Hipparidium Jeannel, 1946

Serrano, Artur R. M. & Capela, Rúben A., 2013, The tiger beetles (Coleoptera: Carabidae, Cicindelinae) of Angola: a descriptive catalogue and designation of neotypes, Zootaxa 3731 (4), pp. 401-444 : 427-434

Hipparidium Jeannel, 1946 (Faune Emp. Fr. 11: 151)

Ten known species for continental Africa are included in this genus. There is no revisionary work on the African representatives of the genus. Rivalier (1948) included this taxon in a study of the species of “French Tropical Africa”. Only a single species is known to occur in Angola so far.

Hipparidium interruptum (Fabricius, 1775) (Syst. Ent.: 225)

Distribution in Angola

Old records:

—Quango (LUNDA NORTE) (= Cuango) (117) (Quedenfeldt 1883 sub Cicindela interstincta Schönherr ). —Dundo (LUNDA NORTE) (51), Luachimo Forest (= Dundo, R. Luachimo) (LUNDA NORTE) (51), Tshikapa River (50 km SW Dundo) (=Chingufo) (LUNDA NORTE) (68) (Basilwesky 1955; Ferreira 1965). Material examined: Dundo (LUNDA NORTE) (51), 2.V.1946, 1♂, Companhia Diamantes de Angola leg., reg.nº 704, IICT.

Remarks. A tiger beetle species with a wide distribution throughout western and central Africa (Werner 2000b; Serrano 2007). As for other species recorded for Angola, Werner (2000b) did not mention it for this country. It occupies the gallery forests of the rivers and in the moist shaded paths inside the forests (Serrano 2007, 2012).

Calomera Motschulsky, 1862 (Études Entomologiques 11: 22)

This genus (type species: Cicindela decemguttata Fabricius, 1801 ), was formerly included in the entomological literature under the junior synonym Lophyridia Jeannel, 1946 (type species: Cicindela dongalensis Klug, 1832 ), but in fact Calomera has priority (Lorenz 2005). The genus Calomera is wide-spread throughout Oriental, Palaearctic and Afrotropical Regions. There is no revisionary work on the African representatives of the genus.

Calomera asperula (Dufour, 1821) (Ann. Genn. Sc. Phys. Brux. 8: 359, Pl. 130: fig.1)

Distribution in Angola

Old records:

—Quango (= Cuango) (LUNDA NORTE) (117) (Quedenfeldt 1883 sub Cicindela asperula ). —Kuango (Mechow) (= Cuango) (LUNDA NORTE) (117) (Wellman & Horn 1908 sub Cicindela asperula ). — Angola (Ferreira 1965 sub Cicindela asperula ).

Remarks. A species known from some western countries of Africa ( Senegal, Ghana, Nigeria) and West- Central African countries ( Central African Republic, D. R. of the Congo and Angola) (Werner 2000b). It is found on the sandy or stony banks and shores of rivers and lakes.

Calomera aulica aulica (Dejean, 1831) (Spec. Col. 5: 250)

Distribution in Angola

Old records:

— Angola (without locality) (Putzeys 1880; Wellman & Horn 1908; Ferreira 1965 all sub Cicindela aulica ). —Loanda (= Luanda) ( LUANDA) (89) (Wellman & Horn 1908 sub Cicindela aulica ).

Remarks. A polymorphic tiger beetle species that is the nominate basis for the description of five subspecies (see Werner 2000b). The nominal population is wide-spread throughout the Palaearctic Region, but it is found also in several African countries. It occupies saline habitats like sea shores and mangroves (Serrano 2007). The presence of C. aulica in Angola is paradoxical as the nearest African country from which it is known is in Guinea-Bissau (Werner 2000b; Serrano op. cit.). Werner (op. cit.) did not include C. aulica for Angola, except for a commentary by Wellman & Horn (1908). They reported on this species saying “an intermediate form between the typical aulica and the subsp. polysita Guér. occurs in Loanda”.

