Eupsenella barada Ramos & Azevedo

Ramos, Magno S. & Azevedo, Celso O., 2012, Revision of Eupsenella Westwood, 1874 (Hymenoptera, Bethylidae), Zootaxa 3539 (1), pp. 1-80 : 24-25

Eupsenella barada Ramos & Azevedo , sp. nov.

( Figs 16 , 61, 106)

Description. FEMALE (holotype). Body length 9.8 mm. LFW 5.14 mm.

Coloration. Head dark castaneous almost black; scape dark castaneous; pedicel, flagellum and mandible castaneous; palpi light castaneous; mesosoma dark castaneous almost black; tegula dark castaneous; wing venation predominantly light castaneous with veins darking apicad; procoxa and mesocoxa dark castaneous almost black; metacoxa dark castaneous; trochanters castaneous; profemur dark castaneous almost black; mesofemur castaneous; metafemur dark castaneous; tarsi dark castaneous; metasoma dark castaneous with light castaneous portions along its length.

Head ( Fig. 16 ). Head wider than long, subtriangular in profile; gena without concavity in profile. Mandible with base wider than apex, two lowermost teeth larger than upper ones, lower tooth curved inward, lower margin strongly setose. Clypeus polished, median clypeal lobe truncate, median concavity of median clypeal lobe absent, 0.07 LH, well marked off from frons, median carina absent. Malar space conspicuous, 0.41 mandibular base, not carinate. Antenna 2.33 mm; first five antennomeres in ratio of about 14:14:14:14:14; flagellomere 1 1.4 longer than wide, flagellar pubescence dense, with many outstanding erect setae. Region between torulus and eye straight. Area between the anterior margin of eyes and demarkation of median clypeal lobe 0.17 LH. Eye protuberant, 0.55 LH, with sparse hairs, located more laterally, lower margin without strong depression. Frons coriarious-punctate. WH 1.1 LH. WF 0.69 WH. WF 1.41 HE. OOL 0.8 WOT. VOL 0.3 HE. Distance from posterior ocellus to vertex crest 0.5 DAO. Ocelli small, DAO 0.07 WF, ocellar triangle not compact, distance between posterior ocelli 4.5 DAO, anterior ocellus surpassing imaginary top eye line, posterior ocellus almost reaching vertex. Vertex straight, long setae, 0.21 LH, temple profile slightly convergent in dorsal view, with corner convex, not projected posterolaterad. Hipostomal carina arched, low.

Mesosoma (Fig. 61). Pronotal collar regular-sized. Pronotal disc coriaceous-punctate, long, 0.5 as long as wide, anterior corner rounded, posterior margin concave medially, pronotal disc without short longitudinal furrow. Parapsidal furrow conspicuous only at third posterior part. Notauli convergent posterad. Mesoscutum 0.90 length pronotal disc. Scutellum coriaceous-punctate, 0.81 length pronotal disc, almost flat in profile, posterior margin almost straight medially, scutellar pit elliptical, narrowed, distant each other 9.0 its own diameter. Metanotum with median region almost flat, coriaceous-puncticulate, as high as scutellum in profile, with fovea oval laterally. Propodeal disc wider than long; anterior margin without foveolate or transverse furrow; median carina 0.75 length propodeal disc, incomplete posterad; discal carina 0.20 length propodeal disc, parallel to median carina, straight; space between median and discal carina imbricate and confused-rugulose. Mesopleuron with subtegular furrow elliptical; mesepimeral suture wholly conspicuous; mesopleural epicoxal lobe inconspicuous. Propleuron with sequence of foveae from anterior margin of prosternum until cervical collar absent. Prosternum with median excavation complete, median suture present. Mesopectus with posterior depression composed by two large foveae almost fused; median suture evident.

