punctures end at about discal ½ ( Costa Rica, Panamá) ( Fig. 145)..... ........................................................................................ Muisca octonotata (GORHAM)

22' Distribution of elytral punctures extend well beyond discal ½ ( Colombia) ( Fig. 146) ........................................................................... Muisca tetraspilota (CHEVROLAT)

23(18') Pronotum black ( Bolivia, Peru) ( Fig. 153)................................ Muisca peruviana (PIC)

23' Pronotum not black ....................................................................................................24

24(23') Pronotum with dark markings .....................................................................................25

24' Pronotum without dark markings................................................................................27

25 Each elytral disc with 7 light brown spots ( Peru) ( Fig. 151)........................................... ..................................................................................... Muisca malakela OPITZ, nov.sp.

25' Elytral color not as above............................................................................................26

26(25') Antennal capitulum much longer than length of funicle ( Bolivia) ( Fig. 154) ................. ......................................................................................... Muisca sigilla OPITZ, nov.sp.

26' Antennal capitulum not longer than length of funicle ( Venezuela, Ecuador) ( Fig. 149) .............................................................................. Muisca biordinis OPITZ, nov.sp.

27(24') Antennal capitulum black ...........................................................................................28

27'. Antennal capitulum not black .....................................................................................30

28(27) Seventh and 8 th antennomeres not infuscated ( Brazil) ( Fig. 155).................................... ................................................................................................. Muisca testacea (KLUG)

28' Seventh and 8 th antennomeres infuscated....................................................................29

29(28') Elytral length/width ratio 1.89 ( Brazil) ( Fig. 147) ...... Muisca adamanta OPITZ, nov.sp.

29' Elytral length/width ratio 1.61 ( Brazil) ( Fig. 156) ............. Muisca variabilis (SPINOLA)

30(27') Short narrow black line projecting from elytral anterior margin.................................31

30' Elytral color not as above............................................................................................32

31(30) Middle of elytral disc with irregular-shaped fascia ( Colombia, Venezuela) ( Fig. 152) ............................................................................... Muisca nigrosignata (SPINOLA)

31' Midelytral fascia fragmented ( Colombia, Venezuela) ( Fig. 148).................................... ........................................................................................... Muisca agma OPITZ, nov.sp.

32(30') Elytral disc traversed by 3 black fasciae ( Bolivia) ( Fig. 157) Muisca zona OPITZ, nov.sp.

32' Elytra not traversed by fascia ( Panamá, Colombia, Venezuela) ( Fig. 150)..................... .......................................................................................... Muisca bitaeniata (SPINOLA)