Calomera fimbriata imperatrix (Srnka, 1891) (Deutsch. Ent. Zeitschr.: 13)

Distribution in Angola

Old records:

—Humbi (=Humbe) (border Angola / Namíbia) (CUNENE) (422) (Horn 1935; Monard et al. 1956; Ferreira 1965

all sub Cicindela dongalensis imperatrix ).

Remarks. A form wide-spread throughout West-Central ( Angola, Namibia, southern D. R. of the Congo, Burundi, Rwanda) to East African countries ( Mozambique, Malawi, Tanzania, Kenya, Somalia) (Werner 2000b). It is found on the sandy or stony banks and shores of rivers and lakes. It is likely it is more wide spread in Angola than presently known.

Lophyra Motschulsky, 1859 (Etud. entom. 8: 25)

A genus widespread throughout the Oriental, Palaearctic and Afrotropical Regions. Most of its subgenera and species occur in Africa, which is most likely the centre of origin of the genus. The Afrotropical species of this genus were revised by Rivalier (1958), but with the subsequent description of more subgenera and species/ subspecies a new revision is needed.

Lophyra (s. str.) differens (W. Horn, 1892) (Deutsch. Ent. Zeitschr.: 82)

Distribution in Angola.

Localities reported: 43 km W Quipungo (HUILA) (337) (P. Schüle pers. comm.).

Remarks. A new record to the tiger beetles fauna of Angola, Lophyra differens is widespread throughout Namibia, South Africa, Zimbabwe, Mozambique, Malawi, Tanzania, Kenya and Somalia (Werner, 2000b). The adults occur on river banks or riverbeds with white or yellow sand (Werner op. cit.). Peter Schüle (pers. comm.) found it on the moist edges of a riverine.

Lophyra (s. str.) endroedyi Cassola, 1993 [Rev. Zool. Afr. 107 (4): 358]

Distribution in Angola

Old records:

—Mossamedes (Anchieta) (=Namibe) (NAMIBE) (353) (Putzeys 1880 sub Cicindela senegalensis Dejean , see Wellman & Horn 1908).

—Mossamedes (=Namibe) (NAMIBE) (353) (Wellman & Horn 1908 sub Cicindela brevicollis clathrata , see also Cassola 1993, p. 360).

—Moçâmedes (=Namibe) (NAMIBE) (353) (Ferreira 1965 sub Cicindela clathrata ).

—Moçamedes (=Namibe) (NAMIBE) (353), west of Cubal (=Bero river) (NAMIBE) (353) (Cassola 1993). New localities reported: Cassoço (= Cassoco) (KWANZA SUL) (185) (Th. Bouyer pers. comm.) Material examined: Mossamedes (=Namibe) (NAMIBE) (353), Anchieta leg., 1♀, reg.nº 1720, “Old

Collection” MZUC.

Remarks. A tiger beetle species restricted to south-western Angola and Namibia (Cassola 1993; Werner

2000b). The female here examined is considerably damaged (lack of antennae and intermediate and hind legs).

This specimen could be ascribed as well to Lophyra damara (Péringuey) by the elytral markings (anterior band

connected largely with the second juaxtasutural elongate spot). The elytral drawing is similar to the figures 301,

301.2 presented in Werner (2000b) or between the fourth and fifth stages of figure 7 given by Cassola (1975a). A

male specimen could help to clarify this judgement. Interestingly Wellman & Horn (1908) under Cicindela

brevicollis neglecta Dejean point out also that “occurs near Mossamedes an intermediate form between this last and

the var. damara Pér. ” (see also Cassola 1993, p. 360). Lophyra endroedyi occurs in dry riverbeds, a type of habitat

very common in the Namibe region also. The presence of this species in Cassoço locality increases its distribution

far to the north of Angola.

Lophyra (s. str.) herero (Péringuey, 1892) (Trans. S. Afr. Phil. Soc. 7: 34)

Distribution in Angola

Old records:

— Angola (without locality) (Werner 2000b).

