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Pleuronichthys lighti Wu, 1929

Yokogawa, Kōji, Watanabe, Ken-Ichi & Ogihara, Gota, 2016, Redescriptions of two closely related East Asian flatfish species of the genus Pleuronichthys, Zootaxa 4205 (1), pp. 1-23 : 4-13

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 https://doi.org/10.11646/zootaxa.4205.1.1

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DOI

 https://doi.org/10.5281/zenodo.5695907

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scientific name

 Pleuronichthys lighti Wu, 1929
status

 

Pleuronichthys lighti Wu, 1929 View in CoL

(New English name: Common frog flounder) ( Figs. 3 View FIGURE 3 A–C, 4A, B, 5A–E; Tables 1–5)

Platessa cornuta View in CoL (not of Temminck & Schlegel): Boeseman, 1947: 145 –146, RMNH 3496 About RMNH a, 3496b, 3501 (synonymy; Japan).

Parophrys cornuta (not of Temminck & Schlegel): Günther, 1862: 455 (synonymy and description; Japan); Günther, 1880: 70 [synonymy; Japan (Kobe)]; Nyström, 1887: 42 –43 (synonymy; Japan); Otaki, 1897: 32 –33, pl. 8 (synonymy, brief description and illustration; Japan).

Pleuronichthys cornutus View in CoL (not of Temminck & Schlegel): Steindachner, 1896: 217 [synonymy and description; Japan (Kobe or Nagasaki)] ; Jordan & Snyder, 1900: 379 [list; Japan (Hakodate and Tokyo Bay)] ; Jordan & Snyder, 1901: 769 [list; Japan (Yokohama)]; Jordan & Starks, 1906 (part): 201–202 [synonymy and description; Japan (Kobe, Aomori, Hiroshima, Nagasaki, Hakodate, Onomichi, Wakanoura, Tsuruga, Tokyo and Misaki)]; Starks & Thompson, 1910: 279 –282 [description; Japan (Tsuruga and Aomori)]; Snyder 1912: 440 [ Japan (Hakodate, Tokyo and Tsuruga)] ; Jordan & Thompson, 1914: 309 [ Japan (Osaka and Misaki)]; Hubbs, 1915: 475 –476 [synonymy and description; Japan (waters near Kagoshima, Goto Is., Genkai Sea and Tsuruga)]; Matsuura & Izuka, 1920: 117 [list; Japan (Takamatsu)] ; Jordan & Hubbs, 1925: 298 [ Japan (Tokyo, Kobe, Toba, Kagoshima, Mikawa Bay, Misaki and Fukui)]; Wu, 1932 (part): 109–110 [synonymy and description; China (Ting-Hai, Techou-San, Tchefou, Tsing-Tao, Pekin and Amoy)]; Wang, 1933: 52 –53 [synonymy and description; China (Chefoo)]; Norman, 1934: 319 –320, fig. 236 (synonymy, description and illustration; Japan (Kobe, Bay of Mutsu , Tokyo, Matsuyama, Hiroshima, Wakanoura and Inland Sea) , China (Yenting and Amoy)]; Okada et al., 1935: 239, pl. 141 (description and monochrome photo; Japan); Kamohara, 1952: 82, fig. 79 [illustration; Japan (off Kochi City)]; Chyung, 1954: 473 –474 (synonymy and description; Korea) ; Tanaka & Abe, 1955: 223 (brief description and illustration; Japan); Zheng, 1955 (part): 286–287, fig. 177 [synonymy, description and illustration; China (Zhuanghe, Pikou, Qinhuangdao, Longkou, Yantai, Shidao, Qingdao and Shijiusuo)]; Okada, 1955: 384 –385 (description; Japan); Tomiyama et al., 1958: 40 (brief description and illustration; Japan); Hiyama & Yasuda, 1961: 124, pl. 169(291) (brief description and illustration; Japan); Zheng, 1962: 981 –982, fig. 763 [synonymy, brief description and illustration; China (Haimen, Jiazi, Zhelang, Shanwei and Jieshi)]; Tchang & Wang, 1963: 527 –528, fig. 394 [synonymy, brief description and illustration; China (Shenjiamen, Kanmen, Dachen Is., Sijiao and Suao)]; Fitch, 1963: 18 –20, fig. 3 (monochrome photo, synonymy and description; China, Japan and Korea); Chen & Weng, 1965: 8 –9: [synonymy, description and illustration; Taiwan (Kaohsing, Pescadores Is., Taichung and Keelung]; Ochiai, 1965: 492 (brief description and illustration; Japan) ; Shen, 1967: 191 –192, figs. 77–80 (synonymy, description and monochrome photos; Hong Kong); Hiyama & Yasuda, 1971: 303, fig. 405 (brief description and illustration; Japan); Burgess & Axelrod, 1974: 1097 [color photo; Taiwan (Pescadore Is.)]; Masuda et al., 1975: 345, pl. 148(I) (color photo; Japan); Chyung, 1977: 569, pl. 301, color pl. 126-1 [brief description, monochrome and color photos; Korea (Pusan and Yeosu)]; Shirano, 1979: 108 [(illustrations; Japan (Kagoshima)]; Fisheries College of Shanghai et al., 1980: 146 –147, fig. 113 [brief description and illustration; China (Fujian Prov.)]; Kataoka & Tomida, 1981: 76, [list; Japan (various sites around Mie Pref.)]; Shen, 1982: 199 –201, fig. 2 [description and monochrome photo; Taiwan (Ta-chi, Tungkong and Su-au)]; Masaki & Ito, 1984: 11 –51 [ Japan ( western Seto Inland Sea )]; Shen, 1984a: 137, pl. 137(440-1a, b) (brief description and color photos; north-eastern waters of Taiwan) ; Shen, 1984b: 451 –452 (key and illustration; Taiwan) ; Chen & Yu, 1984: 821 –823, fig. 6-173(2) (key and illustration; Taiwan) ; Sakamoto & Nakabo, 1984: 337, 352, pl. 315(G) (brief description and color photo; Japan); Matsuura, 1985: 620 –621, fig. 393 [description and color photo; Japan (Okinawa Trough)]; Kamohara & Okamura, 1985: 60, pl. 131(652) (illustration; Japan) ; Miao, 1985: 539, fig. 738 [synonymy, brief description and illustration; China (Shacheng, Taishan, Dongxiang, Xiamen, Sangou and Waixie fishing grounds)]; Mochizuki, 1985: 126 –127 [color photo; Japan (Awaji I.)]; Yamada, 1986: 380 –381 (brief description and color photo; East China and Yellow Seas); Ochiai, 1986: 313 (brief description and color photo; Japan) ; Masuda & Allen, 1987: 464 (color photo; Japan); Cheng & Zheng, 1987: 500 –502, 1404, fig. 2574 (key and illustration; China); Chen, 1987: 428 –429, fig. 290 [brief description and illustration; China (northern Yellow Sea and Liaodong Bay )]; Abe, 1987: 925 (brief description and illustration; Japan) ; Minami, 1988a: 938 –940 [description and illustration of larvae and juveniles; Japan (Seto Inland Sea, coastal waters of Honshu, Kyushu and Hokkaido), southern Korean Peninsula and Yellow Sea]; Sakamoto & Nakabo, 1988a: 337, 352, pl. 315(G) (brief description and color photo; Japan) ; Xu, 1990: 361, fig. 229 [brief description and illustration; China ( Yangtze River estuary)]; Shao & Lin, 1991: 196 (brief description and color photo; sandy coastal waters around Taiwan) ; Li & Zhang, 1991 (part): 78–79, fig. 87 [brief description and illustration; China (Bohai Sea)]; Lindberg & Fedorov, 1993: 153 –156, fig. 52 [synonymy, description and illustration; Japan (Tokyo, Nagasaki, Misaki, Tsuruga, Ohama and Aomori), Korea (Pusan), China (Dalian, Sin-tsin, Qingdao and Shanghai), East China Sea )]; Kim & Kang, 1993: 102, 408 (description and color photo; Korea) ; Shen, 1993: 576, pl. 194(6) (synonymy, brief description and color photo; Taiwan) ; Masuda & Kobayashi, 1994: 405, fig. 2 [color photo; Japan (Izu Pen.); Tchang et al., 1994 (part): 286–287, fig. 177 [synonymy, description and illustration; China (Zhuanghe, Pikou, Qinhuangdao, Longkou, Yantai, Shidao, Qingdao and Shijiusuo)]; Kim & Youn, 1994: 121 –122, pl. 7(A) [synonymy, brief description and monochrome photo; Korea (Pusan and Yeosu)]; Nakabo, 1995: 204 –205 [pictorial key and color photo; Japan (Awaji I.)]; Li & Wang, 1995 (part): 238–241, fig. II-62-1 [synonymy, description and illustration; China (Dalian, Qingdao, Shenjiamen, Xiamen)]; Amaoka et al., 1995: 523 (brief description and color photos; northern Japan ) ; Su, 1995: 242 –243, fig. 299 (brief description and illustration; China); Yamada et al., 1995: 50, 206 (pictorial key and color photo; East China and Yellow Seas); Cheng, 1997: 453 –454, fig. 334 [synonymy, brief description and illustration; China (waters around Shandong Prov.)]; Suzuki & Kataoka, 1997: 243, pl. 138(733) [brief description and monochrome photo; Japan (Mie Pref.)]; Sanada, 1997: 72 [brief description and color photo; Japan (Seto Inland Sea)]; Sakamoto et al., 1997: 674 [brief description and color photos; Japan (Izu Pen.)]; Kim, 1997: 494 –495, pl. 46(189) [brief description, illustration and color photo; [Estuaries of Korea (Kunsan, Mokpo, Yeosu, Masan, Jinhae and Pusan)]; National Fisheries Research and Development Institute ( NFRDI), 1999: 184, 456, fig. 514 (description and color photo; Taiwan, East China Sea , Japan and Korea ) ; Yoshino, 1999: 247 (color photos; Japan); Nakabo, 2000: 1372 (pictorial key; Japan); Kim et al., 2001: 156, 288, fig. 467 (brief description and color photo; Korea); Shimizu, 2001: 44, 94, fig. 148 [monochrome photo; Japan (Iyo City)]; Wang, 2001: 310 –311, fig. 206 [synonymy, brief description and illustration; China ( Hebei Prov. )]; Wu, 2002: 603 –604, color fig. 73, fig. 435 [synonymy, brief description, color photo and illustration; China (Qingdao, Shanghai and Xiamen)]; Kim & Park, 2002: 413–414, brief description, color photo and illustration [Estuaries of Korea (Kunsan, Mokpo, Yeosu, Masan, Jinhae and Pusan)]; Youn, 2002: 438, 686 (synonymy and pictorial key, Korea) ; Myoung et al., 2002: 229 (brief description and color photos; Korea); Choi et al., 2002: 542 –543, color photo (Korea); Chen, 2003: 215, fig. 859 [color photo; Taiwan (Penghu Is.)]; Kim et al., 2004: 260 (brief description, pictorial key and color photo; Korea) ; Kim I. et al., 2005: 482, (brief description and color photo; Korea) ; Taki et al., 2005: 925 (brief description and illustration; Japan); Kim Y. et al., 2005: 288–289, fig. 489 (brief description and color photo; Korea) ; Yao & Yoshida, 2006: 68 (brief description and color photo; China); Zhao & Zhong, 2006: 47, 165–166, fig. 380 [brief description and color photo; China (Zhoushan)]; Ni & Wu, 2006: 818 –819, fig. 429 [synonymy; brief description and illustration; China (Lianyungang, Lusi, Yangtze River estuary and southern Yellow Sea)]; NFRDI, 2006: 336, fig. 558 (description and color photo; Taiwan, East China Sea , Japan and Korea ) ; Sokolovsky et al., 2007: 157 (illustration; Russian waters of the Japan Sea); Yamada et al., 2007: 1033 –1041, pl. 46(6) (pictorial key and color photo; East China and Yellow Seas); Choi, 2007: 267 (brief description and color photo; Korea) ; Konishi & Nakabo, 2007: 338 (brief description and color photos; Japan); NFRDI, 2008: 350, fig. 585 [description and color photo; northwestern Pacific ( Korea, Japan, East China Sea and Taiwan)]; Suzuki et al., 2009: 276 –291, fig. 1(d) (synonymy; description and monochrome photo; all around Japan) ; Sokolovsky et al., 2009: 324, 326: [brief description and color photo; Russia (Peter the Great Bay )]; Cao, 2010: 164 (brief description and illustration; China) ; Ishikawa & Senou, 2010: 331 [color photo; Japan (Fukushima)]; Yokogawa & Watanabe, 2011: 24 –32, fig. 1(a) [monochrome photo; Japan (Kii Channel)]; Zhang et al., 2011a: 51 –54 [ China (around Zhoushan)]; Ohashi & Motomura, 2011: 98 –99, fig. 28 [synonymy, brief description and color photos; Japan (northern Kagoshima Pref.)]; Choi et al., 2011: 158 (brief description and color photo; Korea) ; Uyeno & Sakamoto, 2011: 139, 299 (brief description and illustration; Sea of Japan, East China, Yellow, Bohai and South China Seas)]; Zhang et al., 2011b: 967 –972, fig. 1(upper photo) (color photo; East China Sea ); Shen & Wu, 2011: 746 (brief description and color photo; Taiwan) ; Chen & Yang, 2013: 317 (synonymy, brief description and color photo; Taiwan Strait); Nakabo & Doiuchi, 2013: 1676 (pictorial key; Japan); Igarashi & Ito, 2013: 9 –14 [ Japan (off Fukushima Pref.)]; Ito, 2013: 15–22 [ Japan (off Fukushima Pref.)]; Tsuji, 2013: 12, 20, fig. 64 [monochrome photo; Japan (Hakata I.)]; Liu et al. 2013: 250 –251 [description and color photos; China (southern Fujian Prov.)]; Jo & Myoung, 2013: 310 –311 (brief description and illustrations; Korea) ; Minami, 2014a: 1462 –1463 [description and illustration of larvae and juveniles; Japan (Seto Inland Sea, coastal waters of Honshu, Kyushu and Hokkaido), southern Korean Peninsula and the Yellow Sea]; Myoung et al., 2015: 257 (brief description and color photos; Korea) .

