Thevenetimyia Bigot, 1875

Li, Xuankun, Rodrigues, Paula Fernanda Motta, Lamas, Carlos José Einicker & Yeates, David K., 2018, A Review of the Australian Species of Thevenetimyia Bigot, 1875 (Bombyliidae, Bombyliinae, Eclimini), with Description of Four New Species and the Pupal Case of T. longipalpis (Hardy), Records of the Australian Museum 70 (3), pp. 331-375 : 333

Genus Thevenetimyia Bigot

Thevenemyia Bigot, 1875 a: 196 [1875b: clxxiv]. Type species: Thevenetimyia californica Bigot, 1875 , by monotypy. [Unavailable name; incorrect original spelling.]

Thevenetimyia Bigot, 1875 a: 196 [1875b: clxxiv] (justified emendation of Thevenetimyia [by Bigot, 1892: 339]). Type species: Thevenetimyia californica Bigot, 1875 , automatic.

Epibates Osten Sacken, 1877: 268 . Type species: Epibates funestus Osten Sacken, 1877 , by subsequent designation (Coquillett, 1910b: 538).

Diagnosis. Hall (1969) summarized as “Quite similar to Eclimus Loew. The chief differences lie in the slightly swollen occiput, the smooth integument, partial development of the alula, the narrowly open anal cell and the absence of transverse abdominal depressions in Thevene[ti]myia.” Based on the Australian species, we further diagnose this genus as follows:

Mostly medium-sized, slender Bombyliinae . Male eyes mostly connected or narrowly separated ( Figs 1e, 4e, 7e, 10e, 13e, 16e, 18e, 20d, 23e, 27e). Female frons about 2.5× as wide as ocellar tubercle ( Figs 2d, 8c, 14c, 21c, 24c, 26e). Clypeus swollen. Flagellum one-segmented with apical style. Palp thin and long, extending beyond oral cavity, black with blackish brown hairs, two-segmented, with palpal pit. Mouthparts slender, labella thin and filiform ( Figs 1f, 2e, 4f, 7f, 8d, 10f, 13f, 14d, 16f, 18f, 20e, 21d, 23f, 24d, 27f). Scutum and scutellum with admixed hairs and scales. Three to four notopleural setae present, postalar seta normally absent. Scutum sometimes with small short spines ( Fig. 31d). Anepisternum, katepisternum, metepisternum, and mediotergite with hairs and/or setae, anepimeron, meron and laterotergite bare. Hind femora sometimes with short ventral bristles. Wing membrane various, from hyaline to mostly infuscated to having distinct dark marking on otherwise transparent wings. Female wing colour lighter in some species. Cell r 5 widely open; cell br much longer than cell bm; crossvein m-m long; cell cup narrowly open ( Figs 1d, 2c, 4d, 7d, 8e, 10d, 13d, 16d, 18d, 19c, 21e, 23d, 26d, 27d). Male costa often with two rows of small spines ( Fig. 31b), absent in females ( Fig. 31a). Abdomen slender, tergites often with pale scales on posterior margin, apex tergites sometimes with dense long lateral hairs. Male epandrium wide and short, posterior margin slightly convex ( Figs 3d, 5d, 9d, 11d, 15d, 17d, 22d, 25d, 28d). Hypandrium normally present. Gonocoxal apex slightly narrower than the base in dorsal view; gonocoxal apodeme strong and incurved; inner apex of gonocoxite rounded, elongate and broad; outer apex of gonocoxite normal length, rounded; gonostylus slender and pointed dorsally ( Figs 3a,b; 5a,b; 9a,b; 11a,b; 15a,b; 17a,b; 22a,b; 25a,b; 28a,b). Female posterior margin of tergite 7 with slender dorsal median apodeme. Tergite 7 with ventral spines. Tergite 8 with a row of sparse hairs, acanthophorite spines present, lateral spines long, spines in middle short, apex of spines expanded. Furca connected at apex. Sperm pump strong and nearly as long as furca, not clothed in longitudinal muscle; distal spermathecal duct short but strong; spermatheca elongate, cylindrical ( Figs 3g, 9g, 15g, 25g, 28g).

Distribution. Nearctic ( Canada, Mexico, USA), Palaearctic ( Algeria, Greece, Iran, Japan, Tunisia, Turkey), Afrotropical ( Mauritania, Madagascar), Neotropical ( Mexico [Oaxaca]) and Australian (all Australian states and territories except TAS).

Australian species. Thevenetimyia australiensis Hall, 1969 ; Thevenetimyia fergusoni Li and Rodrigues , sp. nov.; Thevenetimyia furvicostata ( Roberts, 1929) ; Thevenetimyia infuscata Li and Yeates , sp. nov.; Thevenetimyia longipalpis ( Hardy, 1921) ; Thevenetimyia major Li and Yeates , sp. nov.; Thevenetimyia mimula Hall, 1969 ; Thevenetimyia nigrapicalis ( Roberts, 1929) ; Thevenetimyia nuri Rodrigues and Lamas , sp. nov.; Thevenetimyia tenta Hall, 1969 .