Eupsenella bugulmara Ramos & Azevedo

Ramos, Magno S. & Azevedo, Celso O., 2012, Revision of Eupsenella Westwood, 1874 (Hymenoptera, Bethylidae), Zootaxa 3539 (1), pp. 1-80 : 29

Eupsenella bugulmara Ramos & Azevedo , sp. nov.

( Figs 20 , 65, 110)

Description. FEMALE. Body length 9.9 mm. LFW 5.48 mm.

Coloration. Head dark castaneous almost black; scape dark castaneous; pedicel and flagellum castaneous; mandible dark castaneous; palpi castaneous; mesosoma dark castaneous almost black; tegula and wing venation dark castaneous; coxae dark castaneous almost black; trochanters dark castaneous; femora and tarsi castaneous; metasoma dark castaneous almost black.

Head ( Fig. 20 ). Head wider than long, subtriangular in profile; gena without concavity in profile. Mandible with base wider than apex, teeth sub-equal, lower tooth curved inward, lower margin strongly setose. Clypeus coriaceous-punctate, median clypeal lobe angulate, median concavity of median clypeal lobe absent, 0.15 LH, well marked off from frons, median carina present, 0.1 LH. Malar space conspicuous, 0.42 mandibular base. Antenna 2.38 mm; first five antennomeres in ratio of about 14:14:14:14:14; flagellomere 1 1.28 longer than wide, flagellar pubescence dense, with few outstanding erect setae. Region between torulus and eye concave. Area between the anterior margin of eyes and demarkation of median clypeal lobe 0.14 LH. Eye not protuberant, 0.40 LH, with sparse hairs, located more laterally, lower margin without strong depression. Frons coriaceous-puncate. WH 1.06 LH. WF 0.66 WH. WF 1.53 HE. OOL 1.2 WOT. VOL 0.67 HE. Distance from posterior ocellus to vertex crest 1.5 DAO. Ocelli small, DAO 0.10 WF, ocellar triangle not compact, distance between posterior ocelli 3.0 DAO, anterior ocellus not surpassing imaginary top eye line, posterior ocellus distant from vertex. Vertex straight, long setae, 0.18 LH, temple profile approximately parallel in dorsal view, with corner convex, not projected posterolaterad. Hipostomal carina angulate, high.

Mesosoma (Fig. 65). Pronotal disc coriaceous-punctate, 0.36 as long as wide, anterior corner rounded, posterior margin concave medially, pronotal disc without short longitudinal furrow. Parapsidal furrow wholly conspicuous. Notauli parallel to each other. Mesoscutum 1.1 length pronotal disc. Scutellum coriaceous-punctate, 1.0 length pronotal disc, almost flat in profile, posterior margin almost straight medially, scutellar pit elliptical, narrowed, distant each other 11.0 its own diameter. Dorsal axillar surface with fovea oval. Metanotum with median region almost flat, foveolate, lower than scutellum in profile, with fovea oval laterally. Propodeal disc wider than long; anterior margin without foveolate or transverse furrow; median carina 0.83 length propodeal disc, incomplete posterad; discal carina 0.31 length propodeal disc, converging posterad, straight; followed by depressed narrow area, foveolate, median and discal carina disc predominantly slightly confused-rugulose. Lateral of propodeum predominantly coriaceous, anterior region with pits upper one shallower than the others. Mesopleuron with subtegular furrow oval; mesepimeral suture almost inconspicuous posterad; mesopleural epicoxal lobe inconspicuous. Propleuron with sequence of foveae from anterior margin of prosternum until cervical collar absent. Prosternum with median excavation complete, median suture present. Mesopectus with posterior depression heartshaped; median suture not evident.

Wings. Macropterous. Forewing ( Fig. 110 ) with 1M cell subpentagonal; 1M and 1R1 cells not aligned basally; C cell evenly narrow;1R1 cell present, distinctly longer than 2R1 cell, evenly wide; Rs b straight; Cu b present and vestigial; 2R1 cell 0.16 LFW; distal margin of Rs c convex; r-rs vein strongly inclined; R1 b arching distally; Cu b absent; Rs+M vein 1.60 Rs a; M vein 1.44 Rs a; Rs a subperpendicular to anterior margin of forewing; stigma rectangular; distal stigmal margin truncate. Hind wing with five distal hamuli, last hamulus adjacent each other.

Legs. Profemur swollen, 0.5 as wide as long. Metacoxa with median region 0.75 as wide as long. Metacoxal projection in dorsal region conspicuous, apex rounded.

Metasoma. 3.14 as wide as long, ventral carina of petiole present.

MALE. Unknown.

Material examined. Holotype ♀. AUSTRALIA, Tas [mania], 43.25S 146.10E, Melaleuca, Bathurst, Harbour, 12–17 Feb [ruary] 1960, I.D.Naumann, trays at base of Malaise; margin Leptospermum scrubs heathy sedgeland, ANIC.

Distribution. Australia (Tasmania).

Remarks. This species is very different of the others by having the head rectangular in profile, the median clypeal lobe projected, the median carina of propodeal disc slightly longer than half length of propodeal disc, the

Figures

FIGURES 15–26. Head in dorsal view. 15. Eupsenella baada sp. nov. 16. E. barada sp. nov. E. barna sp. nov. 18. E. batjala sp. nov. 19. E. bilingara sp. nov. 20. E. bugulmara sp. nov. 21. E. dalla sp. nov. 22. E. djagaraga sp. nov. 23. E. eora sp. nov. 24. E. ilba sp. nov. 25. E. inawonga sp. nov. 26. E. inggarda sp. nov. (Scale bar = 300 µm).

FIGURES 103–112. Right forewing. 103. Eupsenella araba sp. nov. 104. E. arabana sp. nov. 105. E. baada sp. nov. 106. E. barada sp. nov. 107. E. barna sp. nov. 108. E. batjala sp. nov. 109. E. bilingara sp. nov. 110. E. bugulmara sp. nov. 111. E.