Pleuronichthys lighti: Wu, 1929: 66 View in CoL , fig. 54 [original description and illustration; China (Amoy)]; Yokogawa et al., 2014: 385 – 392; Yokogawa, 2015; Yokogawa, 2016.

Pleuronichthys cornutus View in CoL , MDH-A type: Kato & Fujio, 1979: 10 –12, fig. 2(upper photo) [monochrome photo; Japan ( Sendai Bay , off Fukushima and Ibaraki Prefs.)].

Pleuronichthys cornutus View in CoL , honmeita-type: Nozawa & Kato, 1981: 1 –8, fig. 1(upper photo) [monochrome photo; Japan (off Tottori Pref.)]; Matsuoka et al., 1989: 23 –36, fig. 1(A) (color photo; East China and Yellow Seas); Watanabe et al., 1992: 2253 –2260 [ Japan (Kii Channel)]; Watanabe , 1994: 1–73, fig. 1(4) [illustration of larvae and juvenile; Japan (waters around Tokushima Pref.)]; Watanabe et al., 1994: 515–520 [ Japan ( Kii Channel )]; Watanabe et al., 1996a: 17 –22 [ Japan ( Kii Channel )]; Watanabe et al., 1996b: 23 –27 [ Japan ( Kii Channel )]; Matsuoka , 1996a: 47 –54 ( East China and Yellow Seas); Matsuoka , 1996b: 55 –62 ( East China and Yellow Seas).

Pleuronichthys cornutus View in CoL , WK-A type: Watanabe & Numachi, 1988: 765 –772 [ Japan (Kii Channel)].

Pleuronichthys sp. View in CoL : Nakabo, 1995: 204–205 [pictorial key and color photo; Japan (Maizuru)]; Kim et al., 2001: 156, 288, fig. 468 (brief description and color photo; Korea); Kim Y. et al., 2005: 288–289, fig. 490 (brief description and color photo; Korea) .

Diagnosis. Scale length (trunk area) less than 1.23% of SL (Table 1), whole scale outline fan-shaped ( Fig. 3 View FIGURE 3 A–C), exposed margin rhombic ( Fig. 4 View FIGURE 4 A, B). Scales non-deciduous, longitudinal and vertical scale rows essentially regular ( Fig. 4 View FIGURE 4 A, B). Upper eye area usually less than lower eye area, area ratio 0.70–1.08 (Table 1). Central longitudinal scales 88–105, scales below lateral line 59–74, abdominal vertebrae 12–14 (usually 13) (Tables 1, 2).

Description. Measurement data for length-measured characters subject to isometric growth and meristic counts are shown in Tables 4 and 5, respectively. Body deep, rhombic or ovate, rather compressed, ocular and blind sides convex and almost flat, respectively.

Entire body covered with very small cycloid scales in essentially regular longitudinal and vertical scale rows ( Fig. 4 View FIGURE 4 A, B). Scales basically fan-shaped ( Fig. 3 View FIGURE 3 A–C), exposed margin rhombic ( Fig. 4 View FIGURE 4 A, B), except on caudal peduncle (both sides), most blind side scales with sharp posterior margins ( Fig. 3 View FIGURE 3 C). Scales deeply embedded, nondeciduous, entirely covered by thin tough pigmented epithelial membrane. Minute scales covering entire membrane of dorsal, anal and caudal fins (both sides). Ocular and blind side pectoral fin bases covered broadly and narrowly with minute scales, respectively. Scales absent on pelvic fins.

Eyes on right side of head, globular, extremely protuberant. Upper eye usually scarcely posterior to and of less mass than lower eye. Eye diameters proportional to HL relatively large, stable with growth, proportional to SL decreasing considerably with growth. Inter-orbital space very narrow, forming a sharp bony ridge terminating with strong conical spiny prominences on each edge, directed forward and backward, respectively. Snout very short, rounded. Two closely placed nostrils on each side, anterior nostril larger, with fleshly flaps anteriorly and posteriorly. Ocular side nostrils located on anterior extension of inter-orbital ridge, blind side nostrils on higher level than ocular side. Mouth very small, oblique, lips fleshy, lower jaw slightly projecting. Jaw lengths proportional to HL stable with growth, proportional to SL decreasing considerably with growth. Small acute conical teeth in two or three rows on blind side of both jaws, teeth in villiform bands on ocular side of both jaws. Gill rakers less developed, a few small tubercles on upper and lower limbs of first arch.

Dorsal and anal fin base very long. Dorsal fin anteriorly twisted toward blind side, originating from middle of upper eye position on blind side, terminating on upper middle ridge of caudal peduncle. Anal fin originating from just posterior to vent, terminating on lower middle ridge of caudal peduncle. Caudal fin fan-shaped with rounded or rhombic edge. Pectoral fins slender, ocular side fin longer than that on blind side, located at uppermost positions of gill slits on both sides. Pelvic fins small, blind side fin placed slightly more anterior to ocular side fin. Caudal and pelvic fin ray lengths (both sides) proportional to SL decreasing with growth, those of dorsal, anal and pectoral fins (both sides) stable.