Wings. Macropterous. Forewing ( Fig. 106 ) with 1M cell subpentagonal; 1M and 1R1 cells not aligned basally; C cell evenly narrow; 1R1 cell present, distinctly longer than 2R1 cell, evenly wide; Rs b convex; 2R1 cell 0.15 LFW; distal margin of Rs c convex; r-rs vein inclined; R1 b entirely on anterior margin of forewing; Cu b absent;

Legs. Profemur very swollen, 0.68 as wide as long. Metacoxa with median region 0.7 as wide as long. Metacoxal projection in dorsal region conspicuous, apex sharpened.

Metasoma. 5.0 as wide as long, ventral carina of petiole present.

MALE. Unknown.

Material examined. Holotype ♀. AUSTRALIA, A[ustralian] C[apital] T[erritoy], 35.35S 149.00E, Honeysuckle Creek , 23 Apr [il]– 8 May 1985, I.D.Naumann & J.C.Cardale, Malaise trap /ethanol, ANIC . Paratypes. AUSTRALIA, Q[ueens]L[an]d, 1 ♀, 26.50S 151.33E, Burtons Well , Bunya M [oun]t[ain]s Nat [ional] P[ar]k, 6–7 Oct [ober] 1984, I.D.Naumann & J.C.Cardale, ex[tracted] ethanol ; 1 ♀, 26.52S 151.34E, n[ea]r Westcott Plain, I.D. Naumann & J.C.Cardale, Malaise /ex[tracted] ethanol, ANIC ; N[ew] S[outh] W[ales], 1 ♀, Coocumbac I [sland]s, Natural Reserve , Taree, 1–8 Dec [ember] 1994, G.&T.Williams, ex[tracted] floodplain/rainforest, ANIC ; A[ustralian] C[apital] T[erritory], 1 ♀, Weston , 13 June 1981, T. E. Bellas, ANIC ; 1 ♀, Black M [oun]t[ai]n, 1–6 Jan [uary] 1980, Malaise Site 1, D.H.Colless, ANIC , 1 ♀, Mar [ch] 1982, Malaise Trap, I.D.Naumann & J.C.Cardale, ex[tracted] ethanol, ANIC ; 1 ♀, 12–30 Apr [il] 1980, Malaise Site 2, D.H.Colless, ANIC ; 1 ♀, Dec [ember] 1982, I.D.Naumann & J.C.Cardale, Malaise trap /ethanol, ANIC ; 1 ♀, 35.16S 149/ 149.10E, M[oun]t[ain] Ainslie, June 1991, under bark, C. Reid, ANIC.

Distribution. Australia (Australian Capital Territory, Queensland, New South Wales).

Variation. Body length 4.52–8.57 mm; LFW 4.04–5.00 mm; clypeus and pronotal disc short to moderately long.

Remarks. This species is similar to E. baada sp. nov. by having the area between the anterior margin of eyes and demarkation of median clypeal lobe long, the clypeus with median clypeal lobe truncate and without median carina. However this species has the vertex crest slightly concave, the head nearly subtriangular in profile, the eyes large, the pronotal disc without anterior median pit, apical margin of pronotal disc nearly straight, whereas E. baada sp. nov. has the vertex crest slightly convex, the head not subtriangular in profile, the eyes smaller, the pronotal disc with anterior median pit, the apical margin of pronotal disc convex.

Figures

FIGURES 15–26. Head in dorsal view. 15. Eupsenella baada sp. nov. 16. E. barada sp. nov. E. barna sp. nov. 18. E. batjala sp. nov. 19. E. bilingara sp. nov. 20. E. bugulmara sp. nov. 21. E. dalla sp. nov. 22. E. djagaraga sp. nov. 23. E. eora sp. nov. 24. E. ilba sp. nov. 25. E. inawonga sp. nov. 26. E. inggarda sp. nov. (Scale bar = 300 µm).

FIGURES 103–112. Right forewing. 103. Eupsenella araba sp. nov. 104. E. arabana sp. nov. 105. E. baada sp. nov. 106. E. barada sp. nov. 107. E. barna sp. nov. 108. E. batjala sp. nov. 109. E. bilingara sp. nov. 110. E. bugulmara sp. nov. 111. E.