Thevenetimyia tenta Hall, 1969

Li, Xuankun, Rodrigues, Paula Fernanda Motta, Lamas, Carlos José Einicker & Yeates, David K., 2018, A Review of the Australian Species of Thevenetimyia Bigot, 1875 (Bombyliidae, Bombyliinae, Eclimini), with Description of Four New Species and the Pupal Case of T. longipalpis (Hardy), Records of the Australian Museum 70 (3), pp. 331-375 : 367

Thevenetimyia tenta Hall, 1969

Figs 26–28

Thevenetimyia tenta Hall, 1969: 75 . Type-locality: Australia (NSW) ; holotype, ANIC .

Types and other specimens examined. Holotype ♂, Australia, NSW, Currajong [spelt “Currajang” in Hull (1969: 76), might be a synonym of the Blue Mountains town “Kurrajong”], Macl [eay] M[useum], now in ANIC 29-009852 ( Figs 26 , 28g ). QLD, 1♀, Rockhampton , Macl [eay] M[useum], ANIC 29-041303 ; 2♂♂, QLD, Rockpool Gorge , Mt Walsh NP, Biggenden, 4 Oct 1976, H. Frauca, ANIC 29-041299 – 300 ; 1♂, QLD, Bluff Ra. foothills, Biggenden, Sep 1974, H. Frauca, ANIC 29-041301 ; 1♀, QLD, Sunnybank, 15 Oct 1939, C. F. Ashby, ANIC 29-041302 .

Diagnosis. Medium to small-sized Thevenetimyia , wing membrane with large dark brown infuscation on apical half, middle of cell r 2+3 with a narrow lighter brown spot, wing apex hyaline; costa with two rows of small spines; scutum and scutellum with golden scales; abdominal tergites 6 to 8 with dense lateral long black hairs.

Redescription. Female. Body length 8.1–13.2 mm, wing length 7.9–11.7 mm.

Head. Head about 1.4× wider than long, mostly blackish with thick pale pruinescence and covered in admixed white to blackish brown hairs. Frons blackish brown with thick pale pruinescence, 2.5× as wide as ocellar tubercle, frons with blackish brown hairs ( Fig. 26e ). Ocellar tubercle slightly raised, brown to blackish brown with grey pruinescence, with blackish brown hairs. Face with thin pale pruinescence, lateral area of antenna with some brown hairs, parafacial area dark brown and bare. Gena with thin pale pruinescence and long white hairs. Clypeus swollen, with thick pale pruinescence and admixed white and brown hairs. Occiput with long white hairs except dorsal margin with blackish brown hairs. Posterior eye margin slightly sinuous. Antennae blackish brown; scape and pedicel with thin pale pruinescence and black hairs, ventral face of scape admixed with white hairs. Scape 3.4× as long as wide, and 3.4× as long as pedicel, uniform from base to apex. Pedicel 0.9× as long as wide. Flagellum 5.0× as long as wide, 1.4× as long as scape+pedicel (1.8× as long as scape), conical and slightly laterally compressed, one-segmented with apical style ( Fig. 26c ). Palp thin and long, extending beyond oral cavity, black with blackish brown hairs, two-segmented, with palpal pit. [Mouthparts broken, apex of proboscis missing.]

Thorax. Integumental colour of scutum and scutellum mostly black with sparse grey pruinescence, pruinescence more dense around margins. Scutum and scutellum with golden scales admixed with a few blackish brown hairs, except margin with dense white scales. Three notopleural setae present, postalar seta absent. Postalar callus and posterior margin of scutellum with strong hairs. Notopleural setae pale yellow, strong hairs on postalar callus and posterior margin of scutellum pale yellow to white. Pleura and coxae black with thick grey pruinescence, anepisternum, dorsal half of katepisternum and metepisternum with long white hairs, mediotergite with dense white hairs and setae, anepimeron, meron and laterotergite bare.

