Mniotype dubiosa amitayus Volynkin & Han

Volynkin, Anton V., Matov, Alexei Yu., Behounek, Gottfried & Han, Hui-Lin, 2014, A review of the Palaearctic Mniotype adusta (Esper, 1790) species-group with description of a new species and six new subspecies (Lepidoptera: Noctuidae), Zootaxa 3796 (1), pp. 1-32 : 15-16

Mniotype dubiosa amitayus Volynkin & Han , ssp. n.

( Figs 46–48 , 83, 84 , 119 )

Type material. Holotype: male, East Tibet, Taba, 3500 m, 18.vi.1996, leg. W. Fickler. Slide 3210 Gyulai (Coll. PGM). Paratypes: 2 females, China, Sichuan prov., Lingshan Mts, env. Mianning, 3760 m., 1–3.vii.2010, leg. S. Murzin (Coll. PGM); 1 male, China, NW Yunnan, Baima Xue Shan, near Yak La pass, 3900 m, N 28°24.900’, 98°59.800’, 21–22.v.2012, leg. A. Floriani (Coll. PGM); 1 male, China, prov. Shaanxi, Tsinling Mts, Taibaishan Mt., 33°50' N, 107°44' E; 1500 m, 20–30. vi. 2005, leg. V. Siniaev & Team (Coll. PGM); 1 female, East-Tibet, Prov. Quamdo, Taba, 4000 m, 26.vi.1991, leg. W. Fickler (Coll. PGM); 1 female, China, prov. Shaanxi, Tsinling Mts, Fopin Mt., 33°35’ N, 108°01’ E, 01–30.xi.2004, leg.: V. Siniaev (Coll. PGM); 2 females, China, Sichuan, 3100 m, Aba pref., Heishui, 12.vii.2013, leg. M. Murzin & O. Shulga (Coll. PGM); 1 male, 1 female, China, Sichuan, 3950 m, Garzé Tibetan, Aut. Pref. Litang, 23.vi.2009, leg. Balázs Benedek (Coll. BBT); 1 male, [ China, SE Tibet] Xizang, Linzhi, Mt. Sejilashan, 20.vi.2011 (Coll. NEFU); 1 male, [W China, SE Tibet] Aut. Reg. Xizang, Linzhi, 15.vi.2011, Mt. Sejilashan, 3900 m. Zhao-Hui Pan [leg.] (Coll. NEFU); 1 female, Prov. Yunnan, China, 2007. vi.9., Xianggelila, Coll. H.-L. Han (Coll. NEFU); 1 female, East-Tibet, Taba, 3900 m, 18.vi.1996, leg. Willi Fickler (Coll. GRB); 2 males, 1 female, China, Gansu S.W., near Xiahe (Labrang), Sangke grassland, N 34”59.626’, E 102”33.808’, h- 3560 m., 07.vii.2010, Floriani & Saldaitis leg. (Coll. AFM); 1 female, China, Gansu S.W., near Xiahe (Labrang), N 35”11.968’, E 102”33.545’, h- 2930 m., 08.vii.2010, Floriani & Saldaitis leg. (Coll. AFM); 2 males, China, Gansu S.W., road from Zoige to Lugu, N 34”20.780’, E 102”19.938’, h- 3660 m., 06.vii.2010, Floriani & Saldaitis leg. (Coll. AFM); 3 females, China, Qinghai N.E., Lenglong Ling mt., Ning Chan pass, N 37”32.245’, E 101” 52.275’, h- 4000 m., 14.vii.2010, Floriani & Saldaitis leg. (Colls ASV and AFM); 1 female, China, Qinghai N.E., Daban Shan Mts N from Datong, N 37”13.230’, E 101”46.988’, h- 3800 m., 12. vii. 2010, Floriani & Saldaitis leg. (Coll. ASV); 1 male, China, N Sichuan, near Barkam, Zhe Gu Shan pass, N 31”51.528’, E 102”40.312’, h- 4100 m., 28-29.vii.2011, Floriani & Saldaitis leg. (Coll. AFM); 1 female, China, N Sichuan, road Barkam/Hong Yuan, N 32”10.353’, E 102” 29.692’, h- 3800 m., 22.vii.2011, Floriani & Saldaitis leg. (Coll. AFM); 2 males, 1 female, China, N Sichuan, road Songpan-Juizhaigou, N 33”08.770’, E 103”43.624’, h- 3000 m., 04.vii.2010, Floriani & Saldaitis leg. (Colls ASV and AFM); 1 female, China, N Sichuan, road Songpan- Juizhaigou, N 33”08.770’, E 103”43.624’, h- 3000 m., 23.vii.2011, Floriani & Saldaitis leg. (Coll. AFM); 1 male, 1 female, China, N Sichuan, road Songpan-Zoige, N 32”55.511’, E 103” 24.959’, h- 3500 m., 05.vii.2010, Floriani & Saldaitis leg. (Coll. AFM); 6 males, 3 females, China, Sichuan, Minshan, h- 2900 m., road Songpan-Juizhaigou, 5 km S Jarpo town, 5-8.vii.2012, Della Bruna & Floriani leg. (Coll. AFM); 2 males, China, Sichuan, Minshan, h- 3100 m., road Songpan-Juizhaigou, 16 km S Jarpo town, 5-11.vii.2012, Della Bruna & Floriani leg. (Coll. AFM); 1 male, China, Sichuan, Juizhaigou vic., Yazha vill., h- 2000 m., 07.vii.2012, Della Bruna & Floriani leg. (Coll. AFM); 4 males, China NW Yunnan, Baima Xue Shan, near Yak La pass, N 28”24.900’, E 98”59.800’, 21- 22.v.2012, h- 3900 m., leg. Floriani (Colls ASV and AFM); 1 female, China, Sichuan, 80 km south-west from Danba, 4000–4350 m. a.s.l., 8–16.vii.2012 (Coll. MDS); 1 male, 1 female, Bhutan (West), Prov. Paro, Weg z. Jomolhari, N Shinkarap, S Takengthanka (Seitental bei Sejande Zumba), LF, 3610 m, N 27º41'56'', E 89º18'39'', 15.v.2010, lg. Ratzel, Hauenstein, M. Falkenberg & Trush, ex coll. Ratzel, Karlsruhe, Germany (Coll. URK); 4 females, Bhutan (West), Prov. Paro, Weg z. Jomolhari, S Takethang N 27º44'09'', E 89º17'44'', LF, 13–14.v. 2010, 3840 m, leg. Ratzel, Falkenberg, Trush & Hauenstein, ex coll. Ratzel, Karlsruhe, Germany (Coll. URK); 2 females, Bhutan, Paro (NO), Haus v. Schwester Dasho Kado, N 27º27'38'', E 89º25'48'', LF, 19.v. 2010, 2350 m, leg. Ratzel, Falkenberg, Trush & Hauenstein, ex coll. Ratzel, Karlsruhe, Germany (Coll. URK).