Two lateral lines on each side. Lateral line scales developed but usually embedded subcutaneously. Main lateral line running across center of trunk to caudal base, curving gently upward before middle of trunk. Supratemporal lateral line (SLL) running along dorsal fin base, reaching middle of trunk (occasionally extending more posteriorly), curving downward sharply at supratemporal region (both sides, but often indistinct, particularly on ocular side), occasionally a short antrose branch from curve, running forward along supratemporal edge (Table 3).

Coloration. Ocular side fresh coloration variable ( Fig. 5 View FIGURE 5 A–E), base color brown with a dark ( Fig. 5 View FIGURE 5 A), red ( Fig. 5 View FIGURE 5 B, E) or pale tinge ( Fig. 5 View FIGURE 5 D), vermiculated with brown to dark-brown (occasionally rather pale) ( Fig. 5 View FIGURE 5 D) blotches or spots, varying in size and density. Occasionally, such patterns rather modest, a fine dark mesh pattern surrounding scales ( Fig. 5 View FIGURE 5 E). Some individuals with additional larger dark blotches with indistinct outline ( Fig. 5 View FIGURE 5 B, E), or a broad ivory vertical band (sometimes indistinct) ( Fig. 5 View FIGURE 5 C) posterior to opercle ( Fig. 5 View FIGURE 5 B). Ocular side dorsal, anal and pelvic fins with similar blotches or spots, caudal membrane usually dark, pectoral fin usually with a broad blackish band distally. Blind side uniformly white to ivory, dorsal, caudal and anal fins dusky or dark distally, pectoral and pelvic fins white. Preserved specimens (formalin or alcohol) with ocular side base color more brownish, with darkish blotches but ivory post-opercular band mostly lost; blind side yellowish.

Distribution and habitat. Pleuronichthys lighti is broadly distributed in coastal waters around the Japanese archipelago (southward from southern Hokkaido) and along the entire coast of the Korean Peninsula, China and Taiwan, as well as occurring in offshore waters of the Bohai, Yellow and East China Seas, the latitudinal range extending from Peter the Great Bay ( Sokolovsky et al., 2009) (northern limit) to Hong Kong ( Shen, 1967) (southern limit). The species inhabits mainly sandy bottoms in shallow coastal waters, being relatively dominant over P. cornutus in Tokyo and Ise Bays, and the Seto Inland, Ariake and Yatsushiro Seas. It may also have some tolerance for reduced salinity, occasionally occurring in river estuaries in China ( Xu, 1990; Ni & Wu, 2006) and Korea ( Kim, 1997; Kim & Park, 2002) and brackish waters in Japan ( Hiyama & Yasuda, 1961, 1971).

Remarks. Until Suzuki et al.'s (2009) taxonomic revision, Pleuronichthys lighti had been regarded as a junior synonym of P. cornutus ( Wu, 1932; Wang, 1933; Zheng, 1955; Li and Wang, 1995; Evseenko, 2003, 2004), there being only a single taxon of " P. cornutus " recognized for East Asian Pleuronichthys . Suzuki et al. (2009), who formally indicated two taxa of East Asian Pleuronichthys , also regarded P. lighti as a junior synonym of their " P. cornutus " (common frog flounder), owing to their mis-identification of the P. cornutus lectotype. However, because the name P. cornutus has been transferred to the phantom frog flounder (new common name, see below), P. lighti should be resurrected as the valid name for the former ( Yokogawa et al., 2014).

Although some authors ( Reeves, 1927; Wu, 1929; Chu, 1931) credited P. lighti to Herre, there is no evidence for a species of this name described in Pleuronichthys by Herre ( Fowler, 1933; Eschmeyer, 2015). This suggests that P. lighti has been cited from unpublished or unavailable publication(s) authored by Herre. In this case, therefore, the author of this nominal taxon is regarded as Wu (1929), his publication being the first available description of the new taxon (ICZN 1999: Art 12.2). On the other hand, neither Reeves (1927) nor Chu (1931) provided a description, definition or indication of the status (ICZN 1999: Art. 12.1) of P. lighti . In addition, their material could not be identified as either of the two Pleuronichthys species.

Pleuronichthys lighti is clearly distinguished from P. cornutus by squamatological characteristics, including scale length ( SCL), scale shape, and counts of central longitudinal scales and scales below the lateral line (Table 1). For practical purposes, examination of scales removed from a specimen allows ready identification. For example, Shen's (1967) description of " P. cornutus " from Hong Kong included photographs of individual scales. Since they were clearly fan-shaped, the specimen could be readily identified as P. lighti . SCL proportions (% of SL) of 0.48– 0.92 (reciprocal numbers calculated from SCL proportion data), falling within the range for P. lighti (Table 1), supported that identification. On the other hand, when the removal of scales from a specimen is problematic, the exposed scale area index ( ESAI) substitutes for SCL, Yokogawa et al. (2014) demonstrating such in their reidentification of the P. cornutus lectotype using squamatological characteristics.

Other than squamatological characteristics, meristic counts of central longitudinal scales (CLS) and scales below the lateral line (SBL), which have no range overlap between P. lighti and P. cornutus (Table 1), are very useful for identification, being supported by eye area ratio, and proportions of ocular side pectoral fin length (% of SL) and inter-orbital ridge length (% of HL), which were new key characters proposed by Yokogawa (2015) (Table 1).

Youn (2002) provided a pictorial key for distinguishing between the two Pleuronichthys species, including "presence or absence of antrorse branch of SLL," and was followed by Suzuki et al. (2009), who emphasized that character as key for such identification. However, this is a variable condition, not clearly separating the two species (Table 3). In addition, the very deciduous nature of P. cornutus scales, resulting in frequent damage to landed P. cornutus individuals, results in difficulty in assessing the SLL character (blind side) [unclear in 46.5% of the total P. cornutus individuals examined by Watanabe et al. (1996b) (Table 3)]. In the present study also, an antrose branch of the SLL was unclear in 68.3% (ocular side) and 11.1% (blind side) of P. cornutus specimens (Table 3).

Although Suzuki et al. (2009) showed significantly lower percentages of an indistinct antrose branch in their specimens (Table 3), this may have simply reflected a high proportion of well-fixed undamaged specimens in their material examined. Although Table 3 suggests specific trends in the presence / absence of an antrose branch of the SLL, the condition is clearly variable and obscured by the likelihood of damage to specimens. In conclusion, the SLL character is impractical for identification of the two species.

Suzuki et al. (2009) also emphasized counts of dorsal and anal fin rays (DFR and AFR) as diagnostic characters, frequently using them for specific identification of literature records of Pleuronichthys species. However, the ranges of these meristics for the two Pleuronichthys species broadly overlap ( Table 5 View TABLE 5 ). In addition, some inter-populational differences have been suggested ( Yamada et al., 2007). Although DFR and AFR counts in Suzuki et al. (2009) (based on specimens predominantly from Japan) suggested some separation of the two species [DFR: 72–88 vs. 67–80; AFR: 52–65 vs. 48–59 for P. lighti (= their P. cornutus ) and P. cornutus (= their P. japonicus ), respectively], Yamada et al. (2007), examining both Japanese and non-Japanese specimens, found far greater count overlaps [DFR: 68–89 vs. 65–83; AFR: 50–69 vs. 50–59 for P. lighti (= their P. cornutus ) and P. cornutus (= their P. sp.)]. Therefore, only a few individuals at the extreme ranges of DFR and / or AFR counts may be unequivocally identified as one or other species.

Pleuronichthys lighti Pleuronichthys cornutus

1 Count tends to increase with growth in P. cornutus .

On the other hand, differences in counts of abdominal vertebrae (AV) between the two Pleuronichthys species are significant (Tables 1, 2), with little range overlap (Table 2). However, being an internal character, it is problematic on practical grounds.

Blind side colored individuals have been reported for P. lighti , some specimens having all or part of the blind side colored similarly to the ocular side, with both sides generally convex ( Yamada, 1969). Such individuals may not be so rare in the field, comprising 9.2% of 195 specimens from the Yellow Sea, for example ( Yamada, 1969). They also appeared in higher percentages during artificial production ( Kitajima et al., 1987). In addition, ocular side albinism individuals are also common in this species ( Kitajima et al., 1987; Sakamoto et al., 1997).

Kim & Youn (1994) described P. lighti (as P. cornutus ) from Korea, based on 12 specimens, noting almost equal pectoral fin lengths on both sides (repeated in Kim, 1997). However, the ocular side pectoral fin was found to be clearly longer than that on the blind side in the present study (Table 1).

Because squamation and ocular side coloration are generally useful for distinguishing between the two Pleuronichthys species, many of the " P. cornutus " literature records including photograph(s) and / or illustration(s) could be specifically identified as P. lighti (see synonymy above). Similarly, photographs of Pleuronichthys sp. from Maizuru, Japan, in Nakabo (1995), but strangely omitted by Suzuki et al. (2009), and Korean specimens of P. sp. in Kim Y. et al. (2001, 2005) were considered as this species.

Suzuki et al. (2009) and Nakabo & Doiuchi (2013) questioned the identification of a photographed specimen of " P. cornutus " (= P. lighti ) in Amaoka et al. (1995), regarding it as P. japonicus (= P. cornutus ) due to its similar color pattern to that species ( Fig. 5 View FIGURE 5 H). However, the identification by Amaoka et al. (1995) is here considered to have been correct, the longitudinal and vertical scale rows of the specimen following regular patterns, and the CLS count determined from the photograph being ca. 94 (within the range of P. lighti ) (Table 1).