Legs. Legs brownish yellow, mostly covered with blackish brown scales, except posterior face of femora with white scales. Other hairs and bristles on legs short and blackish brown to black. Fore tibia 2.0× longer than fore basitarsus, mid tibia 2.0× longer than mid basitarsus. [Hind tarsus broken.]

Wings. Wing membrane with large brown infuscation on apical half, but wing apex hyaline. Apical 0.4 of cell r 1, apical 0.9 to 0.6 of cell r 2+3, basal 0.4 of cell r 4, basal 0.8 of cell r 5, apical 0.3 of cell dm, basal 0.9 of cell m 1, apical 0.8 of cell m 2, and apical 0.5 of cell cua 1 all covered by the infuscation, anterior darker than posterior, posterior without a distinct margin. Cell r 5 widely open; cell br much longer than cell bm, crossvein r-m arising a little over half way from the base of cell dm; crossvein m-m long, and 2.0× as long as crossvein r-m; cell cup open ( Fig. 26d ). Costa without small spines. Haltere stem pale yellow, knob black.

Abdomen. Integumental colour of tergites dark brown with thin pale pruinescence, except lateral margin with thick pruinescence. Tergites mostly covered with blackish brown scales and short black hairs. Tergite 1 covered with admixed white and pale yellow scales lateral with long white hairs. Tergites 2 to 5 with white scales laterally. Tergites 6 and 7 with long black hairs laterally. Sternites with dark brown integumental colour except posterior margins yellow, with thick grey pruinescence and white hairs. Genitalia. Posterior margin of tergite 7 with slender dorsal median apodeme. Tergite 7 with ventral spines. Tergite 8 with a row of sparse hairs, 11 acanthophorite spines present, lateral spines long, spines in middle short, apex of spines expanded. Furca narrow and straight, connected at apex. Sperm pump strong and nearly as long as furca, not clothed in longitudinal muscle, with few lateral papillae; distal spermathecal duct short but strong; spermatheca elongate, cylindrical ( Fig. 28g ).

Male. Body length 13.7–15.0 mm, wing length 12.1–13.5 mm. Very similar to female, except wing costa with two rows of short, thick spines. Genitalia. Epandrium wide and short, posterior margin slightly convex ( Fig. 28d ). Hypandrium present. Gonocoxal apex slightly narrower than the base in dorsal view, ejaculatory apodeme large ( Fig. 28a,b ); gonocoxal apodeme strong and incurved; lateral ejaculatory process strong, 2.2× longer than wide, apex expanded; inner apex of gonocoxite rounded, elongate and broad; outer apex of gonocoxite rounded; dorsal bridge with long lateral hollow ( Fig. 28c ); gonostylus slender and pointed dorsally, 3.2× longer than wide.

Remarks. Thevenetimyia tenta Hall, 1969 is redescribed and the male is described for the first time. The holotype of Thevenetimyia tenta Hall, 1969 is smaller than all other specimens we examined. Some specimens middle of wing cell r 2+3 with a narrow lighter brown spot.

Distribution. NSW, QLD.

Figures

Figure 26. Thevenetimyia tenta Hall, 1969 ♀ holotype: (a) dorsal; (b) lateral; (c) flagellum; (d) wing; (e) head, dorsal; (f) head, lateral; (g) head, frontal; (h) head, profile. Scale bars = 1 mm (a, b, d–h); = 0.1 mm (c).

Figure 27. Thevenetimyia tenta Hall, 1969 ♂: (a) dorsal; (b) lateral;(c) flagellum; (d) wing; (e) head, dorsal; (f) head, lateral;(g) head, frontal; (h) head, profile. Scale bars = 1 mm (a, b, d–h); = 0.1 mm (c).

Figure 28. Thevenetimyia tenta Hall, 1969 genitalia (a–e ♂; f, g ♀): (a) genital capsule, dorsal; (b) genital capsule, ventral;(c) genital capsule, lateral; (d) epandrium, dorsal;(e) epandrium, lateral;(f) sternite 8, ventral; (g) genitalia and spermathecae. Scale bars = 0.1 mm.