Remarks. A species common all over Namibia and only included for Angola on the basis of Werner (2000b) who included no locality. Cassola (1975b) who reviewed this species included only Namibia as its distribution. The species occurs on the shores of riverbeds and lakes shores with light sand. The species remains hypothetical for Angola until confirmed by a specimen.

Lophyra (s. str.) neglecta neglecta (Dejean, 1825) (Spec. Col. I: 114)

Distribution in Angola

Old records:

— Angola (Welwitsch) (Putzeys 1880 sub Cicindela pudica Boheman and C. senegalensis Dejean ). —Malange (=Malanje) (Mechow leg.) (MALANJE) (131) (Quedenfeldt 1883 sub Cicindela neglecta ).

—Malange (=Malanje) (Mechow leg.) (MALANJE) (131), Angola (Welwitsch) (Wellman & Horn 1908 sub

Cicindela brevicollis neglecta ).

—Kuango (=Cuango) (LUNDA NORTE) (117) (Wellman & Horn 1908 sub Cicindela brevicollis discoidalis

Dejean).

Material examined: Angola, Welwitsch leg., 1♂, 1♀, regº nº 1907, “Old Collection” MZUC; Angola, 1♂, regº nº 1907, “Old collection” MZUC; Tando Nzinze ( CABINDA ) (5), R. Capela leg., 8.II.2011, 6 ♂, 1♀, ASC; Tando Pala (=N` Pala) ( CABINDA ) (5), R. Capela leg., 11.II.2011, 2 ♂, 6 ♀, ASC; Tando Pala (Nwgo Caio= Caio) ( CABINDA ) (5), R. Capela leg., 12.II.2011, 2 ♂, ASC; Sassa Zau ( CABINDA ) (5), R. Capela leg., 1.III.2011, 4 ♂, 2 ♀, ASC; Camama ( Luanda) ( LUANDA (89), R. Capela leg., 18.IV.2013, 7 ♂, 3 ♀, ASC.

Remarks. A polymorphic tiger beetle species common in West and Central Africa (Werner 2000b), it can be found in sandy habitats near the edges of rivers, lakes and ponds or even on paths through fields or along the road edges (e.g., Cabinda , R. Capela pers. comm.).

Lophyra (s. str.) neglecta intermediola (W. Horn, 1921) (Ent. Blätt. 17: 175)

Distribution in Angola

Old records:

—Quanza (=Cuanza, Barra do) ( LUANDA) (107), Luanda ( LUANDA) (107) (Wellman & Horn 1908 sub Cicindela brevicollis intermedia Klug ).

Remarks. A subspecies more common and widespread throughout Central-southern and eastern Africa

(Werner 2000b). It occurs in the same type of habitats as the nominal form.

Lophyra (s. str.) obliquograciliaenea (W. Horn, 1920) [Ark. Zool. 13 (11)]

Distribution in Angola

Old records:

—Dundo (LUNDA NORTE) (51) [Alves 1956 sub Cicindela (Lophyra) brevicollis obliquograciliaenea ].

Material examined: Dundo (LUNDA NORTE) (51), Companhia dos Diamantes leg., 2.V.1946, 1 ♀, reg. nº 705, IICT,?. II.1949, 3 ♂, 4 ♀, reg. nº 2147, IICT; Lago Dilolo (MOXICO) (223), Estudos Apícolas Ultramar leg., 29.IX.1959, 1 ♀, reg nº 3077, IICT; Chitato (Dundo) (LUNDA NORTE) (51), L. Mendes leg., 26.IX–7. XI.1995, 1 ♂, reg nº 4829, IICT.