On the other hand, color photographs of " P. cornutus " in Ueno (1981) ( Japan) , Iwai & Ito (1986) ( Japan), Chen et al. (1997) ( southern China ), Sakamoto (1998) ( Japan) and Lei et al. (1999) ( Macao) were not identifiable due to ambiguous ocular side coloration and unclear squamation, in addition to a lack of any additional morphological data.

A number of descriptions of " P. cornutus " have included lateral line scale (LLS) counts, that were consistent with the range of this species ( Table 5 View TABLE 5 ), i.e., 100–108 ( Starks & Thompson, 1910), 95 ( Wang, 1933), 98–110 ( Norman, 1934), 93–118 ( Okada et al., 1935), 93–118 ( Chyung, 1954, 1977), 93–118 ( Okada, 1955), 98–110 ( Zheng, 1962), 92–102 ( Tchang & Wang, 1963), 92–105 ( Fitch, 1963), 97–103 (Ochiai, 1965), 97–103 ( Masuda et al., 1975), 98–110 ( Shen, 1982, 1984a, 1993), 98–110 ( Chen & Yu, 1984), 104 ( Matsuura, 1985), 103–109 ( Miao, 1985), 92–113 ( Chen, 1987), 98–101 ( Xu, 1990), 102–106 ( Lindberg & Fedorov, 1993), 93–118 ( Kim & Kang, 1993), 92–102 ( Cheng, 1997), 92–102 ( Wang, 2001), 103–109 ( Wu, 2002), 100–125 ( Kim et al., 2004), 93–118 ( Kim I. et al., 2005), 103–109 ( Ni & Wu, 2006), 98–110 ( Sokolovsky et al., 2009), 103–109 (Chen & Yang, 2013) and 98–110 ( Liu et al., 2013). Because other meristic counts included (e.g., DFR and AFR) also conformed to the ranges of P. lighti (see summary of Yamada et al., 2007), the materials on which the above were based are regarded as that species.

Although Hubbs' (1915) description of " P. cornutus " did not record LLS counts, it included raw measurement data of some body regions for 7 specimens from Swatow , China and waters near Kagoshima , Japan. Calculated proportions from that data for ocular side pectoral fin length and upper eye horizontal diameter resulted in 9.4–15.9 % of SL (average 12.9) and 29.2–33.3 % of HL (average 31.5), respectively, both corresponding closely with figures for P. lighti (Tables 1, 4), which species they apparently represented.

However, the specific identities of specimens listed as " P. cornutus " in the following, could not be determined due to the lack of morphological information: Rutter (1897) [China (Swatow)], Jordan & Starks (1904, 1905) [ Japan ( Suruga Bay ) and Korea], Smith & Pope (1906) [ Japan (Kagoshima and Yamagawa)] , Jordan et al. (1913) [ Japan (Hakodate to Nagasaki)] , Jordan & Metz (1913) [Korea (Fusan)], Reeves (1927) [ China (Swatow, Amoy)], Mori (1928) [Korea (Fusan)], Fowler (1929) ( Hong Kong), Chu (1931) ( China), Schmidt (1931a) [ Japan (Obama)], Schmidt (1931b) [ Japan (Nagasaki and Misaki) and Korea (Fusan)], Schmidt (1931c) [ Korea (Fusan)], Okada & Matsubara (1938) ( Japan, China, Korea and Taiwan), Kuronuma (1940) [ Japan (from Hokkaido to Kyushu)], Herre (1945) [ China (Tinghai and Hong Kong)], Mori (1952), [Korea (Fusan, Kunsan)], Matsubara (1955), ( Japan, Korea, Taiwan and China), Mori (1956) [ Japan (Kasumi and Hamada)], Kamohara (1964) [ Japan (Kochi Pref.)], Hayashi & Ito (1974) [ Japan (Sagami Bay)], Fuwa (1976) [ Japan (Shibushi Bay)], Hayashi & Nishiyama (1980) [ Japan ( Sagami Bay )], Yoo & Lee (1995) (recorded as Pleuronichthys cornuta ) [Korea (Cheju I.)], Sokolovskaya et al. (1998) [ Russia (Peter the Great Bay )], Amaoka (2000) ( South China Sea ), Evseenko (2003, 2004) [ Russia (Peter the Great Bay ) and western North Pacific ], Chen (2004) [ Taiwan (Penghu Is.)], Shao et al., (2008) ( northern South China Sea ) .

Notwithstanding the unknown specific identities of " P. cornutus " material, the following reports are considered likely to include at least P. lighti , due to the dominance of that species over P. cornutus at their collection localities (including regions around Tokyo and Ise Bays, and the Seto Inland Sea): Steindachner (1896) [ Japan (Kobe or Nagasaki)], Jordan & Snyder (1900, 1901) [ Japan (Hakodate, Tokyo and Yokohama)], Snyder (1912) [ Japan (Hakodate, Tokyo and Tsuruga)], Jordan & Thompson (1914) [ Japan (Osaka and Misaki)], Matsuura & Izuka (1920) [ Japan (Takamatsu)], Jordan & Hubbs (1925) [ Japan (Tokyo, Kobe, Toba, Kagoshima, Mikawa Bay, Misaki and Fukui)], Kataoka & Tomida (1981) [ Japan (various sites around Mie Pref.)].

On the other hand, although records of " P. cornutus " by Shinohara et al. (2001) [ Japan ( Tosa Bay )] and Hagiwara & Kimura (2005) [ Japan (Tateyama Bay)] were presently not identifiable, they can be identified when examining their registered museum (voucher) specimens listed therein.

Regarding other synonymies, although Bleeker (1854 –57, 1859) listed " Platessa cornuta " (listing material from Nagasaki, Japan in each case), the latter could not be identified as either of the two Pleuronichthys species owing to insufficient morphological information. The genus Heteroprosopon , which was established by Bleeker (1862) for a monotypic taxon ( Platessa cornuta ), is regarded as a synonym of Pleuronichthys , potentially including either or both of the two Pleuronichthys species considered here.

Specimens comprising " Parophrys cornuta ," recorded by Günther (1862) and Otaki (1897), were identifiable as Pleuronichthys lighti due to the respective descriptions and illustrations of their ocular side coloration patterns. Nyström's (1887) specimens, however, were identified on the basis of 61 AFRs, a count peculiar to P. lighti ( Table 5 View TABLE 5 , see also Yamada et al., 2007). Günther's (1880) specimens, collected from Kobe (Seto Inland Sea), where P. cornutus does not occur, must therefore have been P. lighti , whereas specimens reported by Bleeker (1872) (China), and Namiye (1881) and Ishikawa & Matsuura (1897) (both Niigata, Japan) are now unidentifiable due to a lack of specific information.

Suzuki, S., Kawashima, T. & Nakabo. T. (2009) Taxonomic review of East Asian Pleuronichthys (Pleuronectiformes: Pleuronectidae), with description of a new species. Ichthyological Research, 56 (3), 276 - 291. http: // dx. doi. org / 10.1007 / s 10228 - 009 - 0100 - 7

Yokogawa, K., Ogihara, G. & Watanabe, K. (2014) Identity of the lectotype of the East Asian flatfish Pleuronichthys cornutus (Temminck and Schlegel 1846) and reinstatement of Pleuronichthys lighti Wu 1929. Ichthyological Research, 61 (4), 385 - 392. http: // dx. doi. org / 10.1007 / s 10228 - 014 - 0405 - z

Watanabe, K., Numachi, K., Ueta, Y. & Ishida, Y. (1996 b) Morphological characteristics in the two forms of flatfish, genus Pleuronichthys, in Kiisuido Channel waters off the Pacific coast of central Japan. Nippon Suisan Gakkaishi, 62 (1), 23 - 27. [in Japanese with English abstract]

Yamada, U., Tokimura, M., Horikawa, H. & Nakabo, T. (2007) Fishes and Fisheries of the East China and Yellow Seas. Tokai University Press, Hadano, 1262 pp. [in Japanese]

Amaoka, K., Nakaya, K. & Yabe, M. (1995) The Fishes of Northern Japan. Kita-Nihon Kaiyo Center Co., Ltd., Sapporo, 390 pp. [in Japanese]

Chen, Q., Cai, Y. & Ma, X. (1997) Fishes from Nansha Islands to South China Coastal Waters 1. Science Press, Beijing, 202 pp. [in Chinese and English]

Lei, K., Yang, T., Leung, C. & Huang, X. (1999) Fishes of Macao. Maritime Museum, Association for Promotion of Science and Technology of Macao, Macao, 236 pp. [in Chinese and English]

Sokolovskaya, T. G., Sokolovsky, A. S. & Sobolevsky, E. I. (1998) A list of fishes of the Peter the Great Bay (the Sea of Japan). Voprosy Ikhtiologii, 38 (1), 5 - 15. [in Russian]

Shao, K., Ho, H., Lin, P., Lee, P., Lee, M., Tsai, C., Liao, Y., Lin, Y., Chen, J. & Yeh., H. (2008) A checklist of the fishes of southern Taiwan, Northern South China Sea. The Raffles Bulletin of Zoology, Supplement 19, 233 - 271.

Shinohara, G., Endo, H., Matsuura, K., Machida, Y. & Honda, H. (2001) Annotated checklist of the deepwater fishes from Tosa Bay, Japan. National Science Museum Monographs, 20, 283 - 343.

