Calloeneis francis Gordon and Hanley, 2020

Gordon, Robert D., F, Guillermo González & Hanley, Guy A., 2020, South American Coccinellidae (Coleoptera), part XX: systematic revision of South American Calloeneis Grote (Cryptognathini), Insecta Mundi 2020 (766), pp. 1-25 : 13

15. Calloeneis francis Gordon and Hanley , new species

Description. Male holotype. Length 1.6 mm, width 1.4 mm; dorsal surface shiny. Color reddish brown; head, pronotum yellow; elytra with large, rounded, black basomedian macula on suture ( Fig. 64 ); ventral surface yellow except meso-, metasternun brown; abdomen yellow except apex of median 1/3 of basal ventrite dark brown. Head with dense punctures separated by a diameter or less; pronotal punctures small, separated by a diameter or less; elytral punctures as large as on pronotum, separated by up to three times a diameter; prosternal punctures small, indistinct, separated by about a diameter; mesosternal punctures larger than on prosternum, separated by less than a diameter; metasternal punctures small, separated by two or three times a diameter medially, absent in lateral 1/3; basal abdominal ventrite with small punctures separated by about three times a diameter medially, ventrite 2 with coarse punctures in median 2/3 separated by less than a diameter, ventrites 3, 4 finely, densely punctured medially; ventrite 5 finely, densely punctured. Head slightly narrowed from vertex to clypeus, medially depressed, 1.5 times as wide as eye, clypeal apex truncate ( Fig. 65 ); eye canthus short, about half width of eye. Prosternum with lateral carina on each side curved, extended from apex of intercoxal process to lateral margin of prosternum. Epipleuron slightly descending externally. Postcoxal line on basal abdominal ventrite curved throughout, extended 2/3 distance to rear margin of ventrite. Genitalia with penis guide 7/8 as long as paramere, oval, narrowed to abruptly rounded apex in apical 1/4; paramere slender, curved, parallel sided, apex rounded ( Fig. 66, 67 ); penis lost.

Female. Similar to male except genitalia with ramus of spermathecal capsule large, bulbous, cornu bent at base, narrowed before apex, apex acute ( Fig. 68 ).

Variation. Length 1.6 to 2.1 mm. Size of median black macula somewhat variable.

Type material. Holotype male; Perú Madre de Dios, Res. Biológica CICRA, 27/ 28.XI.2006, 25/ 150m., Leg. Angélico Asenjo, Flight trap intercep,( MUSM) . Paratypes, 4, 1, PERU: Madre de Dios Dept., CICRA Field Stn., trl 6, research plot, 12.55207°S 70.110962°W. 295m 11–13.VI.2011 Chaboo team, Malaise trap, PER-11-MAT-029. Paratype, 1, PERU: Cusco Dept. Villa Carmen field station, 1.7 km west cafeteria research transect 12.89213°S 71.41920°W 547m 22–24.V.2011, D.J. Bennett & E. Razuri Malaise trap PER-11-MAT-010 ; 2, Peru, Satipo , VII, VIII 1942, Paprzycki ( MUSM, USNM) .

Figures

Figures 54–68. Calloeneis spp. 54–58) Calloeneis johnnie. 54) Habitus. 55) Frons. 56) Penis guide ventral. 57) Penis guide lateral. 58) Penis. 59–63) Calloeneis robyn. 59) Habitus. 60) Frons. 61) Penis guide ventral. 62) Penis guide lateral.63) Penis.64–68) Calloeneis francis. 64) Habitus.65) Frons.66) Penis guide ventral.67) Penis guide lateral. 68) Spermathecal capsule.