Remarks. A species with a distribution throughout D. R. of the Congo, People`s Republic of Congo and Angola (Werner 2000b). Basilewsky (1955) and later Ferreira (1965) mentioned this species as a synonymy of Lophyra reliqua Barker. As the first author identified L. reliqua based on specimens from Dundo deposited in the Museu da Companhia dos Diamantes de Angola, and our examined specimens come from the same region (one of them from the same Museum), we are confident that all specimens belong to L. obliquograciliaenea .

Lophyra (s. str.) reliqua (Barker, 1920) [Ann. Durban Mus. 2 (6): 279]

Distribution in Angola

Old records:

—Dundo (LUNDA NORTE) (51) (Basilewsky 1955; Ferreira 1965).

— Angola (without locality) (Rivalier 1948; Werner 2000b).

New localities reported: 10 km S Caluquembe (HUILA) (317) (P. Schüle pers. comm.). Remarks. A species widespread throughout Central and southern Africa, including Angola (Werner 2000b). Lophyra reliqua , as we indicated under the remarks of the preceding species, was probably mistaken for L. obliquograciliaenea by previous authors (e.g. Rivalier 1948; Basilewsky 1955), who considered the latter species a junior synonym of the former. As a consequence of the new confirmed captures of L. reliqua near Caluquembe, the species is confirmed for Angola. Peter Schüle (pers. comm.) found the adults on the edges of a big pond.

Lophyra (Stenolophyra) infuscatula (W. Horn, 1915) (Wytsman, Gen. Ins. Cic.: 266)

[= infuscata Quedenfeldt, 1883 (Berl. Ent. Zeitschr. 27: 245)]

Distribution in Angola

Old records:

— Type locality: Malange (Mechow), 1 ex., MNHN?

—Malange (=Malanje) (MALANJE) (131) (Quedenfeldt 1883 sub Cicindela infuscata ).

—Malange (=Malanje) (MALANJE) (131), Chipeyo (=Pedra do Chipeio) (BENGUELA) (253), Chiyaka (=Tchiacana) (BENGUELA) (251), Ekuiva River (=Kuiva River) (BENGUELA or HUAMBO) (Wellman & Horn 1908 sub Cicindela infuscata ).

—Bimbi (=Bimbe) (HUAMBO) (210) (Horn 1935; Monard et al. 1956; Ferreira 1965 all sub Cicindela infuscatula ). New localities reported: 10 km and 20 km S Caluquembe, 3,5 km SW Negola, 20 km NE Cacula (HUILA)

(317) (P. Schüle pers. comm.).

Remarks. A species described from Angola based on collections by Mechow. We were unable to determine

where the type specimen is deposited. Presently it is known from Angola, D. R. of the Congo and Tanzania (Werner

2000b). It seems to occur on paths of meadows. Peter Schüle (pers. comm.) found the adults in open secondary

forests.

Lophyra (Stenolophyra) obtusidentata (Putzeys, 1880) (Journ. Sc. Math. Phys. Nat. Lisboa: 22)

Distribution in Angola

Old records:

— Type locality: Angola, (Welwitsch), 1♂, 3 Paratypes, MLZA.

— Luanda (south outskirts) ( LUANDA) (89) (Werner 2000b).

New localities reported: Cassoço (= Cassoco) (KWANZA SUL) (185) (Th. Bouyer, pers. comm.); Luanda (south outskirts) ( LUANDA) (89) (A. Kudrna pers. comm.).

Material examined: Angola (Welwitsch), 1♂, reg. nº 1901, “Old Collection” MZUC; Angola (Welwitsch), 2♀, reg. nº 1907, “Old Collection” MZUC; Veados, Morro dos ( LUANDA) (89), A.Serrano leg., 1♀, 25.IV.1975, ASC; same locality, R. Capela leg., 6 ♂, 5♀, 9.II.2013, ASC, 1♂, 1♀, PSC; same locality, R. Capela leg., 5 ♂, 1♀, 12.IV.2013, ASC; Palmeirinhas ( LUANDA) (89), R. Capela leg., 25.III.2013, 1♀, ASC.