Abe, T. (Ed.) (1987) Illustrated Fishes of the World in Colour. Hokuryu-kan, Tokyo, 1029 pp. [in Japanese]

Amaoka, K. (2000) Family Pleuronectidae. In: Randall, J. E. & Lim, K. K. P. (Eds.), A checklist of the fishes of the South China Sea. The Raffles Bulletin of Zoology, Supplement (8), National University of Singapore, pp. 645 - 645.

Bleeker, P. (1854 - 57) Nieuwe nalezingen op de ichthyologie van Japan. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen, 26, 1 - 132. [8 plates. (in Dutch)]

Bleeker, P. (1862) Sur quelques genres de la famille des Pleuronectoides. Verslagen en Mededeelingen der Koninklijke Akademie van Wetenschappen, Afdeling Natuurkunde, 13, 422 - 429. [in French and Latin]

Bleeker, P. (1872) Memoire sur la faune ichthyologique de Chine. Nederlandsch Tijdschrift voor de Dierkunde, 4, 113 - 154. [in French]

Boeseman, M. (1947) Revision of the fishes collected by Burger and von Siebold in Japan. Zoologische Mededelingen, 28, 1 - 242. [plates. 1 - 5.]

Burgess, W. E. & Axelrod, H. R. (1974) Fishes of Taiwan and Adjacent Waters (Pacific Marine Fishes, Book 4). T. F. H. Publications Inc. Ltd., Hong Kong, 1110 pp.

Cao, Y. (Ed.) (2010) Atlas of Sea Fishes of China. Encyclopedia of China Publishing House, Beijing, 197 pp. [in Chinese]

Chen, J. T. F. & Weng, H. T. C. (1965) A review of the flatfishes of Taiwan. Biological Bulletin of Tunghai University, 25 / 27, 1 - 103.

Chen, J. T. F. & Yu, M. (1984) A Synopsis of the Vertebrates of Taiwan, revised and enlarged edition, Volume 2. The Commercial Press, Taipei, 633 pp. [in Chinese]

Chen, J. (1987) Order Pleuronectiformes. In: Liu, C. & Qin, K. (Eds.), Fauna Liaoningica, Pisces. Liaoning Science and Technology Press, Shenyang, pp. 416 - 445. [in Chinese]

Chen, C. (2003) Fishes of Penghu (FRI Special Publication No. 3). Fisheries Research Institute, Council of Agriculture, Keelung, 379 pp. [in Chinese]

Chen, C. (2004) Checklist of the Fishes of Penghu (FRI Special Publication No. 4). Fisheries Research Institute, Council of Agriculture, Keelung, 175 pp. [in Chinese]

Cheng, Q. & Zheng, B. (Eds.) (1987) Systematic Synopsis of Chinese Fishes. Science Press, Beijing, 1458 pp. [in Chinese]

Cheng, Q. (1997) Order Pleuronectiformes. In: Cheng, Q. & Zhou, C. (Eds.), The fishes of Shandong Province. Shandong Science and Technology Press, Jinan, pp. 442 - 467. [in Chinese]

Choi, Y., Kim, J. & Park, J. (2002) Marine Fishes of Korea (Korean Nature Series 19). Kyo-Hak Publishing Co., Ltd, Seoul, 645 pp. [in Korean]

Choi, Y. (2007) Sea Fishes for Food (Kyo-hak mini guide 7). Kyo-Hak Publishing Co., Ltd, Seoul, 314 pp. [in Korean]

Choi, Y., Kim, J., Lee, H., Yoon, S., Kim, D. & Lee, S. (2011) A Pictorial Guide to Fishes with Tidbits Manufactured by Teachers. Kyo-Hak Publishing Co., Ltd., Seoul, 279 pp. [in Korean]

Chu, Y. (1931) Index piscium sinensium. Biological Bulletin St. John's University, 1, i - iv + 1 - 290.

Chyung, M. (1954) Korean Fishes. Ministry of Commerce and Industry, Republic of Korea, Seoul, 517 pp. [in Korean]

Chyung, M. (1977) The fishes of Korea. Iljisa, Seoul, 727 pp. [in Korean]

Eschmeyer, W. N. (Ed.) (2015) Catalog of Fishes, Online Version. California Academy of Science, San Francisco. Available from: http: // research. calacademy. org / research / ichthyology / catalog / fishcatmain. asp (accessed 2 November 2015)

Evseenko, S. A. (2003) An annotated catalogue of pleuronectiform fishes (order Pleuronectiformes) of the seas of Russia and adjacent countries. Journal of Ichthyology, 43 (Supplemental 1), S 57 - S 74.

Evseenko, S. A. (2004) Family Pleuronectidae Cuvier 1816, righteye flounders. California Academy of Science, Annotated Checklists of Fishes, 37, 1 - 37.

Fitch, J. E. (1963) A review of the fishes of the genus Pleuronichthys. Los Angeles County Museum Contributions in Science, 76, 3 - 33.

Fowler, H. W. (1929) Notes on Japanese and Chinese fishes. Proceedings of the Academy of Natural Sciences of Philadelphia, 81, 589 - 616.

Fowler, H. W. (1933) A synopsis of the fishes of China, part V. Hong Kong Naturalist, 4, 156 - 175.

Fuwa, S. (1976) On the small trawl in Shibushi Bay fishing ground and catch. Memoirs of Faculty of Fisheries, Kagoshima University, 25 (1), 125 - 135. [in Japanese with English abstract]

Gunther, A. (1862) Catalogue of the Acanthopterygii, Pharyngognathi and Anacanthini in the collection of the British Museum. Catalogue of the Fishes in the British Museum, 4, 534 - 534.

Gunther, A. (1880) Report on the shore fishes procured during the voyage of H. M. S. Challenger in the years 1873 - 1876. In: Report on the scientific results of the voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 1 (6), 1 - 82. [plates. 1 - 32.]

Hagiwara, K. & Kimura, K. (2005) Catalogue of fishes of the Yokosuka City Museum (IV), Sagami Bay Marine Biological Research Club's fish collection of Hasama, Tateyama Bay. Miscellaneous Report of Yokosuka City Museum, (29), 1 - 34. [plates. 1 - 8. (in Japanese with English abstract)]

Hayashi, M. & Ito, T. (1974) Fishes collected and observed from the coastal waters of Tenjin-jima and Kasa-jima near Sajima, Yokosuka City. Miscellaneous Report of Yokosuka City Museum, 20, 37 - 50. [in Japanese with English abstract] Hayashi, M. & Nishiyama, K. (1980) Fishes collected by the stationary net in the west coast of Sagami Bay, the catalogue of fish collection of Sagami Bay- 1. Natural History Report of Kanagawa, 1, 15 - 27. [in Japanese]

Herre, A. W. C. T. (1945) Marine fishes from the Chusan Archipelago and the Chinese coast. Lingnan Science Journal, 21, 107 - 122.

Hiyama, Y & Yasuda, F. (1961) Nippon Suisan Gyofu (Scientific Illustrations of Food Fish of the World). Central Research Laboratory of Nippon Suisan Co., Ltd., Tokyo, 155 pp. [210 plates. (in Japanese)]

Hiyama, Y. & Yasuda, F. (1971) Japanese Fishes. Kodansha Co., Ltd., Tokyo, 342 pp. [in Japanese]

Hubbs, C. L. (1915) Flounders and soles from Japan collected by the United States Bureau of fisheries steamer Albatross in 1906. Proceedings of the United States National Museum, 48, 449 - 496. [plates. 25 - 27.]

Igarashi, S. & Ito, T. (2013) Feeding habits of congeneric flatfishes Pleuronichthys cornutus and P. japonicus off Fukushima Prefecture. Bulletin of Fukushima Prefectural Fisheries Experimental Station, 16, 9 - 14. [in Japanese]

Ishikawa, C. & Matsuura, K. (1897) Preliminary Catalogue of Fishes Including Dipnoi, Cyclostomi and Cephalochorda in the Collection of the Natural History Department, Imperial University. Imperial Museum, Tokyo, 64 pp. [in Japanese with English preface]

Ishikawa, H. & Senou, H. (2010) Grand Illustrated Encyclopedia of Marine Fishes. Nitto Shoin Honsha Co., Ltd., Tokyo, 399 pp. [in Japanese]

Iwai, T. & Ito, K. (1986) Identification Guide, Illustrated Encyclopedia of Fishes- 2. Hoikusha, Osaka, 215 pp. [in Japanese]

Jo, G. & Myoung, J. (2013) A Pictorial Guide to Marine Fishes. Bori Publishing Co., Ltd., Paju, 351 pp. [in Korean]

Jordan, D. S. & Snyder, J. O. (1900) A list of fishes collected in Japan by Keinosuke Otaki, and by the United States steamer Albatross, with descriptions of fourteen new species. Proceedings of the United States National Museum, 23, 335 - 380. [plates. 9 - 20.]

Jordan, D. S. & Snyder, J. O. (1901) List of fishes collected in 1883 and 1885 by Pierre Louis Jouy and preserved in the United States National Museum, with descriptions of six new species. Proceedings of the United States National Museum, 23, 739 - 769. [plates. 31 - 38.]

Jordan, D. S. & Starks, E. C. (1904) List of fishes dredged by the steamer Albatross off the coast of Japan in the summer of 1900, with descriptions of new species and a review of the Japanese Macrouridae. Bulletin of the United States Fish Commission, 22, 577 - 630. [plates. 1 - 8.]

Jordan, D. S. & Starks, E. C. (1905) On a collection of fishes made in Korea, by Pierre Louis Jouy, with descriptions of new species. Proceedings of the United States National Museum, 28, 193 - 212.

