Janalychas, Kovařík, 2019

Kovařík, František, 2019, Taxonomic reassessment of the genera Lychas, Mesobuthus, and Olivierus, with descriptions of four new genera (Scorpiones: Buthidae), Euscorpius 288, pp. 1-27 : 6

Genus Janalychas gen. n.

( Figures 10, 14, 38–43, 66–71, 95–100, 102–103) http://zoobank. org/urn:lsid:zoobank. org:act:D3B3A7AA-9947-

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Lychas (in part) C. L. Koch, 1845: 3; Tikader & Bastawade, 1983: 40–107, figs. 185–215, 240–255; Fet & Lowe, 2000: 158–169 (complete references list until 2000); KovařÍk & Ojanguren Affilastro, 2013: 194–210, figs. 1424–1431, 1437–1438, 1450–1470, 1557–1558, 1593–1600.

TYPE SPECIES. Lychas srilankensis Lourenço, 1997 .

ETYMOLOGY. The generic name is a matronym honoring Jana ŠtundlovÁ (née PlÍškovÁ) (Prague, Czech Republic) for her contributions to scorpion systematics, especially by applying karyotype study.

DIAGNOSIS. Total length 30–65 mm. Carapace granular, lacking distinct carinae, flat, subrectangular with concave anterior margin. Median eyes on ocular tubercle in anterior half of carapace; usually with 4, or sometimes 5 pairs of lateral eyes (3 major ocelli, 1–2 minor ocelli). Sternum type 1, triangular in shape. Tergites I–VI granular, with single median carina, tergite VII with 5 carinae. Metasoma elongate, segment I with 10 carinae, II–III with 8–10 carinae, IV with 8 carinae. Telson ellipsoidal in shape, with distinct subaculear tooth. Pectinal tooth counts 21–26. Pectines with fulcra. Chelicerae with typical buthid dentition, fixed finger armed with single denticle on ventral surface. Pedipalps orthobothriotaxic, type Aβ, femur trichobothrium d 2 obviously internal, patella d 3 external to dorsomedian carina. Dentate margins of chela movable finger armed with 3 terminal (two subterminal and one distal) denticles, apical row composed of 3–7 denticles, and imbricated overlapping six rows of denticles, rows 1–5 terminated proximally flanked by two enlarged outer accessory denticles, and single inner accessory denticle displaced distally. Sixth row without outer and inner accessory denticles. Elongated tibial spurs present on leg III and leg IV, tibia and tarsus without bristle combs, ventral surfaces of tarsomeres II densely equipped with two rows of setae, ungues stout.

SUBORDINATE TAXA (7 SPECIES).

Janalychas albimanus (Henderson, 1919) , comb. n. ( India)

Janalychas farkasi (KovařÍk, 1997) , comb. n. ( Nepal)

Janalychas heurtaultae (KovařÍk, 1997) , comb. n. ( Nepal)

Janalychas laevifrons (Pocock, 1897) , comb. n. ( India, Nepal)

Janalychas shoplandi (Oates, 1888) , comb. n. ( Myanmar)

=? Isometrus feae Thorell, 1889

Janalychas srilankensis ( Lourenço, 1997) , comb. n. ( Sri Lanka)

= Lychas ceylonensis Lourenço & Huber, 1999

Janalychas tricarinatus (Simon, 1884) , comb. n. ( India)

COMMENTS. Janalychas srilankensis comb. n. was originally described as Lychas by Lourenço (1997) where it was also placed so far (e. g. Fet & Lowe, 2000; KovařÍk & Ojanguren Affilastro, 2013; KovařÍk et al., 2016). Morphological characters cited in the diagnosis above indicate that this placement was not correct, and this species, together with six other species from India, Nepal, and Myanmar, represents a separate new genus related to Sri Lankan species of Reddyanus Vachon, 1972 and Isometrus Ehrenberg, 1828 (see KovařÍk et al., 2016).