Remarks. A large Lophyra species restricted to Angola and Namibia (Werner & Wiesner 1995). The type locality in Angola is unknown, but more recent colections in this country indicate that the species occurs south of Luanda toward the Cuanza River, a region visited by Welwitsch also. From October 1853 to March 1854 this Austrian botanist visited several litoral sites around Luanda from Quicembo to Cuanza river mouth (Crawford- Cabral & Mesquitela 1989). A. Bandinelli (in Werner 2000b) indicated that this species lives in open bush habitat on sandy yellow-reddish soil with sparse grassy vegetation. When disturbed it flies or runs, trying to hide under the thorny bushes (R. Capela pers. comm.).

The type series of this species deposited in MLZA (Museu Bocage) was destroyed during a catastrophic fire in 1978. As a consequence of this event and in the interest of the stability of the type designation a neotype is here designated based on one of the 3 specimens deposited in the MZUC. The specimens of MZUC bear the same data as that of the destroyed type series and were probably identified by Paulino de Oliveira. I therefore designate the male specimen, explicitly labelled with a similar data mentioned in Putzeys (1880) as the neotype. The three specimens of MZUC are damaged. We selected the male specimen with part of both antennae, part of the left anterior leg and the right intermediate leg. The aedeagus of one male specimen from the new material examined was extracted and illustrated here ( Fig. 5 B) to avoid damaging the neotype specimen.

Type specimen ( Fig. 5A ): Neotype ♂ (designated here) in MZUC, nº 1901, two labels, one labelled “ Angola Welwitsch” (white label printed), other labelled “M.Z.U.C., registo nº 1901” (blue label, printed and handwritten), design. A. Serrano (red printed);

Redescription.Diagnosis. A robust and distinct species within the subgenus Stenolophyra , with head and thorax metallic bronze, sulci golden greenish iridescents, elytra dark bronze dull, largely marginate with a continuous white-yellowish scalariform band. Legs green metallic. Labrum with 2 submarginal setae. Cheeks glabrous or only slightly pilose; antenomeres 5–11 orange-brownish coloured. Male antennae devoid of penicillum.

Pronotum strongly sculptured, with sparse decumbent white pubescence on anterior and lateral margins. Underside covered with white long decumbent posteriad pubescence on the the sides of mesosternum, metasternum and side margins of the abdominal sternites, except the last one that is glabrous.

Head metallic bronze, with violet or golden greenish reflections on clypeus and frons, darker on vertex. Supraocular striae distinct, interocular region with irregular strong ridges, finer and transverse on the vertex; frons with strong longitudinal ridges congregated posteriad forming a rosete. Surface glabrous, only two juxtaorbital sensorial steae near both eyes, internal angle of eyes without decumbent hairs; cheeks strongly striated, only with some semi-erect white hairs. Gular area blueish, the tooth of mentum violet, long. Labrum quite large, entirely testaceous, darker in the middle region which is humped posteriorly, narrowly blackened on the margins, only the central tooth is distinct and slightly bent down; two setigerous punctures near forward edge. Mandibles stout, similar in both sexes, testaceous at base, black apically. Labial palpi testaceous, the last article metallic green;maxillary palpi darker, the penultimate and last articles less metallic green. Antennae slightly longer in males than in females, in the former reaching posteriorally and sometimes surpassing the large humeral band, but in the latter rarely reaching this band; scape metallic bronze with apical region and the antenomeres 2–4 metallic green with green reflections; a single long seta on the scape near the tip, three and one bristles on the external edge of the third and fourth antenomeres, articles 5–11 orange-brownish coloured, gradually darkened to the last ones, covered with a very fine regular pubescence: The male antennae devoid of a penicillum.

Pronotum sub-squared, metallic bronze, sulci with golden greenish reflections, 1.4 times broader than long, narrowed forwards, the maximum width after the middle; only the episternal suture slightly visible from above.