Jordan, D. S. & Starks, E. C. (1906) A review of the flounders and soles of Japan. Proceedings of the United States National Museum, 31, 161 - 246.

Jordan, D. S. & Metz, C. W. (1913) A catalog of the fishes known from the waters of Korea. Memoirs of the Carnegie Museum, 6 (1), 1 - 65. [plates. 1 - 10.]

Jordan, D. S. & Thompson, W. F. (1914) Record of the fishes obtained in Japan in 1911. Memoirs of the Carnegie Museum, 6 (4), 205 - 313. [plates. 24 - 42.]

Jordan, D. S. & Hubbs, C. L. (1925) Record of fishes obtained by David Starr Jordan in Japan, 1922. Memoirs of the Carnegie Museum, 10 (2), 93 - 346. [plates. 5 - 12.]

Kamohara, T. (1952) Revised descriptions of the offshore bottomfishes of Prov. Tosa, Shikoku, Japan. Reports of the Kochi University, Natural Science, 3 (3), 1 - 122.

Kamohara, T. (1964) Revised catalogue of fishes of Kochi Prefecture, Japan. Reports of the Usa Marine Biological Station, 11 (1), 1 - 99.

Kamohara, T. & Okamura, O. (1985) Colored Illustrations of the Marine Fishes of Japan, Volume. II. Hoikusha Publishing, Osaka, 160 pp. [in Japanese]

Kataoka, T. & Tomida, Y. (1981) Fish fauna of Mie Prefecture. Bulletin of the Mie Prefectural Museum, Natural Science, 3, 1 - 110. [in Japanese]

Kato, Y. & Fujio, Y. (1979) On the sibling species of the flounder, Pleuronichthys cornutus. Fish Genetics and Breeding Science, 4, 10 - 12. [in Japanese]

Kim, I. & Kang, U. (1993) Coloured Fishes of Korea. Academy Publishing Company, Seoul, 477 pp. [in Korean]

Kim, I. & Youn, C. (1994) Taxonomic revision of the flounders (Pisces: Pleuronectidae) from Korea. Korean Journal of Ichthyology, 6 (2), 99 - 132. [in Korean with English summary]

Kim, I. (1997) Illustrated Encyclopedia of Fauna and Flora of Korea, Volume 37, Freshwater Fishes. Korean Ministry of Education, Seoul, 629 pp. [in Korean with English preface]

Kim, Y., Myoung, J., Kim, Y., Han, K., Kang, C. & Kim, J. (2001) The Marine Fishes of Korea, 1 st Edition. Hanguel, Pusan, 382 pp. [in Korean]

Kim, Y., Han, K, Kang, C. & Kim, J. (2004) Commercial Fishes of the Coastal and Offshore Waters in Korea, 2 nd Edition.

Kim, I., Choi, Y., Lee, C., Lee, Y., Kim, B. & Kim, J. (2005) Illustrated Book of Korean Fishes. Kyo-Hak Publishing Co., Ltd., Seoul, 615 pp. [in Korean]

Kitajima, C., Hayashida, G. & Yasumoto, S. (1987) Ambicoloration and albinism in hatchery-reared frog flounder juveniles, Pleuronichthys cornutus. Bulletin of Nagasaki Prefectural Institute of Fisheries, 13, 17 - 23. [in Japanese with English abstract]

Konishi, H. & Nakabo, T. (2007) Color Guide to the Japanese Fishes for Sportfishermen. Enterbrain Inc., Tokyo, 400 pp. [in Japanese]

Kuronuma, K. (1940) Check list of the flatfishes taken by the Kisen-sokobibi along the Pacific coast of Japan from Hokkaido to Kyushu. Suisan Kenkyushi, 35 (8), 211 - 216.

Li, M. & Zhang, H. (1991) Fish Biology in the Bohai Sea. China Science and Technology Press, Beijing, 141 pp. [in Chinese]

Li, S. & Wang, H. (1995) Fauna Sinica, Ostichthyes, Pleuronectiformes. Science Press, Beijing, 433 pp. [in Chinese]

Lindberg, G. U. & Fedorov, V. V. (1993) Fishes of the Sea of Japan and the adjacent parts of the Sea of Okhotsk and the Yellow Sea, Part 6, Teleostomi, Osteichthyes, Actinopterygii, XXXI Pleuronectiformes. Handbook on the Identification of Animals, Zoological Institute of the Russian Academy, 166, 1 - 272. [in Russian]

Liu, M., Chen, X. & Yang, S. (2013) Marine Fishes of Southern Fujian, China, Volume 1. China Ocean Press, Beijing, 297 pp. [in Chinese]

Masaki, Y. & Ito, H. (1984) Stock identification of finespotted flounder, Pleuronichthys cornutus (Temminck and Schlegel) in the western Seto Inland Sea. Bulletin of Nansei Regional Fisheries Research Laboratory, 16, 11 - 51. [in Japanese with English abstract]

Masuda, H., Araga, C. & Yoshino, T. (1975) Coastal Fishes of Southern Japan, 1 st Edition. Tokai University Press, Tokyo, 379 pp. [in English and Japanese]

Masuda, H. & Allen, G. R. (1987). Sea Fishes of the World (Indo-Pacific Region). Yama-to-keikokusha, Tokyo, 527 pp. [in Japanese]

Masuda, H. & Kobayashi, A. (1994) Grand Atlas of Fish Life Modes. Tokai University Press, Tokyo, 465 pp. [in Japanese]

Matsubara, K. (1955) Fish Morphology and Hierarchy. Ishizaki-Shoten, Tokyo, 789 pp. [in Japanese]

Matsuoka, M., Taniguchi, N., Fujita, H., Kitajima, T. & Tokimura, M. (1989) Comparative studies on two types of frog flounder, Pleuronichthys cornutus, in the East China Sea and the Yellow Sea - I, distributional, morphological and genetic differences. Bulletin of Seikai Regional Fisheries Research Laboratory, 67, 23 - 36. [in Japanese with English abstract]

Matsuoka, M. (1996 a) Comparative studies on two types of frog flounder, Pleuronichthys cornutus, in the East China Sea and the Yellow Sea - II, age and growth. Bulletin of Seikai National Fisheries Research Institute, 74, 47 - 54. [in Japanese with English abstract]

Matsuoka, M. (1996 b) Comparative studies on two types of frog flounder, Pleuronichthys cornutus, in the East China Sea and the Yellow Sea - III, sexual maturation and seasonal change of distribution. Bulletin of Seikai National Fisheries Research Institute, 74, 55 - 62. [in Japanese with English abstract]

Matsuura, K. & Izuka, A. (1920) Catalogue of the Zoological Specimens Exhibited in the Natural History Department, Tokyo Imperial Museum (Vertebrata). Tokyo Imperial Museum, Tokyo, 192 pp. [in Japanese]

Matsuura, K. (1985) Family Pleuronectidae. In: Okamura, S. & Kitajima, T. (Eds.), Fishes of the Okinawa Trough and the Adjacent Waters II. Japan Fisheries Resource Conservation Association, Tokyo, pp. 618 - 623. [in Japanese]

Miao, X. (1985) Order Pleuronectiformes. In: Chu, Y. (Ed.), The Fishes of Fujian Province, Part II. Fujian Science and Technology Press, Fuzhou, pp. 518 - 563. [in Chinese]

Minami, T. (1988 a) Pleuronichthys cornutus. In: Okiyama M. (Ed.), An Atlas of Early Stage Fishes in Japan, 1 st Edition. Tokai University Press, Tokyo, pp. 938 - 940. [in Japanese]

Minami, T. (2014 a) Pleuronichthys cornutus. In: Okiyama M. (Ed.), An Atlas of Early Stage Fishes in Japan, 2 nd Edition. Tokai University Press, Hadano, pp. 1462 - 1463. [in Japanese]

Mochizuki, K. (1985) Pleuronichthys cornutus. In: Konishi, K. (Ed.), Grand Illustrated Encyclopedia of Fishes. Weekly Fishing Sunday Co., Ltd., Osaka, pp. 126 - 127. [in Japanese]

Mori, T. (1928) A catalogue of the fishes of Korea. Journal of the Pan-Pacific Research Institution, 3 (3), 3 - 8.

Mori, T. (1952) Check list of the fishes of Korea. Memoirs of the Hyogo University of Agriculture, 1 (3), 1 - 228.

Mori, T. (1956) Fishes of San-in District including Oki Islands and its adjacent waters (Southern Japan Sea). Memoirs of the Hyogo University of Agriculture, 2 (3), 1 - 62. [in Japanese]

Myoung, J., Km, B., Lee, S. & Jeon G. (2002) The Sea Fishes of Korea. Darakwon Press, Seoul, 287 pp. [in Korean]

Myoung, J., Ko, D. & Kim, J. (2015) Fishes of Cheju Island. Jisungsa, Seoul, 295 pp. [in Korean]

Nakabo, T. (1995) Pleuronichthys cornutus, Pleuronichthys sp. In: Konishi, K. (Ed.), Fishes, New Color Guide for Sports Fishermen. Weekly Fishing Sunday Co., Ltd., Osaka, pp. 204 - 205. [in Japanese]

Nakabo, T. (2000) Family Pleuronectidae. In: Nakabo, T. (Ed.), Fishes of Japan with Pictorial Keys to the Species, 2 nd Edition. Tokai University Press, Tokyo, pp. 1371 - 1379. [in Japanese]

Nakabo, T. & Doiuchi, R. (2013) Family Pleuronectidae. In: Nakabo, T. (Ed.), Fishes of Japan with Pictorial Keys to the Species, 3 rd Edition. Tokai University Press, Hadano, pp. 1675 - 1683. [in Japanese]

Namiye, M. (1881) Classified Catalogue of the Specimens of Vertebrates in the Collections of Kiyoiku Hakubutsukwan (Educational Museum). Educational Museum, Tokio, 117 pp. [in Japanese]

Ni, Y. & Wu, H. (2006) Order Pleuronectiformes. In: Ni, Y. & Wu, H. (Eds.), Fishes of Jiangsu Province. China Agriculture Press, Beijing, pp. 800 - 838. [in Chinese]

Norman, J. R. (1934) A Systematic Monograph of the Flatfishes (Heterosomata), vol. I, Psettodidae, Bothidae, Pleuronectidae. The Trustees of the British Museum, London, 459 pp.

