Physornis fortis Ameghino, 1895
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https://doi.org/ 10.1590/S0031-10492003000400001 |
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https://treatment.plazi.org/id/03D18A23-C84F-1850-59DC-2A5294D0FBBD |
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Carolina |
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Physornis fortis Ameghino, 1895 |
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Physornis fortis Ameghino, 1895
Physornis fortis Ameghino, 1895:576 ; Brodkorb, 1967. Aucornis euryrhyncus Ameghino, 1898:9 ; Brodkorb, 1967
(syn. of Physornis fortis ).
Type – A fragment around 137 cm in length, constituting the ventral portion of the mandibular symphysis and the adjacent part of the right mandibular branch ( BMNH-A583 : Fig. 14 View FIGURE 14 ) .
A proximal portion of the left tarsometatarsus (MACN-A-52-185, Figs. 2A View FIGURE 2 and 13 View FIGURE 13 ). A rostral extremity of the mandibular symphysis (MACN-A-52-186). A phalanx 1 of the toe II from the left foot (MACN-A-52-187); these last three bones were associated and served as type for Ancornis euryrhyncus Ameghino, 1898 . A phalanx 1 of the toe IV from the left foot (MACN-A-52-188). In the Museum of Amherst College , Massachussetts, U.S.A., there is also a right femur, incomplete on the proximal extremity, without number (?), refered to and figured by Loomis (1914:226) .
Horizon and Locality – Middle to Upper Oligocene of Argentina (Deseadan). Province of Santa Cruz, Argentina: Puerto Deseado, Punta Nova, La Flexa, in the levels of occurrence of Pyrotherium . For a better indication of the geological age, see Mac Fadden (1985) and Marshall et al. (1986).
Measurements – Table 3.
Illustrations – Loomis, 1914: Fig. 149.
Remarks – The size of the mandibular symphysis (type: BMNH-A583), originally described by Ameghino (1895), is in perfect agreement with the two measured specimens in the Field Museum of Chicago ( Table 3). A flat (not convex) region in the mid-ventral portion of the symphysis is noted in all the specimens.
The branches of the mandible of Physornis , in comparison with those of Phorusrhacos , are really more further apart from the median line, as in the original description by Ameghino (op. cit.), which implies a skull with a wider base. Ameghino perfectly described the only large built Phorusrhacidae of the stratigraphic layers characterized by the presence of Pyrotherium (Deseadan) . Thus, there is no reason for the rejection of the genus Physornis and the species P. fortis , as being indeterminate, as suggested by Patterson (1941).
Although the complete shape of the tarsometatarsus of Physornis is unknown, the proximal portion of this is shown to be quite flattened in the dorsoventral extent. Associating this to the short and wide mandibular symphysis, besides the quite large build, it fits in perfectly amongst the Brontornithinae .
Aucornis solidus Ameghino, 1898 , considered by Brodkorb (1967) as synonymous of Physornis fortis , was created only based on a proximal extremity of a phalanx (MACN-A-52-110), of a much smaller-sized bird than Physornis fortis , possibly synonymous of Andrewsornis abbotti Patterson, 1941 , which should be left as a species inquirenda, seeing that one is dealing with an absolutely insufficient segment to diagnose a species.
The two symphysis examined in the Field Museum of Chicago, showed an interesting difference on the dorsal surface, which, almost flat in FM-P13340, forms a longitudinal channel in FM-P13340 ( Figs. 12C and 12D View FIGURE 12 ). These differences were interpreted as being individual variation.
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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