Surface strongly sculptured, somewhat granited; disk very convex, with irregular, transverse waves; anterior, medium and posterior sulci strong. Anterior and lateral margins with sparse decumbent white pubescence, posterior margin and disk glabrous. Scutellum glabrous, first half green and second half purple.

Elytra almost paralled-sided, 1.8–1.9 times longer than broad, faintly convex, maximum breadth in the last third in both sexes. Shoulders nearly square, apical edge serrulated, apex similar in both sexes, with a sutural spina. Colour dark bronze dull, elytral markings large constituted by: a broad continuous marginal band from base to apex; a wide humeral lunule almost reaching the scutellum anteriorly and transverse-parallel sided posteriorly, ending sometimes in a large button; a very broad transverse-parallel sided middle lunule, not touching the suture at distal end; a broad apical lunule largely protruding on the disk; one juxtasutural dot imediately below the middle band rarely connected with it. A single, longitudinal row of large punctuations runs parallel to, but some distance from, the suture. Extreme lateral margins of the elytra metallic blue-green in the first third, blachish in the remaining part; epipleura rufous-testaceous.

Underside of the thorax metallic violet with violaceous reflections, especially on the prosternum; mesosternum dark brown, lateral parts greenish with some reflections; metasternum dark brown, usually with metallic greenviolet reflections anteriorly. Anterior and middle coxae reddish or dark brown with green reflections, hind coxae dark brown-reddish. Episterna almost black, epimera dark blue-violet, covered moderately with decumbent long white hairs; proepimerite with a few long, erect white hairs near the margins of coxae. Metasternum covered moderately with decumbent white long hairs at their lateral regions. Abdomen dark brown-reddish with greenviolet metallic reflections mostly in the first two sternites, all sternites except the last one, covered on the side margins with sparse decumbent white long hairs; hind margin of the last sternite moderately sinuate inwards (males), straight and excavated in the middle region (females).

Legs long and robusts. Femora metallic bronze at the front, metallic green at the back, with bronze and green reflections respectively, moderately pilose all around (anterior and middle femora) or only at the front part (hind femora); one row of longer white erect setae above and underneath. Tibiae and tarsi green, with metallic reflections.

Male aedeagus as in figure 5B.

Length: Neotype ♂ 11.3 mm; material examined: ♂♂ (N=7) 11.3–12.1 mm, ♀♀ (N=6) 12.9–13.2 mm (without labrum).

Lophyra (Stenolophyra) saraliensis saraliensis (Guérin-Méneville, 1849) (Rev. Mag. Zool. 2 I: 80)

Distribution in Angola

Old records:

—Malange (=Malanje) (MALANJE) (131) (Quedenfeldt 1883 sub Cicindela flammulata Quedenfeldt ).

—Malange (=Malanje) (MALANJE) (131), Elende (= Elendé) (HUAMBO) (255), Chiyaka (=Tchiacana) (BENGUELA) (251), Ekuiva River (=Kuiva River) (BENGUELA or HUAMBO) (Wellman & Horn 1908, sub Cicindela saraliensis ).

—Bimbi (=Bimbe) (HUAMBO) (210) (Horn 1935; Monard et al. 1956; Ferreira 1965 all sub Cicindela saraliensis ). New localities reported: 75 km N (near Cuima) and 75 km NE Caconda (HUAMBO) (280); 10 km S

Caluquembe, 3,5 km SW Negola (HUILA) (317) (P. Schüle pers. comm.).

Remarks. A polymorphic tiger beetle species widespread throughout the western, central and eastern

countries of Africa (Werner 2000b). In Angola it was found on dark soil (Wellman & Horn 1908). According to

these authors the adults do not fly and are “often found in the edge of woods near certain Cosmenae, but not within

them”. According to Werner (2000b) the adults are attracted to lights. Peter Schüle (pers. comm.) found the adults

during daytime in meadows, arable fields and open secondary forests.