Nozawa, M. & Kato, F. (1981) Morphological comparison of two types of frog flounder, Pleuronichthys cornutus, collected off Tottori Prefecture, in the Japan Sea. Bulletin of Japan Sea Regional Fisheries Research Laboratory, 32, 1 - 8. [in Japanese with English abstract]

Nystrom, E. (1887) Redogorelse foorden Japanska Fisksalmingen i Upsala Universitets Zoologiska Museum. Bihang till Kongl, Svenska Vetenskaps-Akademiens Handlingar, 13 (4), 1 - 54. [in Swedish]

Ochiai, A. (1986) Order Pleuronectiformes. In: Abe, T. (Ed.), A Definitive Grand Illustrated Encyclopedia of Creatures, Fishes. Sekai-bunkasha, Tokyo, pp. 310 - 317. [in Japanese]

Ohashi, Y. & Motomura, H. (2011) Pleuronectiform fishes of northern Kagoshima Prefecture, Japan. Nature of Kagoshima, 37, 71 - 118. [in Japanese]

Okada, Y., Uchida, K. & Matsubara, K. (1935) Color Atlas of Fishes of Japan. Sanseido Co., Ltd., Tokyo, 471 pp. [83 plates. (in Japanese)]

Okada, Y. & Matsubara, K. (1938) Keys to the Fishes and Fish-like Animals of Japan, Including Kuril Islands, Southern Sakhalin, Bonin Islands, Ryukyu Islands, Korea and Formosa, 1 st Edition. Sanseido Co., Ltd., Tokyo, 584 pp. [in Japanese]

Okada, Y. (1955) Fishes of Japan, Illustrations and Descriptions of Fishes of Japan. Maruzen, Tokyo, 434 pp. [28 plates.]

Otaki, K. (1897) Identification of pleuronectids from Japan. Research Report for Fisheries by Ministry of Agriculture and Commerce, 6 (1), 17 - 36. [plates. 5 - 8. (in Japanese)]

Reeves, C. D. (1927) A catalogue of the fishes of north-eastern China and Korea. Journal of the Pan-Pacific Research Institution, 2 (3), 3 - 16.

Rutter, C. M. (1897) A collection of fishes obtained in Swatow, China, by Miss Adele M. Fielde. Proceedings of the Academy of Natural Sciences of Philadelphia, 49, 56 - 90.

Sakamoto, K. & Nakabo, T. (1984) Pleuronichthys cornutus. In: Masuda, H., Amaoka, K., Araga, C., Uyeno, T. & Yoshino, T. (Eds.), The Fishes of Japanese Archipelago, 1 st Edition. Tokai University Press, Tokyo, pp. 337, 352. [plate. 315. (in Japanese and English)]

Sakamoto, K. & Nakabo, T. (1988 a) Pleuronichthys cornutus. In: Masuda, H., Amaoka, K., Araga, C., Uyeno, T. & Yoshino, T. (Eds.), The Fishes of Japanese Archipelago, 2 nd Edition. Tokai University Press, Tokyo, pp. 337, 352. [plate. 315. (in Japanese and English)]

Sakamoto, K. (1998) Order Pleuronectiformes. In: Nakabo, T. & Mochizuki, K. (Eds.), The Encyclopedia of Animals in Japan, vol. 6, Bony Fishes. Heibonsha Limited, Publishers, Tokyo, pp. 184 - 187. [in Japanese]

Sanada, Y. (1997) Pleuronichthys cornutus. In: Ikehara, K. (Ed.), Fishes of the Seto Inland Sea. Fisheries Developing Committee of the Seto Inland Sea, Kobe, pp. 72 - 72. [in Japanese]

Schmidt, P. J. (1931 a) A list of fishes, collected in Japan and China by Dr. A. Bunge and N. Grebnitzky. Bulletin de l'Academie des Sciences, Leningrad, Ser. 7, 101 - 123.

Schmidt, P. J. (1931 b) Fishes of Japan, collected in 1901. Transactions of the Pacific Committee of the Academy of Sciences of the U. S. S. R., 2, 1 - 176.

Schmidt, P. J. (1931 c) On a collection of flat-fishes from Fusan (Korea). Doklady Akademii Nauk SSSR, Seriya A, 12, 313 - 320.

Fisheries College of Shanghai, Xiamen University & Fisheries Research Institute of Fujian (Eds). (1980) Economic Marine Fishes of Fujian. Fujian Science & Technology Publishing House, Fuzhou, 162 pp. [in Chinese]

Shao, K. & Lin, P. (1991) Marine Fishes for Fishing (Fishes Inhabited Sandy Coastal Waters). Dujia Publishing Co. Ltd., Taipei, 238 pp. [in Chinese]

Shen, S. (1967) Studies on the flatfishes (Pleuronectiformes or Heterosomata) in the adjacent waters of Hong Kong. Quarterly Journal of the Taiwan Museum, 20 (1 / 2), 149 - 281.

Shen, S. (1982) Study on the pleuronectid fishes (family Pleuronectidae) from Taiwan. Quarterly Journal of the Taiwan Museum, 35 (3 / 4), 197 - 213.

Shen, S. (1984 a) Coastal Fishes of Taiwan. Department of Zoology, Taiwan University, Taipei, 190 pp. [152 plates. (in English and Chinese)]

Shen, S. (1984 b) Synopsis of Fishes of Taiwan. Southern Materials Center, Inc., Taipei, 533 pp. [in Chinese]

Shen, S. (Ed.) (1993) Fishes of Taiwan. Department of Zoology, National Taiwan University, Taipei, 960 pp. [in Chinese]

Shen, S. & Wu, G. (Eds.) (2011) Fishes of Taiwan. National Museum of Marine Biology & Aquarium, Checheng Township, Pingtung County, 896 pp. [in Chinese]

Shimizu, T. (2001) An annotated list of the coastal fishes from Iyo City, Ehime Prefecure, Japan [sic]. Bulletin of the Tokushima Prefectural Museum, 11, 17 - 99. [in Japanese with English abstract]

Shirano, K. (Ed.) (1979) Geikai Gyofu, New Edition. Shimazu Press, Tokyo, 230 pp. [in Japanese]

Smith, H. M. & Pope, T. E. B. (1906) List of fishes collected in Japan in 1903, with descriptions of new genera and species. Proceedings of the United States National Museum, 31, 459 - 499.

Snyder, J. O. (1912) Japanese shore fishes collected by the United States Bureau of Fisheries steamer Albatross expedition of 1906. Proceedings of the United States National Museum, 42, 399 - 450. [plates. 51 - 61.]

Sokolovsky, A. S., Dudarev, V. A., Sokolovskaya, T. G. & Solomatov, S. F. (2007) Sea fishes of the Russian waters of the Sea of Japan, Annotated and Illustrated Catalogue. Dalnauka, Vladivostok, 199 pp. [in Russian]

Sokolovsky, A. S., Sokolovskaya, T. G. & Yakovlev, Y. M. (2009) Fishes of the Peter the Great Bay. Dalnauka, Vladivostok, 375 pp. [in Russian]

Starks, E. C. & Thompson, W. F. (1910) A review of the flounders belonging to the genus Pleuronichthys. Proceedings of the United States National Museum, 38, 277 - 287.

Steindachner, F. (1896) Bericht uber die wahrend der Reise Sr. Maj. Schiff Aurora von Dr. C. Ritter v. Microszewski in den Jahren 1895 und 1896, gesammelten Fische. Annalen des Naturhistorischen Hofmuseums, 11, 197 - 230. [plate. 4. (in German)]

Su, J. (Ed.) (1995) Ichthyology and Aquaculture of Marine Fishes, 2 nd Edition. China Agriculture Press, Beijing, 507 pp. [in Chinese]

Suzuki, K. & Kataoka, T. (1997) Marine Fishes of Mie Prefecture, Japan. Toba Aquarium, Toba, 297 pp. [152 plates. (in Japanese)]

Taki, Y., Kohno, H., Sakamoto, K. & Hosoya, K. (Eds.) (2005) Illustrated Fishes in Colour Revised Edition. Hokuyu-kan, Tokyo, 242 pp. [in Japanese]

Tanaka, S. & Abe, T. (1955) Pictorial Descriptions of Thousand Species of Useful Fishes. Morikita Publishing Co., Ltd., Tokyo, 294 pp. [in Japanese]

Tchang, T. & Wang, W. (1963) Order Pleuronectiformes. In: Chu, Y., Tchang, T. & Cheng, Q. (Eds.), Fishes of the East Sea.

