Triozidus, Li, 1994

Liao, Yi-Chang, Inoue, Hiromitsu & Percy, Diana M., 2025, The psyllid genus Triozidus Li, 1994 stat. rev. sensu novo (Hemiptera: Psylloidea: Triozidae) in East Asia is redefined with the addition of two new species from Taiwan inducing galls on the leaflet petiolules of Eleutherococcus trifoliatus (Araliaceae), Zootaxa 5727 (1), pp. 113-139 : 117

publication ID	https://doi.org/10.11646/zootaxa.5727.1.8
publication LSID	lsid:zoobank.org:pub:8C71D66D-DACB-43F8-A3F4-6F493B9AF9EB
persistent identifier	https://treatment.plazi.org/id/03D2A373-FFDC-FFAF-FF26-FDB8FA142A30
treatment provided by	Plazi
scientific name	Triozidus
status

Treatment

Key to adults of Triozidus

1. Forewing with Rs vein long and sinuous (terminating distal to a line drawn through termination of vein Cu 1a and bifurcation of vein M); cell cu 1 higher resulting in cell m 1 not being markedly larger than cell cu 1 ( Fig. 9B View FIGURE 9 ); on Eleutherococcus spinosus View in CoL (or other Eleutherococcus spp. where alien................................... Triozidus ukogi ( Shinji, 1940) comb. nov.

- Forewing with Rs short and more or less evenly curving to the fore margin (terminating at or proximal to a line drawn through termination of vein Cu 1a and bifurcation of vein M); cell cu 1 lower and cell m 1 markedly larger than cell cu 1 ( Figs 2E View FIGURE 2 , 5C View FIGURE 5 , 13D View FIGURE 13 ); on other Eleutherococcus View in CoL species (where host plant is known)................................................. 2

2. Antennae shorter than 1.5 x head width; apex of paramere (in lateral profile) tapering to a slender, anteriorly directed point ( Fig. 16 View FIGURE 16 )............................................................ Triozidus ceratophorus ( Li, 2005) comb. nov.

- Antennae longer than 1.5 x head width; apex of paramere (in lateral profile) more or less blunt and directed inward or posteriorly ( Figs 3B View FIGURE 3 , 6B View FIGURE 6 , 14B View FIGURE 14 , 16 View FIGURE 16 )................................................................................. 3

3. Body including forewing shorter than 4.5 mm; distal aedeagus segment lacking medial projections ( Figs 3C View FIGURE 3 , 16 View FIGURE 16 )......... 4

- Body including forewing longer than 4.5 mm; distal aedeagus segment with distinct medial projections ( Figs 6C View FIGURE 6 , 14C View FIGURE 14 , 16 View FIGURE 16 ).. 5

4. Head and antennae entirely black ( Fig. 1A View FIGURE 1 ); paramere (in lateral profile) with distinct angular incision medially on anterior surface, apex rounded ( Figs 3A, 3B View FIGURE 3 , 16 View FIGURE 16 )............................................ Triozidus burckhardti sp. nov.

- Head mostly yellow-brown to orange-brown with probably only terminal antennal segments black; paramere (in lateral profile) without angular medial incision but with slight incision near the base on anterior surface, apex elongate and extended rearward ( Fig. 16 View FIGURE 16 )............................................... Triozidus eleutherococci ( Konovalova, 1980) comb. nov.

5. Genal processes shorter (length just over half vertex length, <0.3 head width) and more blunt apically ( Fig. 13A View FIGURE 13 ); forewing with distinct dark marginal patch around clavus, but without a small dark spot at trifurcation of R+M+Cu ( Fig. 13D View FIGURE 13 ); forewing broader (ratio length:width <2.6), widest distal to middle, length approximately 2 x Rs vein length ( Fig. 13D View FIGURE 13 ); paramere broader (in lateral profile) ( Figs 14A, 14B View FIGURE 14 , 16 View FIGURE 16 ); distal aedeagus segment with medial projections narrow and thumb-like ( Figs 14C View FIGURE 14 , 16 View FIGURE 16 ); female terminalia shorter (length of proctiger less than head width and <3 x anal ring length) ( Fig. 14D View FIGURE 14 ); on Eleutherococcus trifoliatus View in CoL ..................................................................... Triozidus yangorum sp. nov.

- Genal processes longer (length approximately equal to vertex length,> 0.3 x head width) and more acute apically ( Fig. 5A View FIGURE 5 ); forewing without distinct dark marginal patch around clavus, but with a small dark spot at trifurcation of R+M+Cu (in Japanese and South Korean material, see Fig. 5C View FIGURE 5 , but possibly absent in eastern Russia material); forewing narrower (ratio length:width> 2.6), widest in middle, length approximately 2.5 x Rs vein length; paramere narrower (in lateral profile) ( Figs 6A, 6B View FIGURE 6 , 16 View FIGURE 16 ); distal aedeagus segment with medial projections semi-circular ( Figs 6C View FIGURE 6 , 16 View FIGURE 16 ); female terminalia longer (length of proctiger greater than head width and> 3 x anal ring length) ( Fig. 6D View FIGURE 6 ); on Eleutherococcus sessiliflorus View in CoL and E. divaricatus View in CoL (or other Eleutherococcus spp. where introduced).......................... Triozidus stackelbergi ( Loginova, 1967) comb. nov.

Key to 5 th instar immatures of Triozidus

(known for three species)

1. Body covered in narrow truncate sectasetae ( Fig. 7C View FIGURE 7 )............... Triozidus stackelbergi ( Loginova, 1967) comb. nov.

- Body covered in long simple setae ( Figs 11C View FIGURE 11 , 15C View FIGURE 15 ).......................................................... 2

2. Circumanal ring heart-shaped (medial constriction mainly from anterior), outer ring composed of a mix of elongate and irregular shaped pores ( Fig. 15D View FIGURE 15 )......................................................... Triozidus yangorum sp. nov.

- Circumanal ring peanut-shaped (with antero-posterior medial constriction), outer ring composed of a single narrow row of elongate pores ( Fig. 11D View FIGURE 11 )............................................. Triozidus ukogi ( Shinji, 1940) comb. nov.