Lophyra (Stenolophyra) uncivittata (Quedenfeldt, 1883) (Berl. Ent. Zeitschr. 28: 242)

Distribution in Angola

Old records:

— Type locality: Malange (Mechow), 1 ♂, MHNH.

—Malange (=Malanje) (Mechow) (MALANJE) (131), Ekuiva river (BENGUELA) (254) (Wellman & Horn 1908 sub Cicindela uncivittata ).

—Bimbi (=Bimbe) (HUAMBO) (210) (Horn 1935; Monard et al. 1956; Ferreira 1965; all sub Cicindela uncivittata ). —Chiyaka (=Tchiacana?) (BENGUELA) (251) (Werner 2000b).

New localities reported: 75 km N (near Cuima) and 75 km NE Caconda (HUAMBO) (280); 20 km N Cacula, 3,5 km SW Negola (HUILA) (317) (P. Schüle pers. comm.).

Remarks. A species with a distribution throughout Peoples Republic of Congo, D. R. of the Congo, Angola, Zambia, Zimbabwe and Malawi (Werner 2000b, 2003b). Lophyra uncivittata was described from material collected by Major von Mechow in Angola. It was found in the clayey soil of the Kuiva river banks (Wellman & Horn, 1908). On the other hand, P. Schüle (pers. comm.) found the adults during day on sandy areas in open secondary forests.

Lophyra (Bothryolophyra) wellmani (W. Horn, 1907) (Deutch. Ent. Zeitschr.: 421)

Distribution in Angola

Old records:

— Type locality: Ciyaka (=Chiyaka=Tchiacana), 1 ♂, SDEI.

—Chiyaka (=Tchiacana) (BENGUELA) (251) (Wellman & Horn 1908 sub Cicindela wellmani ). —Bimbi (=Bimbe) (HUAMBO) (210), Ebanga (BENGUELA) (254) (Horn 1935; Monard et al. 1956; Ferreira

1965 all sub Cicindela wellmani ).

—Ballundo (=Bailundo) (HUAMBO) (232) (Werner 2000b).

New localities reported: 75 km N (near Cuima) and 75 km NE Caconda (HUAMBO) (280); 20 km S Caluquembe, 3,5 km SW Negola (HUILA) (317) (P. Schüle pers. comm.).

Remarks. This is the smallest Lophyra species known, endemic of Angola. Wellman & Horn (1908) included some ecological remarks based on the observations of the first author who found the adults “on paths or on other bare places, clayey soil, on which the coloring of the beetle makes it hard to see”. Moreover it seems to move like the ants. On the other hand, Peter Schüle (pers. comm.) found the adults during day on sandy areas in open secondary forests also.

Lophyra (Juengeria) jungeriorum Mandl, 1973 (Ent. Arb. Mus. Frey: 301)

Distribution in Angola

Old records:

— Type locality: Calulo (KWANZA SUL) ( Angola) (128), 1♀, NHMW.

Remarks. Cassola (2003) tentatively elevated the subgenus Juengeria to the genus status on the basis of some “quite unusual charaters” such as the underside pubescence, the genae fully glabrous, mandibles rather short, the large unidentate, four-haired labrum with hind half dark green instead of testaceous and the pattern of elytral maculation. However, most of these characters can be observed in L. (Stenolophyra) obtusidentata and the forebody of this species and L. (Juengeria) jungeriorum are almost identical. So, without the examination of a male specimen all these characters we judege that they are inadequate to support that decision (P. Schule pers. comm.). By this, we assume provisionally for this species the classic status of Juengeria subgenus, without neglecting the possibility that it belongs to the subgenus Stenolophyra . Only the holotype specimen is known and it was collected on a laterite trail (Jünger 1967 in Cassola 2003).

Figures

FIGURE 5. A) Neotype habitus: Lophyra (Stenolophyra) obtusidentata (Putzeys, 1880), male, “ Angola Welwitsch ”, “ Old collection ” MZUC. B) Lophyra (Stenolophyra) obtusidentata: aedeagus (lateral view).