Thevenetimyia major Li and Yeates, 2018

Li, Xuankun, Rodrigues, Paula Fernanda Motta, Lamas, Carlos José Einicker & Yeates, David K., 2018, A Review of the Australian Species of Thevenetimyia Bigot, 1875 (Bombyliidae, Bombyliinae, Eclimini), with Description of Four New Species and the Pupal Case of T. longipalpis (Hardy), Records of the Australian Museum 70 (3), pp. 331-375 : 353-356

Thevenetimyia major Li and Yeates , sp. nov.

Figs 16–17

Holotype ♂, Australia, QLD, 30 km W of Alpha , T. M. S. Hanlon, ex dead standing Acacia , collected 30 Aug 1998, emerged 16 Nov 1999, AM K.364586 . Paratype 1♂, NT, Koongarra , 15 km E of Mt Cahill, 12°52'S 132°50'E, 6 Oct 1975, A . Allwood & T. Angeles, ANIC 29-041486 .

Diagnosis. Large-sized Thevenetimyia , wing membrane with large brown mark on middle area, but remaining wing apex hyaline, base mostly infuscated; vein R 4 arising far from base, 0.8 from crossvein r-m; costa without small spines; hind femur with a row of short black bristles; abdominal tergites 5 to 8 with dense lateral long black hairs.

Description. Male. Body length 14.6–16.7 mm, wing length 12.7–c. 12.9 mm.

Head. Head about 1.7× wider than long, mostly blackish with thin pale pruinescence and covered in admixed white, pale yellow and black hairs. Eye narrowly separated, by 0.2× width of ocellus. Frons slender and long, upper triangular section small, lower triangular section black, middle narrow part about 0.7× length of frons. Ocellar tubercle slightly raised, black with grey pruinescence, with black hairs. Face with thin pale pruinescence, lateral area of antenna with some black hairs, parafacial area black and bare. Gena with thick pale pruinescence and long white hairs. Clypeus swollen, with thin pale pruinescence and blackish brown hairs. Occiput mostly with pale yellow hairs, except ventral area with white hairs and dorsal margin with black hairs. Posterior eye margin slightly sinuous. Antennae blackish brown; scape and pedicel with thin pale pruinescence and black hairs, ventral face of scape admixed with white hairs. Scape 3.2× as long as wide, and 3.4× as long as pedicel, uniform from base to apex. Pedicel 1.1× as long as wide. Flagellum 8.3× as long as wide, 1.4× as long as scape+pedicel (1.8× as long as scape), conical and slightly laterally compressed, onesegmented with apical style ( Fig. 16c ). Palp thin and long, extending beyond oral cavity, black with blackish brown hairs, two-segmented, with palpal pit. Mouthparts slender, 3.5× as long as eye length (2.0× as long as head length), labella thin and filiform ( Fig. 16f ).

Thorax. Integumental colour of scutum and scutellum mostly black with sparse grey pruinescence, pruinescence more dense around margins. Scutum with short black hairs, anterior half and lateral margins admixed with dense long white scales, posterior half with long golden scales. Scutellum covered with admixed white and yellow scales and black hairs, posterior margin with black setae. Three pale yellow notopleural setae, and six pale yellow postalar setae present. Pleura and coxae black with thin pale pruinescence, anepisternum and katepisternum with admixed white and black hairs, dorsal half of metepisternum with long white scales, mediotergite with dense white hairs and setae, anepimeron, meron and laterotergite bare.

Legs. Legs black, mostly covered with black scales, except basal half of hind femur and posterior face of mid femur and tibia admixed with white scales, ventral face of fore and mid femora with short black hairs, hind femur with a row of short black bristles. Other hairs and bristles on legs short and blackish brown to black. Fore tibia 1.6× longer than fore basitarsus, mid tibia 1.7× longer than mid basitarsus, hind tibia 1.9× longer than hind basitarsus.

Wings. Wing membrane with large brown mark in median area, but wing apex hyaline, base mostly infuscated. Cell r 1, base 0.7 of cell r 2+3, base 0.55 of cell r 5, base 0.1 of cell m 1, base 0.9 of cell m 2, cell cua 1 and cell dm, apex 0.3 of cell cup, apex 0.4 of cell br and apex 0.2 of cell bm all covered with brown infuscation, except cell cua 1 and cell dm with small middle clear area. Cell r 5 widely open; cell br much longer than cell bm, crossvein r-m arising half way from the base of cell dm; crossvein m-m long, and 2.0× as long as crossvein r-m; cell cup open ( Fig. 16d ). Vein R 4 arising 0.8 from crossvein r-m. [Wing apex broken]. Costa without small spines. Haltere stem dark yellow, knob blackish brown.

Abdomen. Integumental colour of tergites dark brown with thin pale pruinescence, except lateral margin with thick pruinescence. Tergite 1 with dense long white hairs. Tergites 2 to 8 mostly covered with black scales and short black hairs, tergite 2 admixed with long white hairs and posterior margin with white scales, tergites 3 to 4 with long white hairs laterally, posterolateral with dense white scales, tergites 5 to 8 with dense long black hairs laterally, tergite 5 with dense white scales on dorsal area. Sternites with black integumental colour except posterior margins yellow, mostly with thick grey pruinescence and white hairs, except sternites 5 to 8 covered with black hairs. Genitalia. Epandrium wide and short, posterior margin slightly convex ( Fig. 17d ). Hypandrium present. Gonocoxal apex slightly narrower than the base in dorsal view, ejaculatory apodeme large ( Figs 17a, b ); gonocoxal apodeme strong and incurved; lateral ejaculatory process strong, 3.7× longer than wide, apex expanded; inner apex of gonocoxite rounded, elongate and broad; outer apex of gonocoxite rounded; dorsal bridge with long lateral hollow ( Fig. 17c ); gonostylus slender and pointed dorsally, 3.0× longer than wide.

Female. Unknown.

Remarks. Thevenetimyia major Li and Yeates , sp. nov. differs markedly from congeners by having a large body size, a large brown mark on middle area of wing, and a vein R 4 arising far from base, 0.8 from crossvein r-m.

The paratype is without clear area in the middle of brown infuscation.

Distribution. QLD, NT.

Etymology. This specific name refers to the large body size of the species.

Figures

Figure 16. Thevenetimyia major Li and Yeates, sp. nov. ♂: (a) dorsal; (b) lateral; (c) flagellum; (d) wing; (e) head, dorsal; (f) head, lateral; (g) head, frontal; (h) head, profile. Scale bars = 1 mm (a, b, d–h); = 0.1 mm (c).

Figure 17. Thevenetimyia major Li and Yeates, sp. nov. ♂ genitalia: (a) genital capsule, dorsal; (b) genital capsule, ventral; (c) genital capsule, lateral;(d) epandrium, dorsal;(e) epandrium, lateral. Scale bars = 0.1 mm.