Slides 3183 Gyulai, hhl-2406-1, hhl-2506-1 Han, AV1116 Volynkin (males), hhl-2503-2, hhl-2504-2 Han, 12096f, 12098f, 12099f Behounek, AV0777, AV1117 Volynkin (females).

Diagnosis. The new subspecies differs externally from M. dubiosa dubiosa in its somewhat larger size, darker colouration of forewings, less contrast pattern and poorly defined streaks of the marginal line.

Description. External morphology ( Figs 46–48 ). Wingspan 40–47 mm. Head, thorax and abdomen dark brown. Forewing moderately broad. Ground colour of forewing dark brown. Crosslines thin, double. Submarginal line pale, thin, strongly dentate. Marginal line present as a series of lengthwise blackish brown streaks between the veins. Basal dash short, thin, black. Claviform dark brown, with thin black border; medial fascia thin, black. Medial area shadowed posteriorly of claviform and medial fascia. Terminal area dark brown. Subterminal area paler, brown, inwards from the submarginal line with blackish cuneate streaks between the veins. Cilia dark brown. Hindwing pale greyish brown, discal spot diffuse, semilunar. Cilia brownish grey. Male genitalia ( Figs 83, 84 ): differ from M. dubiosa dubiosa in the somewhat broader juxta, somewhat longer and apically pointed protruding part of clavus, smaller teeth on the apex of carina and longer eversible dorso-lateral bar of carina. Female genitalia ( Fig. 119 ): differ from M. dubiosa dubiosa in the more curved ductus bursae.

Distribution. Himalayan-Tibetan. West China (Tibetan plateau and Tsinling Mts) and Bhutan (North East Himalaya).

Etymology. Amitayus is the Buddha of Infinite Life in Tibetan Buddism.

Figures

FIGURES 41 – 50. Mniotype spp. adults: 41, M. anilis anilis, male, Italy, Lazio region (GBG / ZSM); 42, M. anilis sylvatica, male, France, Corsica (GBG / ZSM); 43, M. dubiosa dubiosa, lectotype male, " Mustagata Yarkend " (NKMB); 44, ditto, male, Tajikistan, Pamir Mts (ZISP); 45, ditto, female, SE Kazakhstan (AVB); 46, M. dubiosa amitayus, holotype male, China, Tibet Plateau (PGM); 47, ditto, paratype female, China, Sichuan Prov. (BBT); 48, ditto, paratype male, Bhutan (URK); 49, M. krisztina, male, China, Tibet Plateau (BBT) (Photo by B. Benedek); 50, ditto, female, China, Tibet Plateau (CAU).

FIGURES 81 – 86. Mniotype spp., male genitalia: 81, M. dubiosa dubiosa, lectotype, Muztagh-Ata Mt., slide MB 355 Boursin; 82, M. d. dubiosa, Kyrgyzstan, slide AV 0693 Volynkin; 83, M. d. amitayus, holotype, China, Tibet Plateau, slide 3210 Gyulai; 84, M. d. amitayus, paratype, Bhutan, Prov. Paro, slide AV 1116 Volynkin; 85, M. krisztina, W China, slide hhl- 2162 - 1 Han; 86, M. krisztina, W China, slide hhl- 2272 - 1 Han.

FIGURES 113 – 120. Mniotype spp., female genitalia: 113, M. bathensis, South Siberia, slide AV 0725 Volynkin; 114, M. aulombardi, paratype, Taiwan, slide 8144 f Behounek; 115, M. anilis anilis, France, slide GB 8142 f Behounek; 116, M. a. sylvatica, Corsica, slide AV 0787 Volynkin; 117, M. dubiosa dubiosa, paralectotype, " Mustagata Yarkend ", slide 7659; 118, ditto, SE Kazakhstan, slide AV 0878 Volynkin; 119, M. d. amitayus, paratype, W China, Tibet Plateau, slide AV 0777 Volynkin; 120, M. krisztina, China, Tibet, slide hhl- 2271 - 2 Han.