Capobula, Haddad & Jin & Platnick & Booysen, 2021

Haddad, Charles R., Jin, Chi, Platnick, Norman I. & Booysen, Ruan, 2021, Capobula gen. nov., a new Afrotropical dark sac spider genus related to Orthobula Simon, 1897 (Araneae: Trachelidae), Zootaxa 4942 (1), pp. 41-71 : 52

Key to the species of Capobula gen. nov.

1 Females ............................................................................................. 2

– Males (males of C. neethlingi spec. nov. and C. ukhahlamba spec. nov. unknown).................................. 6

2 Abdomen grey, with cream inverted Y-shaped marking dorsally ( Fig. 10); epigyne with large circular depressions incorporating copulatory openings ( Fig. 73)........................................................ C. ukhahlamba spec. nov.

– Abdomen uniformly grey dorsally, without marking (e.g. Fig. 3); epigyne with curved epigynal ridges incorporating copulatory openings............................................................................................ 3

3 Copulatory openings in anterolateral corners of faint M-shaped ridges ( Fig. 54); bursae clearly much larger than primary spermathecae ( Fig. 55)................................................................... C. capensis spec. nov.

– Copulatory openings in small C- or J-shaped ridges (e.g. Fig. 59); bursae same size or smaller than primary spermathecae (e.g. Fig. 60)............................................................................................. 4

4 Anterior margin of ridges incorporating copulatory openings in same transverse plane as anterior margin of primary spermathecae; initial 2/3 of copulatory ducts before posterior bend heavily sclerotized and darker than last 1/3 ( Fig. 55)................................................................................................ C. infima comb. nov.

– Anterior margin of primary spermathecae clearly in front of anterior margin of ridges incorporating copulatory openings; only initial 1/3 of copulatory ducts before posterior bend heavily sclerotized, last 2/3 clearly lighter in colour ( Figs 65, 71)...... 5

5 Ridges incorporating copulatory openings large and almost transverse to longitudinal plane of epigyne ( Fig. 71); heads of bursae converging ( Fig. 72) ( South Africa: Western Cape, Fig. 75)............................... C. neethlingi spec. nov.

– Ridges incorporating copulatory openings small and obliquely orientated, at 45 degrees to longitudinal plane of epigyne ( Fig. 65); heads of bursae diverging ( Fig. 66) (eastern half of South Africa, Fig. 75).................... C. montana spec. nov.

6 Embolus as wide as 1/2 of tegulum width at midpoint, tip sharply curved and almost transverse to longitudinal plane of palp ( Fig. 62)................................................................................................................................................................................ C. infima comb. nov.

– Embolus narrow, width less than ¼ of tegulum width at midpoint, tip directed disto-retrolaterally ( Figs 57, 68)…7

7 Embolus short, with stout tip ( Fig. 57); femoral apophysis a sharp retrolateral spike ( Fig. 58) ( South Africa: Western Cape, Fig. 75)................................................................................ C. capensis spec. nov.

– Embolus longer, with slender tip ( Fig. 68); femoral apophysis a stout ventral lobe ( Fig. 69) (eastern half of South Africa, Fig. 75)................................................................................ C. montana spec. nov.