Tchang, T., Cheng, Q., Zheng, B., Li, S., Zheng, W. & Wang, W. (1994) Fishes of the Yellow and Bohai Seas. The Sueichan Press, Keelung, 353 pp. [in Chinese]

Tomiyama, I., Abe, M. & Tokioka, T. (1958) Encyclopaedia Zoological Illustrated in Colours, Volume II. Hokuryu-kan, Tokyo, 478 pp. [in Japanese]

Tsuji, K. (2013) An annotated list of fishes from Hakata Island, Ehime Prefecture. Bulletin of the Tokushima Prefectural Museum, 23, 1 - 21. [in Japanese]

Ueno, T. (1981) Pleuronichthys cornutus. In: Ochiai, A. (Ed.), Gakken Illustrated Nature Encyclopedia, the Fishes of Japan. Gakushu-kenkyusha, Tokyo, pp. 159, 273. [in Japanese]

Uyeno, T. & Sakamoto, K. (2011) Nippon Suisan Gyofu, Scientific Illustrations of Food Fish of the World. Nippon Suisan Kaisha, Ltd., Tokyo, 364 pp. [in Japanese and English]

Wang, K. (1933) Study of the teleost fishes of coastal region of Shangtung, part 1. Contributions from the Biological Laboratory of the Science Society of China, Zoological Series, 9 (1), 1 - 76.

Wang, Z. (2001) Order Pleuronectiformes. In: Wang, S., Wang, Z., Li, G., Cao, Y., et al. (Eds.), The Fauna of Hebei, Pisces. Hebei Science and Technology Publishing House, Shijiazhuang, pp. 303 - 326. [in Chinese]

Watanabe, K. & Numachi, K. (1988) Biochemical demonstration and morphological examination of the sibling species of the frog flounder in the eastern part of the Seto Inland Sea. Nippon Suisan Gakkaishi, 54 (5), 765 - 772. [in Japanese with English abstract]

Watanabe, K., Tamai, K. & Numachi, K. (1992) Feeding habits and distribution pattern of two forms of flatfishes (Genus Pleuronichthys) in Kiisuido Channel waters off the Pacific coast of central Japan. Nippon Suisan Gakkaishi, 58 (12), 2253 - 2260. [in Japanese with English abstract]

Watanabe, K., Numachi, K., Jo, Y. & Ueta, Y. (1996 a) Sexual maturity, spawning and sex ratio of the two forms of flatfish, genus Pleuronichthys, in Kiisuido Channel Waters off the Pacific Coast of central Japan. Nippon Suisan Gakkaishi, 62 (1), 17 - 22. [in Japanese with English abstract]

Wu, H. (1929) Study of the fishes of Amoy, part 1. Contributions from the Biological Laboratory of the Science Society of China, Zoological Series, 5 (4), i - vi + 1 - 90.

Wu, H. (1932) Contribution a l'Etude Morphologique, Biologique et Systematique des Poissons Heterosomes (Pisces Heterosomata) de la Chine. Theses Presentees a la Faculte des Sciences de l'Universite de Paris, Paris, 179 pp. [in French]

Wu, H. (Ed.) (2002) Icthyotoxic and Medical Fishes of China, New Version. China Agriculture Press, Beijing, 679 pp. [in Chinese]

Xu, C. (1990) Order Pleuronectiformes. In: East China Sea Fisheries Research Institute & Chinese Academy of Fisheries Science, Shanghai Fisheries Research Institute (Eds.), Fishes of Shanghai. Shanghai Science and Technology Press, Shanghai, pp. 357 - 370. [in Chinese]

Yamada, U. (1969) An impression for observation of ambicoloration phenomenon in frog flounder (Pleuronichthys cornutus). Newsletter of Seikai Regional Fisheries Research Laboratory, 30, 2 - 3. [in Japanese]

Yamada, U. (1986) Family Pleuronectidae. In: Yamada, U., Tagawa, M., Kishida, S., Hongo, K. (Eds.), Fishes of the East China Sea and the Yellow Sea. Seikai Regional Fisheries Research Laboratory, Nagasaki, pp. 378 - 401. [in Japanese]

Yamada, U., Shirai, S., Irie, T., Tokimura, M., Deng, S., Zheng, Y., Li, C., Kim, Y. & Kim Y. (1995) Names and Illustrations of Fishes from the East China Sea and the Yellow Sea. Oversea Fishery Cooperation Foundation, Tokyo, 288 pp.

Yao, Z. & Yoshida, N. (2006) Trading Fishes and Shellfishes from China. Seizando-shoten Publishing. Co., Ltd., Tokyo, 142 pp. [in Japanese]

Yokogawa, K. & Watanabe, K. (2011) Squamatological differences between two closely related flatfish species of the genus Pleuronichthys (Pleuronectidae, Pleuronectiformes), with a proposal of unambiguous distinctive characters. Ichthyological Research, 58 (1), 24 - 32. http: // dx. doi. org / 10.1007 / s 10228 - 010 - 0186 - y

Yokogawa, K. (2015) Morphological differences between two closely related East Asian flatfish species of the genus Pleuronichthys, with particular emphasis on growth-related changes. Ichthyological Research, 62 (4), 474 - 486. http: // dx. doi. org / 10.1007 / s 10228 - 015 - 0462 - y

Yokogawa, K. (2016) A long-finned variant of the flounder Pleuronichthys lighti from Tokyo Bay, Japan. Ichthyological Research, 63 (1), 192 - 196. http: // dx. doi. org / 10.1007 / s 10228 - 015 - 0469 - 4

Yoo, J. & Lee, M. (1995) Marine fishes around Cheju Island. Cheju Provincial Office of Education, Cheju, 248 pp. [in Korean]

Yoshino, Y. (1999) Yama-kei Pocket Guide 8, Marine Fishes. Yama-to-keikokusha, Tokyo, 284 pp. [in Japanese]

Youn, C. (2002) Fishes of Korea with Pictorial Key and Systematic List. Academy Publishing Company, Seoul, 747 pp. [in Korean]

Zhang, H., Gao, T., Xu, H. & Li, P. (2011 a) A new record of Pleuronichthys in China: Pleuronichthys japonicus. Periodical of Ocean University of China, 41 (1 / 2), 51 - 54. [in Chinese with English summary]

Zhang, H., Zhang, Y., Gao, T., Li, P. & Xu, H. (2011 b) Genetic identification of two species of Pleuronichthys by DNA barcoding. Chinese Journal of Oceanology and Limnology, 29 (5), 967 - 972. http: // dx. doi. org / 10.1007 / s 00343 - 011 - 0040 - 8

Zhao, S. & Zhong, J. (2006) Colored Encyclopedia of Fishes in Zhoushan Waters. Zhejiang Science and Technology Press, Hangzhou, 200 pp. [in Chinese]

Zheng, B. (1955) Order Pleuronectiformes. In: Tchang, T., Cheng, C., Cheng, B., Li, S., Cheng, W. & Wang, W. (Eds.), Survey Report on Fishes from the Yellow and Bohai Seas. Chinese Academy of Science, Beijing, pp. 274 - 305. [in Chinese]

Zheng, B. (1962) Order Pleuronectiformes. In: Institute of Zoology Academia Sinica, Institute of Oceanology Academia Sinica, Shanghai Fisheries College (Ed.), Fishes of South China Sea, Science Press, Beijing, pp. 950 - 1014. [in Chinese]

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FIGURE 3. Scales of Pleuronichthys lighti (A – C) and P. cornutus (D – F). A, D: ocular side, trunk, area above lateral line; B, E: ocular side, trunk, area below lateral line; C, F: blind side, trunk, area above lateral line. Bars indicate 0.2 mm. Photographs selectively cited from Yokogawa & Watanabe (2011).

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FIGURE 4. Trunk squamation (ocular side) of Pleuronichthys lighti (A, B) and P. cornutus (C, D). A: BSKU 37610; B: SNFR 13740; C: SNFR 1428; D: SNFR 15497 - 2.

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FIGURE 5. Ocular side coloration variations in fresh Pleuronichthys lighti (A – E) and P. cornutus (F – J). A: BSKU 114101; B: BSKU 118425; C: BSKU 118426 (A – C were photographed by K. Yokogawa); D: KAUM – I. 20492; E: KAUM – I. 5064; F, G: Uncatalogued specimens (F and G were photographed by K. Watanabe); H: KAUM – I. 30446; I: KAUM – I. 29267; J: KAUM – I. 4229 (KAUM specimens were photographed by the Kagoshima University Museum).

TABLE 5. Meristic counts of two Pleuronichthys species (other than those shown in TABLE 1).

  Range Average Range Average
Dorsal fin rays 71–83 77.69 67–81 74.24
Caudal fin rays 18–20 19.07 18–20 19.11
Anal fin rays 50–62 57.28 48–58 53.63
Pectoral fin rays (ocular side) 9–13 10.94 9–13 10.92
Pectoral fin rays (blind side) 9–13 10.81 9–12 10.38
Pelvic fin rays (ocular side) 5–7 6.05 6–7 6.02
Pelvic fin rays (blind side) 5–7 5.98 5–6 5.98
Upper limb gill rakers 1 2–4 2.70 2–4 2.70
Lower limb gill rakers 1 4–7 5.42 4–7 5.40
Total gill rakers 1 6–11 8.12 6–11 8.10
Caudal vertebrae 23–26 24.25 23–25 24.24
Total vertebrae 36–39 37.24 35–37 36.29

RMNH

National Museum of Natural History, Naturalis
NFRDI

National Fisheries Research and Development Institute
SCL

St. Cloud State University

Kingdom

Animalia

Phylum

Chordata

Class

Actinopterygii

Order

Pleuronectiformes

Family

Pleuronectidae

Genus

Pleuronichthys

Specimens